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Sedimentological and ecological significance of a biodeformational structure associated with an unusual feeding behavior in gulls (Larus sp.) 与海鸥异常摄食行为相关的生物变形结构的沉积学和生态学意义(Larus sp.)
IF 0.8 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2022-05-29 DOI: 10.1080/10420940.2022.2067535
S. Melnyk, Cody N. Lazowski, M. Gingras
Abstract Shorebirds produce a range of biogenic sedimentary structures related to their feeding behaviors, the most common of which are probing behaviors that result in morphologically simple impressions or indentations on the surface of a sedimentary substrate. Here we describe an unusual biodeformational structure made by a feeding gull (Larus sp.) produced as the tracemaker kneaded the sediment by moving its feet back and forth in place. This process liquified the sediment making it easier to catch invertebrate prey — the most common of which is the varnish clam (Nuttallia obscurata) — that are advected to the top of the liquified sediment. The bird retreated slowly backward as the sediment was liquified, an effort that is interpreted to represent continued and ongoing feeding in the sediment of prey-rich substrates. The resulting structure comprises a series of nested, concavo-convex sediment mounds and a terminal bowl-shaped impression. The aim of this paper is to document the trace–tracemaker association and provide a means of comparison with similar structures. Most notably, the structure bears similarities to Piscichnus, which represents ray feeding traces produced by hydraulic jetting (fodichnia) or nesting behaviors (calichnia) in fish. Piscichnus, however, lacks the nested mounds that result from the tracemaker systematically moving backward as it searches for food.
滨鸟会产生一系列与其摄食行为相关的生物成因沉积结构,其中最常见的是探测行为,这种行为会在沉积基底表面产生形态简单的印痕或凹痕。在这里,我们描述了一种不寻常的生物变形结构,它是由一只进食的海鸥(Larus sp.)在追踪器通过来回移动它的脚来揉捏沉积物时产生的。这一过程使沉积物液化,从而更容易捕获无脊椎动物猎物——其中最常见的是漆蛤(Nuttallia obscurata)——它们被平流到液化沉积物的顶部。随着沉积物被液化,鸟慢慢后退,这一努力被解释为在富含猎物的基质的沉积物中持续不断地进食。由此产生的结构包括一系列嵌套的、凹凸的沉积物丘和一个末端碗形的印象。本文的目的是记录示踪-示踪器关联,并提供与类似结构进行比较的方法。最值得注意的是,这种结构与Piscichnus有相似之处,它代表了鱼的水力喷射(fodichnia)或筑巢行为(calichnia)产生的射线摄食痕迹。然而,在寻找食物的过程中,这种追踪器系统地向后移动,导致了Piscichnus没有巢丘。
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引用次数: 1
Defining and refining principles in ichnotaxonomy: Markus Bertling (1959–2022) 定义和完善技术分类学的原则:马库斯·伯特林(1959-2022)
IF 0.8 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2022-05-14 DOI: 10.1080/10420940.2022.2057481
M. Wisshak, D. Knaust, Lothar H. Vallon, A. Rindsberg
In the peaceful days before Christmas 2021, fate struck hard on fellow ichnologist Markus Bertling (Figure 1), when he learned of a devastating diagnosis that gave him very little chance to turn the tide. On February 13, 2022, Markus passed away, leaving behind a painful void in his family and the ichnologic community alike. Starting off in 1985 with research on Upper Jurassic coral reef palaeoecology and sedimentology in northern Germany, Markus received his doctoral degree in 1990 and quickly picked upon an interest in the trace fossils he found in Jurassic reef settings and established his f irst new ichnogenus, Arachnostega Bertling, 1992. In the following years, it was the bioerosion trace fossils and the lessons to learn from the bioerosion at Mesozoic coral reef settings that caught his particular interest (e.g., Bertling, 1995, 1997a, 1999a, 1999b, 1999c, 2002; Perry & Bertling, 2000). Since 1998, Markus was curator for palaeontology at the Institute of Palaeontology of the Westphalian Wilhelm’s University in Münster, Germany, and in 2007 he became one of the heads of the University’s Geomuseum. For the past decade, Markus was heavily involved in the complete refurnishing and modernization of the museum and had less valency to follow his scientific passion in ichnology and ichnotaxonomy. Nevertheless, he sustained his interest in this field of science and was involved, for instance, in various case studies on bioerosion traces in osteic and xylic substrates (e.g., Feng et al., 2019; Höpner & Bertling, 2017; Mikuláš et al., 2020) and contributed with this expertise to an extensive review and annotated list of all the known bioerosion ichnotaxa (Wisshak et al., 2019). Aside from Arachnostega gastrochaenae, Markus co-authored the erection of at least three ichnospecies, three ichnogenera and fourteen ichnofamilies (Höpner & Bertling, 2017; Wisshak et al., 2019). Markus was best known to the ichnologic research community as a leading authority when it came to both the nomenclature of trace fossils and ichnotaxonomic principles. His effort helped to implement the rules established in the International Code of Zoological Nomenclature and to advocate their application in all kinds of ichnotaxonomic work. This, in turn, enabled the ichnotaxonomist to apply ichnotaxobases more consistently, to focus on the erection of ichnotaxa by a valid procedure and avoiding the erection of too many monotypic taxa. Whenever there was a problem to solve on how to apply the Code, the trickier the better, it became a reflex for many ichnologists to send an e-mail to Markus, asking for his advice. His in-depth knowledge of the Code put him into the position to lead or contribute to several comments on the draft proposal to emend the Code with respect to trace fossils (Bertling et al., 2003, 2004; Genise et al., 2004). From 2017 onward, Markus himself served as a commissioner on the Internat ional Commission for Zoologica l Nomenclature, striving to foster the statu
在2021年圣诞节前的平静日子里,命运沉重地打击了技术专家马库斯·伯特林(图1),他得知了一项毁灭性的诊断,几乎没有机会扭转局势。2022年2月13日,马库斯去世了,给他的家人和科技界留下了痛苦的空白。Markus于1985年开始研究德国北部的上侏罗纪珊瑚礁古生态学和沉积学,于1990年获得博士学位,并很快对他在侏罗纪珊瑚礁环境中发现的痕迹化石产生了兴趣,并于1992年建立了他的第一个新鱼属——Arachnostega Bertling。在随后的几年中,引起他特别兴趣的是生物侵蚀痕迹化石和中生代珊瑚礁环境中生物侵蚀的教训(例如,Bertling, 1995,1997a, 1999a, 1999b, 1999c, 2002;Perry & Bertling, 2000)。自1998年以来,马库斯一直是德国梅恩斯特威斯特伐利亚威廉大学古生物学研究所的古生物学馆长,并于2007年成为该大学地质博物馆的负责人之一。在过去的十年里,马库斯积极参与了博物馆的全面装修和现代化,并没有多少时间去追随他对技术和技术分类学的科学热情。尽管如此,他保持了对这一科学领域的兴趣,并参与了诸如骨和木质基质中生物侵蚀痕迹的各种案例研究(例如,Feng等人,2019;Höpner & Bertling, 2017;Mikuláš等人,2020),并利用这些专业知识对所有已知的生物侵蚀ichnotaxa进行了广泛的审查和注释列表(Wisshak等人,2019)。除了腹chaenae Arachnostega, Markus还参与了至少3个鱼属、3个鱼科和14个鱼科的直立研究(Höpner & Bertling, 2017;Wisshak et al., 2019)。马库斯在技术研究界最为人所知的是,他是痕迹化石命名法和技术分类学原理方面的权威。他的努力有助于实施《国际动物命名法法典》所建立的规则,并倡导将这些规则应用于各种技术分类工作。这反过来又使鱼类分类学家能够更加一致地应用鱼类分类基,通过有效的程序关注鱼类分类群的建立,并避免过多的单型分类群的建立。每当需要解决如何应用《准则》的问题时,越复杂越好,许多技术人员就会本能地给马库斯发电子邮件,征求他的建议。他对《法典》的深入了解使他能够领导或参与对《法典》有关化石修订提案草案的若干评论(Bertling et al., 2003,2004;Genise et al., 2004)。自2017年起,Markus本人担任国际动物命名委员会委员,致力于提升生物分类学在动物命名中的地位,并为正在进行的法典修订做出贡献。他对ICZN的参与受到了更广泛的分类学界的赞赏,为了纪念他的工作,脊椎动物属Bertlinggekko以他的名字命名。Markus致力于改进代码,证明了他在合理(方法)逻辑思维和科学认知理论方面的巨大天赋和浓厚兴趣——这是他在20世纪90年代公认的技术分类学混乱局面中带来的宝贵品质。马库斯在1998年成为技术分类学界的核心人物
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引用次数: 0
Tooth marks, gnaw marks, claw-marks, bite marks, scratch marks, etc: terminology in ichnology 牙痕、咬痕、爪痕、咬痕、抓痕等:技术术语
IF 0.8 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2022-04-12 DOI: 10.1080/10420940.2022.2058937
J. Zonneveld, A. Fiorillo, S. Hasiotis, M. Gingras
Abstract Phrases incorporating the modifier ‘mark’ (e.g., bite mark, tooth mark, gnaw mark, etc.) have recently come under attack. These phrases are wide-spread in their usage, and are, in fact, appropriate to the original definition of the word mark. Phrases such as bite mark and tooth mark are de rigueur as interpretive terms in the larger scientific community and in the archaeological, anthropological, pathological forensic and biological literature and are consistent with the original definition of the word mark. Longstanding convention in the ichnological literature as well as these diverse other disciplines underscores that usage of the word ‘mark’ as a modifier in ichnological analyses is both appropriate and useful for inter-disciplinary communication. Neither the words ‘mark’ nor ‘trace’ are clear terms on their own and become well-defined only when a modifying term is associated (e.g., bite mark or bite trace; trace fossil; ripple mark, fault trace, trace element, gnaw mark, etc.).
含有“标记”修饰语的短语(例如,咬痕、牙痕、咬痕等)最近受到了攻击。这些短语被广泛使用,事实上,它们适合于“标记”一词的原始定义。诸如咬痕和牙痕之类的短语在更大的科学界以及考古学、人类学、病理学法医和生物学文献中作为解释性术语是必要的,它们与“标记”一词的原始定义是一致的。在技术文献中以及这些不同的其他学科中,长期以来的惯例强调,在技术分析中使用“标记”一词作为修饰语,对于跨学科交流既合适又有用。“标记”和“痕迹”这两个词本身都不是明确的术语,只有当与一个修饰术语相关联时(例如,咬痕或咬痕;跟踪化石;波纹痕、故障痕、微量元素、咬痕等)。
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引用次数: 9
Cruziana and Helminthopsis in fluvial deposits of the uppermost Stockton Formation (Late Triassic), west-central New Jersey 新泽西州中西部斯托克顿组最上层(晚三叠世)河流沉积物中的克氏藻和蠕虫藻
IF 0.8 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2022-04-04 DOI: 10.1080/10420940.2022.2056168
R. Metz
Abstract Fluvial deposits of the uppermost Stockton Formation (Late Triassic), west-central New Jersey have yielded the trace fossils Cruziana tenella and Helminthopsis isp. The ichnotaxa belong to the Scoyenia ichnofacies. On the basis of stratification and primary sedimentary structures, the beds are interpreted as deposits in a meandering stream environment. Worm-like forms, nematodes, notostracans, myriapods, and arthropods are probably responsible for most of the animal traces in moist or wet channel and point bar sediments subject to aerial exposure.
摘要在美国新泽西州中西部晚三叠世斯托克顿组最上层河流沉积中,发现了Cruziana tenella和Helminthopsis等微量化石。鱼类群属于斯科耶尼亚鱼相。根据层理和原始沉积构造,认为该层为曲流环境下的沉积。在潮湿或潮湿的水道和点坝沉积物中,受空气暴露的大多数动物痕迹可能是由蠕虫状形式、线虫、无尾纲动物、多足类动物和节肢动物造成的。
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引用次数: 1
Site-selectivity of symbiotic (parasitic?) pits in crinoid column material from the middle Silurian (Wenlock: Sheinwoodian) of western Estonia 爱沙尼亚西部中志留世(Wenlock: Sheinwoodian)海红柱体材料中共生(寄生)坑的选择性
IF 0.8 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2022-03-15 DOI: 10.1080/10420940.2022.2047038
J. Thomka, O. Vinn, U. Toom
Abstract Crinoid specimens containing symbiotic (potentially parasitic) pits are relatively rare in Silurian strata of Estonia. The first specimen of middle Silurian-age (Wenlock: Sheinwoodian) crinoid material containing such pits—a pluricolumnal of uncertain affinity—was recently reported from the Jaagarahu Formation of Saaremaa Island. The presence and morphology of pits were previously described, but a noteworthy paleoecological phenomenon that was not initially documented involves the site-specificity of pit occurrence along the pluricolumnal. All 21 of the pits distributed around the lateral circumference of the specimen are located on plate sutures, demonstrating a degree of site-selectivity that contrasts strongly with the more random distribution of pits observed in most Silurian crinoid assemblages. The underlying paleoecological driver of this site-selectivity remains enigmatic, particularly considering some reports that suggest a preference for the latera rather than the sutures of Paleozoic crinoid columns.
在爱沙尼亚志留系地层中,含有共生(潜在寄生)坑的海百合标本相对较少。最近,在Saaremaa岛的Jaagarahu组中发现了第一个含有这种坑的中志留世(Wenlock: sheinwood dian)海麻类材料标本——一种不确定亲缘关系的多柱状物。坑的存在和形态之前已经被描述过,但一个值得注意的古生态现象最初并没有被记录下来,它涉及沿多柱带出现的坑的地点特异性。所有分布在标本外侧圆周的21个凹坑都位于板缝上,显示出一定程度的位置选择性,与大多数志留系海百合组合中观察到的更加随机的凹坑分布形成强烈对比。这种位置选择性的潜在古生态驱动因素仍然是个谜,特别是考虑到一些报告表明,古生代海百合柱的缝合线更受偏爱。
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引用次数: 2
Possibly the oldest fish-made resting traces 可能是最古老的鱼留下的休息痕迹
IF 0.8 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2022-03-09 DOI: 10.1080/10420940.2022.2047039
K. Ayranci, A. Wetzel, M. Kaminski, Bedri Kurtuluş, L. Rabaoui
Abstract The Silurian Sharawra Member (middle to late Llandoverian) in Saudi Arabia exhibits unique surface traces with exceptional preservation: Seven small, wing-shaped traces occur on a sandstone slab showing well-defined hummocky cross-stratification having a few decimeters wavelength being covered by a thin mud layer. These traces are preserved as concave epirelief along with elongated internal striae and surrounded by slightly elevated levees. Preservation of these delicate, 10–20 mm long and ∼10 mm wide traces suggests that they were formed after a storm event when mud had already settled to the sediment surface. The lack of wave-generated sedimentary structures implies deposition between fair-weather and storm-wave base probably preventing further rapid reworking by waves. Based on their geometry, internal pattern, and spatial distribution, these wing-shaped traces are interpreted to have been produced by relatively stationary bottom-feeding fish, morphologically similar to ancestors of Gobiidae, probably in the Actinopterygii class. The majority of fish-made trace fossils record feeding and locomotion behaviour, whereas the wing-shaped traces are interpreted to represent resting structures made by fish that pressed their fins downward on the muddy sediment surface and hence, represent undertracks. These traces are possibly the oldest fish-made structure documented thus far. These wing-shaped traces are described as Pisquiesichnus dashtgardi igen. et isp. nov. in this study.
沙特阿拉伯志留系Sharawra段(landoverian中晚期)表现出独特的表面痕迹,保存得非常好:在砂岩板上出现了7个小的翼状痕迹,显示出清晰的丘状交叉层状,其波长为几分米,被薄泥层覆盖。这些痕迹被保留为凹形的外浮雕,沿著细长的内部条纹,周围有略高的堤防。这些精致的10 - 20毫米长、10 - 10毫米宽的痕迹保存下来表明,它们是在一场风暴事件之后形成的,当时泥浆已经沉降到沉积物表面。波浪形成的沉积结构的缺乏意味着风平浪静和风暴波基底之间的沉积,这可能阻止了波浪进一步的快速改造。根据它们的几何形状、内部模式和空间分布,这些翅膀形状的痕迹被解释为由相对静止的底食性鱼类产生的,在形态上与虾蛄科的祖先相似,可能属于放线翼类。大多数鱼类留下的痕迹化石记录了进食和运动行为,而翅膀形状的痕迹被解释为鱼类在泥泞的沉积物表面向下按压鳍时留下的休息结构,因此代表了足迹。这些痕迹可能是迄今为止记载的最古老的鱼制结构。这些翅膀状的痕迹被描述为Pisquiesichnus dashgardi igen。isp。在这项研究中。
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引用次数: 0
The trace fossil Gyrolithes lorcaensis from the Lower Cretaceous of the Kopet-Dagh Basin, NE Iran 伊朗东北部Kopet-Dagh盆地下白垩世古旋石化石
IF 0.8 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-12-30 DOI: 10.1080/10420940.2021.2017921
S. Moosavizadeh, D. Knaust
Abstract The helical, vertical trace fossil Gyrolithes lorcaensis is described from marine to marginal-marine deposits of the Lower Cretaceous of the Sarcheshmeh Formation near the city Kalat-e-Naderi in the Kopet-Dagh Basin. This is the first report of Gyrolithes from Iran and thus expands the geographic range of this trace fossil. G. lorcaensis is characterized by its unique morphometric parameters, a small burrow size and whorl radius, which distinguishes it as the smallest Gyrolithes ichnospecies. The most likely producers of the Iranian Gyrolithes were crustacean decapods, which is supported by the paleoenvironment (shallow and warm water ramp) in which it is found. The function of the spiral structure in the described Gyrolithes of the Sarcheshmeh Formation can be explained as a permanent dwelling structure with the main purpose of farming bacteria via increasing the surface area of the burrow margin that allows more microbial activities. The small burrow dimensions may reflect reduced burrower body size in response to the dysoxic to anoxic conditions suggested by sedimentological evidence.
摘要:本文描述了Kopet-Dagh盆地Kalat-e-Naderi市附近下白垩世Sarcheshmeh组海相至边缘海相沉积物中螺旋形、垂直形的Gyrolithes lorcaensis化石。这是来自伊朗的首个Gyrolithes报告,从而扩大了这一痕迹化石的地理范围。G. lorcaensis以其独特的形态参数、较小的洞穴尺寸和螺旋半径为特征,使其成为最小的螺线虫鱼种。伊朗回转类动物最有可能的生产者是甲壳类十足类动物,这是由其发现的古环境(浅水和暖水斜坡)支持的。在Sarcheshmeh地层中描述的Gyrolithes螺旋结构的功能可以解释为一种永久性居住结构,其主要目的是通过增加洞穴边缘的表面积来养殖细菌,从而允许更多的微生物活动。较小的洞穴尺寸可能反映了沉积学证据表明的缺氧或缺氧条件下洞穴体型的缩小。
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引用次数: 1
A new occurrence of archosaur footprints from the Löwenstein Formation (Late Triassic, Middle Norian) of southern Germany 德国南部Löwenstein组(晚三叠世,中诺里亚)新发现的祖龙脚印
IF 0.8 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-12-29 DOI: 10.1080/10420940.2021.2017922
A. Wagensommer, S. D. Porchetti, Hans Dieter Bolter, Matteo Antonelli, Frank-Otto Haderer
Abstract Two sandstone slabs from a new Norian tetrapod tracksite in the Löwenstein Formation of southern Germany preserve a set of tracks including both tridactyl and pentadactyl ichnites, referred to theropod dinosaur and sphenosuchian crocodylomorph trackmakers, respectively. A very large manus print hints at the presence of a hitherto unknown large quadrupedal archosaur in the Norian fauna of southern Germany. The material includes one of the oldest records of crocodylomorph tracks presently known, in agreement with the skeletal record from the same formation.
在德国南部Löwenstein组一个新的挪威四足动物足迹遗址的两块砂岩板上保存了一组足迹,其中包括三足动物和五足动物,分别被称为兽脚亚目恐龙和sphenosuchian鳄鱼足迹制造者。一个非常大的手印暗示在德国南部的诺里亚动物群中存在一种迄今为止未知的大型四足祖龙。该材料包括目前已知的最古老的鳄鱼足迹记录之一,与同一地层的骨骼记录一致。
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引用次数: 0
First record of the Permian nonmarine helical trace fossil Augerinoichnus from outside of New Mexico 新墨西哥州以外发现的二叠纪非海洋螺旋形Augerinoichnus化石的首个记录
IF 0.8 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-12-21 DOI: 10.1080/10420940.2021.2017923
S. Voigt, S. Lucas, M. Raisch, T. Schindler
Abstract Augerinoichnus refers to successions of horseshoe-shaped structures interpreted as sedimentary remains of horizontal helical burrows of worm-like animals. The monospecific trace fossil was hitherto only known from early Permian intertidal to lowland coastal-plain deposits of New Mexico, U.S.A. Recent discoveries of conspecific material in early Permian nearshore lacustrine deposits of southwestern Germany represent the first record of Augerinoichnus outside of New Mexico and extend the paleogeographic and paleoenvironmental distribution of the ichnotaxon. Augerinoichnus evidentally was made in a wide array of nonmarine settings that ranged from tidal flats to inland floodplains where substrate cohesion and stability as well as stable and predictable food resources allowed the tracemaker to forage in open burrows.
Augerinoichnus是一种马蹄形结构序列,被解释为蠕虫类动物水平螺旋洞的沉积遗迹。该单种化石仅在美国新墨西哥州的早二叠世潮间带和低地海岸平原沉积物中发现。最近在德国西南部早二叠世近岸湖泊沉积物中发现的同种化石是新墨西哥州以外的第一个Augerinoichnus记录,扩展了该鱼分类群的古地理和古环境分布。显然,Augerinoichnus是在广泛的非海洋环境中形成的,从潮滩到内陆洪泛平原,在这些环境中,基质的凝聚力和稳定性以及稳定和可预测的食物资源使追踪者能够在开放的洞穴中觅食。
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引用次数: 0
Scratch circle from the Passaic Formation (Late Triassic), West-Central New Jersey 新泽西州中西部Passaic组(晚三叠世)划痕圈
IF 0.8 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-10-02 DOI: 10.1080/10420940.2021.1998038
R. Metz
ABSTRACT Scratching, forming one sharply-defined, somewhat elliptical scratch circle, surrounds a narrow-filled tube probably formed by a plant stem. Water currents carrying sediment onto a lacustrine shoreline swirled around a plant leaf, etching out a partial scratch circle. Later settling of the finest material resulted in a veneer of mud filling the scratch circle.
划痕,形成一个清晰的,有点椭圆的划痕圈,围绕着一个可能由植物茎形成的狭窄填充的管。带着沉淀物的水流在湖泊岸线上绕着一片植物叶子打转,留下了部分划痕。后来最细的材料的沉降导致泥的贴面填充划痕圈。
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引用次数: 0
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Ichnos-An International Journal for Plant and Animal Traces
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