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Sandbian (Late Ordovician) conodonts in Estonia: distribution and biostratigraphy 爱沙尼亚的山边(晚奥陶世)牙形刺:分布和生物地层学
IF 1 4区 地球科学 Q2 Earth and Planetary Sciences
Gff
Pub Date : 2022-01-02 DOI: 10.1080/11035897.2021.2020333
T. Paiste, P. Männik, T. Meidla
ABSTRACT The boundaries of the Sandbian Stage are defined by graptolites. Because of their rare occurrence in the Estonian part of the Baltoscandian palaeobasin, other fossils, including conodonts, are used for biostratigraphy as good alternatives. Since the definition of Sandbian Stage/Age in 2002, information about conodonts from this time interval in Baltsocandia has improved considerably. In this paper, existing data on Sandbian conodont faunas from the Estonian part of the Baltoscandian palaeobasin, including the most recent information from the Velise V-97 core section (western part of mainland Estonia), is evaluated. Dating of these strata in the region is addressed and partly updated. Comparison of conodont successions from other regions of Baltoscandia with those from the Estonian sections is used to check consistency of known conodont zones. The rich conodont succession from the Velise V-97 drill core is considered a representative one for the Sandbian-age carbonate ramp environment in Baltoscandia. Most of the principal conodont zones within the Sandbian Stage were recognised whilst the lower and upper boundary intervals of the stage are poorly resolved. Despite the rich conodont faunas available, the boundaries of the Sandbian Stage are currently not clearly definable in the conodont succession. However, further detailed studies on the morphology and evolution of the earliest Amorphognathus (A. inaequalis and A. tvaerensis in particular), based on rich and well preserved material from different parts of Baltoscandia, might provide useful criteria for location of the Sandbian and Katian boundaries in non-graptolitiferous carbonate sections.
摘要:沙滨阶的边界是由笔石界定的。由于它们在Baltoscandian古盆地的爱沙尼亚部分罕见,包括牙形刺在内的其他化石被用作生物地层学的良好替代品。自2002年定义Sandbian阶段/年龄以来,Baltsocndia这个时间间隔的牙形石信息有了很大的改善。本文评估了Baltoscandian古盆地爱沙尼亚部分Sandbian牙形刺动物群的现有数据,包括Velise V-97核心剖面(爱沙尼亚大陆西部)的最新信息。对该地区这些地层的年代测定进行了处理,并进行了部分更新。将Baltoscandia其他地区的牙形石序列与爱沙尼亚剖面的牙形岩序列进行比较,以检查已知牙形石带的一致性。Velise V-97岩芯中丰富的牙形刺序列被认为是Baltoscandia Sandbian时代碳酸盐岩斜坡环境的代表性序列。Sandbian阶内的大多数主要牙形刺带已被识别,而该阶的下边界层段和上边界层段的分辨率较差。尽管有丰富的牙形石动物群,但目前在牙形石序列中尚不能明确界定沙滨阶的边界。然而,基于Baltoscandia不同地区丰富且保存完好的材料,对最早的无齿龙(特别是A.inaequip和A.tvaerensis)的形态和进化进行进一步的详细研究,可能会为非笔石碳酸盐岩剖面中Sandbian和Katian边界的定位提供有用的标准。
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引用次数: 1
First record of an Early Ordovician brachiopod and conodont fauna from Lapland, Sweden 瑞典拉普兰早奥陶世腕足类和牙形石动物群的首次记录
IF 1 4区 地球科学 Q2 Earth and Planetary Sciences
Gff
Pub Date : 2022-01-02 DOI: 10.1080/11035897.2022.2032822
M. Streng, J. A. Rasmussen, J. R. Ebbestad, Thomas Weidner
ABSTRACT An Early Ordovician fauna of linguliform brachiopods and euconodonts is described from the Alum Shale Formation in northernmost Västerbotten County, northern Sweden. The fauna was recovered from a single carbonate concretion collected at an exposure near the northern shore of Lake Storvindeln. Despite of the fragmentary and tectonized preservation of the obtained fossils, at least eight different taxa of linguliform brachiopods and four species of euconodonts could be identified. Among the brachiopods, the ephippelasmatid Pomeraniotreta biernatae Bednarczyk, 1986 is most common; it is associated among others with the acrotretids Eurytreta cf. sabrinae (Callaway, 1877) and Ottenbyella sp., and a spinose specimen potentially representing the zhanatellid Thysanotos. Two of the brachiopod taxa appear to present new species, Pomeraniotreta n. sp. and Mytoella? n. sp., but additional material would be needed for a formal description. Euconodont specimens are rare and comprise coniform elements of Drepanodus arcuatus Pander, 1856, Drepanoistodus aff. amoenus (Lindström, 1955) sensu Löfgren (1994), Paroistodus numarcuatus (Lindström, 1955) and Rossodus aff. manitouensis Repetski & Ethington, 1983. Evaluation of the stratigraphic distribution of the different taxa of brachiopods and euconodonts under consideration of the local geology, suggests that the investigated sample represents a level in the upper part of the Tremadocian (Tr2) part of the Alum Shale Formation. The fauna represents the northernmost occurrence of age diagnostic Early Ordovician (Tremadocian) fossils in Sweden.
摘要/ ABSTRACT摘要:在瑞典北部Västerbotten县最北端的明矾页岩组中,描述了早奥陶世的舌形腕足动物和真牙形动物。动物群是在Storvindeln湖北岸附近的一个暴露处收集的单一碳酸盐固结物中恢复的。尽管所获得的化石碎片化和构造化保存,但至少可以识别出8个不同的舌形腕足动物分类群和4种真牙形动物。在腕足动物中,最常见的是ephippelasmatid Pomeraniotreta biernatae Bednarczyk, 1986;它与肢肢类Eurytreta cf. sabrinae (Callaway, 1877)和Ottenbyella sp.以及可能代表zhanatellid Thysanotos的棘类标本有关联。两个腕足动物分类群似乎出现了新的物种,Pomeraniotreta n. sp.和Mytoella?N. sp.,但要作正式说明还需要额外的材料。真牙形刺标本是罕见的,包括弓形锥齿(1856年),弓形锥齿(Lindström, 1955年),弓形锥齿(Löfgren, 1994年),numarcuatus (Lindström, 1955年)和Rossodus aff.manitouensis Repetski & Ethington, 1983年)。结合当地地质条件,对腕足类和真牙形刺类不同分类群的地层分布进行了评价,认为所调查的样品代表了明质页岩组Tremadocian (Tr2)段上部的一个水平。该动物群代表了瑞典早奥陶世(Tremadocian)化石最北端的年龄诊断。
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引用次数: 0
Has the tectonic regime of the Baltic Shield always remained the same? 波罗的海地盾的构造机制一直保持不变吗?
IF 1 4区 地球科学 Q2 Earth and Planetary Sciences
Gff
Pub Date : 2022-01-02 DOI: 10.1080/11035897.2021.1993453
A. Lindh
ABSTRACT Recent theories on the growth of continental crust stress its formation in pre-Archean and Archean times with minor additions at later times. The Earth has gone through a unidirectional evolution including surface changes, the genesis of life and an important loss of energy. The energy loss drives tectonic processes, but at a rate declining with available energy and thus with time. Semiquantitative modelling of energy-changes and transport indicates important differences between today’s Earth and the Paleoproterozoic Earth. Actualistic but non-uniformitarian arguments suggest that principal tectonic processes have changed over time. The Baltic Shield displays a fundamental difference in conditions between the c. 0.9 Ga high-pressure metamorphism including the occurrence of eclogite in the western Sveconorwegian Orogen and c. 2 Ga low-pressure metamorphism in the eastern Svecokarelian Orogen. This suggests that modern platetectonic processes were operative at c. 0.9 Ga but leaves the question whether it was active during the Svecokarelian orogeny open. The lack of eclogite in the Svecokarelian Orogen is explained with a change with time of the geotherm slope. The occurrence of two generations of granitic rocks of different compositions in the Svecokarelian is easily explained by an increased mantle heat flow in a stratified continental crust. In contrast to calc-alkaline platetectonic compositions, alkali-calcic rocks occur in a quarter-circle around the Svecokarelian core. The question arises whether modern plate-processes successively replaced proto-plate processes during Paleo- and Mesoproterozoic times.
最近关于大陆地壳生长的理论强调其形成于前太古宙和太古宙,后期略有增加。地球经历了单向进化,包括表面变化、生命起源和重要的能量损失。能量损失推动了构造过程,但随着可用能量的增加,从而随着时间的推移,能量损失呈下降趋势。对能量变化和传输的半定量建模表明了今天的地球和古元古代地球之间的重要差异。现实主义但非统一主义的论点表明,主要的构造过程随着时间的推移而发生了变化。波罗的海地盾表现出0.9Ga高压变质作用(包括Svecoorwegian造山带西部榴辉岩的出现)和2.Ga低压变质作用(Svecokarelian造山带东部)之间的基本条件差异。这表明现代板块构造过程在约0.9 Ga时是有效的,但它在Svecokarelian造山运动期间是否活跃的问题仍然存在。用地热坡随时间的变化解释了Svecokarelian造山带榴辉岩的缺乏。Svecokarelian中两代不同成分的花岗岩的出现很容易用层状大陆地壳中地幔热流的增加来解释。与钙碱性板块构造成分相反,碱性钙质岩石出现在Svecokarelian核心周围的四分之一圈中。问题是,在古元古代和中元古代,现代板块过程是否相继取代了原始板块过程。
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引用次数: 0
Two trepostome bryozoans from the Assistance Formation (Permian, Roadian) near Lake Hazen, Ellesmere Island, Canada 加拿大埃尔斯米尔岛黑森湖附近援助组(二叠纪,Roadian)的两种环口苔藓虫
IF 1 4区 地球科学 Q2 Earth and Planetary Sciences
Gff
Pub Date : 2021-11-17 DOI: 10.1080/11035897.2021.1989716
A. Ernst, H. A. Nakrem
ABSTRACT Rhombotrypella superangustata and Dyscritellina fuglensis (Bryozoa, Trepostomata) were found for the first time in the Assistance Formation of Ellesmere Island, Canada. Both species correlate the Roadian age of the Assistance Formation with contemporary deposits in Northern Russia and Bjørnøya, Norway. They possess robust erect colonies adapted for shallow shelf conditions with moderate energy depositional environment with low sedimentation rate and hard substrate.
摘要在加拿大埃尔斯米尔岛的辅助群中首次发现了超鳗形锥虫(Rhombotrypella superangustata)和富格丽锥虫(Dyscritellina fuglensis)。这两个物种都将辅助组的路迪亚时代与俄罗斯北部和挪威Bjørnøya的当代矿床联系起来。它们具有强壮的直立菌落,适合于浅海陆架条件,中等能量沉积环境,低沉积速率和硬基质。
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引用次数: 1
Holdfasts of Sphenothallus (Cnidaria) from the early Silurian of western North Greenland (Laurentia) 北格陵兰岛西部(劳伦西亚)志留纪早期Sphenothallus(刺胞目)的巢壳
IF 1 4区 地球科学 Q2 Earth and Planetary Sciences
Gff
Pub Date : 2021-10-02 DOI: 10.1080/11035897.2021.1979642
J. S. Peel
ABSTRACT Well-preserved holdfasts and early growth stages of the tubular cnidarian Sphenothallus are described from the early Silurian (Llandovery Series) of Washington Land, North Greenland. The diameter of the holdfast is determined by the formation of a basal disc attached to the substrate, upon which the initially conical, but subsequently tubular shell is formed. The characteristic opposing longitudinal thickenings are developed at each angular margin as the initially circular tube acquires an elliptical form.
摘要描述了北格陵兰华盛顿地志留纪早期(Llandovery系列)管状cnidarian鞘翅目化石保存完好的固着物和早期生长阶段。固定器的直径由连接到基底的基底盘的形成来确定,在基底盘上形成最初的圆锥形但随后的管状壳体。当最初的圆形管获得椭圆形形状时,在每个角边缘处形成了特征性的相对纵向增厚。
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引用次数: 5
Cleaning up the record – revised U-Pb zircon ages and new Hf isotope data from southern Sweden 整理瑞典南部记录修正的U-Pb锆石年龄和新的Hf同位素数据
IF 1 4区 地球科学 Q2 Earth and Planetary Sciences
Gff
Pub Date : 2021-10-02 DOI: 10.1080/11035897.2021.1939777
Å. Johansson
ABSTRACT Ten samples of felsic plutonic rocks from the Eastern Segment of the Sveconorwegian Orogen in southern Sweden, previously dated by ID-TIMS on zircon, have been dated anew using SIMS spot analysis of individual zircon grains, leading to more reliable and in most cases also more precise revised magmatic crystallization ages. A gneissic monzonite within the Protogine Zone in Småland yields a revised U-Pb age of ca. 1725 Ma, four samples of orthogneiss from Skåne all yield revised ages between 1700 and 1690 Ma, while two samples of coarse-grained granitic gneiss in the same region yield ages between 1690 and 1680 Ma. These revised ages are between 15 and 250 m.y. older than previously obtained TIMS ages. Two samples of the Gumlösa-Glimåkra granite along the Protogine Zone in northern Skåne and one sample of related syenite yield ages around 1220 Ma, similar but more precise compared to the previous ages. The U-Pb zircon data have been complemented by Hf isotope analysis by LA-ICP-MS on the same grains, and previously obtained initial Sr and Nd whole-rock isotope data have been recalculated to the revised crystallization ages. The Sr isotope data scatter, while the revised initial εNd values fall between +1 and +2 for the older rocks, and close to 0 for the younger intrusives along the Protogine Zone. Initial εHf in magmatic undisturbed zircons shows relatively little spread within each sample, between 2 and 4 Epsilon units, disregarding a few outliers, with average values for the 1725 to 1680 Ma rocks between +3 and +5.5 and for the 1220 Ma rocks at ca. +1.5. Covariation between initial εNd and initial εHf in the older rocks suggests either mixing between two isotopically distinct magma sources or one magma source which was isotopically heterogeneous. The isotopic signatures of the 1220 Ma intrusive rocks along the Protogine Zone are indicative of juvenile mantle input to their magmas, rather than pure crustal melting.
来自瑞典南部斯韦诺威造山带东段的10个长英质深成岩样品,之前用锆石ID-TIMS定年,现在利用单个锆石颗粒的SIMS斑点分析重新定年,从而得到更可靠的、在大多数情况下也更精确的修正岩浆结晶年龄。sm原生带的一块片麻岩二长岩测得的U-Pb年龄约为1725 Ma, sk地区的4块正长岩样品测得的U-Pb年龄均在1700 ~ 1690 Ma之间,同一地区的2块粗粒花岗质片麻岩样品测得的U-Pb年龄在1690 ~ 1680 Ma之间。这些修订后的年龄比以前获得的TIMS年龄大15至250岁。两个沿斯克纳北部原生带的Gumlösa-Glimåkra花岗岩样本和一个相关正长岩屈服年龄在1220 Ma左右的样本,与之前的年龄相似但更精确。利用LA-ICP-MS对同一颗粒进行Hf同位素分析,补充了U-Pb锆石数据,并对先前获得的Sr和Nd全岩初始同位素数据进行了重新计算,以修正结晶年龄。Sr同位素数据较为分散,古岩体修正后的初始εNd值在+1 ~ +2之间,而沿原生带的新岩体修正后的初始εNd值接近于0。未扰动岩浆锆石的初始εHf分布相对较小,在2 ~ 4 Epsilon单位之间,不考虑少数异常值,1725 ~ 1680 Ma岩石的平均值在+3 ~ +5.5之间,1220 Ma岩石的平均值约为+1.5。古岩石中初始εNd和初始εHf的协变表明两个同位素差异明显的岩浆源或一个同位素不均匀的岩浆源之间存在混合。原生带1220 Ma侵入岩的同位素特征表明其岩浆是由幼年地幔输入的,而不是单纯的地壳熔融作用。
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引用次数: 1
Upper Darriwilian (Middle Ordovician) graptolite biostratigraphy and correlation of the Krapperup drill core, Scania, Sweden 瑞典斯堪的纳维亚克拉珀普岩芯的上达里维里阶(中奥陶世)笔石生物地层学和对比
IF 1 4区 地球科学 Q2 Earth and Planetary Sciences
Gff
Pub Date : 2021-09-13 DOI: 10.1080/11035897.2021.1936156
J. Maletz, P. Ahlberg
ABSTRACT The Krapperup drill core from Scania (Skåne), southern Sweden, includes one of the most important graptolitic Darriwilian (upper Middle Ordovician) successions of Baltica. Only some intervals have been documented previously and especially the uppermost part of the succession has been completely unknown. A fairly complete succession of mid- to late Darriwilian age provides insight into the early evolution of the axonophoran (biserial) graptolites and demonstrates a distinct biogeographical faunal differentiation during this time. Graptolites from the mid-Darriwilian Holmograptus lentus and Nicholsonograptus fasciculatus biozones are discussed and important specimens and species from the Krapperup drill core are illustrated. Ekstroemograptus inexpectatus n. gen., n. sp. from the Holmograptus lentus Biozone is described as new.
瑞典南部Scania (sk ne)的Krapperup岩心包含了波罗的海最重要的笔石系Darriwilian(中奥陶统上)序列之一。以前只有一些间隔被记录下来,特别是演替的最上面部分是完全未知的。一个相当完整的中晚期达里威廉时代序列,提供了对轴索笔石(双序列)早期进化的深入了解,并证明了这一时期明显的生物地理区系分化。讨论了中darriwilian Holmograptus lentus和Nicholsonograptus fasciculatus生物带的笔石,并举例说明了Krapperup岩心的重要标本和种。来自lentus Holmograptus的Ekstroemograptus inexpectatus n. gen., n. sp.被描述为新种。
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引用次数: 1
Authigenic calcium carbonate precipitation in the “bathtub ring” around the anoxic Alum Shale Basin during the Furongian SPICE event (Baltic Basin, northern Poland) 富隆期SPICE事件(波兰北部波罗的海盆地)缺氧明矾页岩盆地周围“浴缸环”自生碳酸钙沉淀
IF 1 4区 地球科学 Q2 Earth and Planetary Sciences
Gff
Pub Date : 2021-09-08 DOI: 10.1080/11035897.2021.1941239
Aleksander Majchrzyk, W. Kozłowski, A. Żylińska
ABSTRACT The precipitation of both biotic and abiotic calcium carbonate is of great importance in modern and ancient global biogeochemical cycles. In the present-day oceans, the widespread precipitation of inorganic CaCO3 on the seafloor or in the water column is possible only under extraordinary circumstances. By contrast, in the geological record, authigenic seafloor carbonate cements were widespread in the supersaturated, anoxic oceans of the Precambrian. Widespread authigenic carbonate precipitation ceased by the end of the Neoproterozoic as a consequence of global oceanic oxygenation: in the Phanerozoic, it occurred only during major anoxic events (for example, at the Permian/Triassic boundary) or in restricted, stagnant basins. Here, we present an anomalous record of CaCO3 precipitation from the Cambrian Alum Shale Basin of the Baltica palaeocontinent with Precambrian-like authigenic, seafloor encrusting, crystalline carbonates. The depositional environment of this well-recognized, cool-water, stagnant anoxic basin favoured local carbonate precipitation via the surplus generation of alkalinity in anoxic bottom waters. However, the correlation of the acme of authigenic carbonate formation with the onset and peak of the Steptoean Positive Carbon Isotope Excursion (SPICE) suggests a driver−trigger relation between the two phenomena. On a smaller scale, carbonate authigenic precipitation is manifested by a cement-supported texture of the limestone intercalations in the Alum Shale facies. Instantaneous calcification of the faunal remains, termed here the Snedronningen phenomenon, must have been of great importance in the formation of Orsten-type Konservat Lagerstätten.
生物和非生物碳酸钙的沉淀在现代和古代全球生物地球化学循环中都具有重要意义。在当今的海洋中,无机物CaCO3在海底或水柱上的广泛沉淀只有在特殊情况下才有可能。相比之下,在地质记录中,自生的海底碳酸盐胶结物广泛存在于前寒武纪过饱和、缺氧的海洋中。由于全球海洋的氧化作用,广泛的自生碳酸盐降水在新元古代末期停止了;在显生宙,它只发生在主要的缺氧事件期间(例如,在二叠纪/三叠纪的边界)或在有限的停滞盆地中。本文对波罗的海古大陆寒武纪明矾页岩盆地的CaCO3降水进行了异常记录,该盆地具有前寒武纪样自生、海底覆壳、结晶碳酸盐。这个众所周知的、冷水、停滞的缺氧盆地的沉积环境有利于通过缺氧底部水的剩余碱度产生局部碳酸盐降水。然而,自生碳酸盐形成的顶峰与阶梯正碳同位素偏移(SPICE)的开始和峰值的相关性表明,这两种现象之间存在驱动-触发关系。在较小的尺度上,碳酸盐自生降水表现为明矾页岩相灰岩夹层的水泥支撑结构。动物遗骸的瞬时钙化,在这里被称为Snedronningen现象,在orsten型Konservat Lagerstätten的形成中一定是非常重要的。
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引用次数: 0
Rock fracturing by subglacial hydraulic jacking in basement rocks, eastern Sweden: the role of beam failure 瑞典东部基岩冰下液压顶进岩石破裂:梁破坏的作用
IF 1 4区 地球科学 Q2 Earth and Planetary Sciences
Gff
Pub Date : 2021-07-15 DOI: 10.1080/11035897.2021.1939776
M. Krabbendam, Romesh Palamakumbura, C. Arnhardt, A. Hall
ABSTRACT Dense networks of dilated fractures occur locally in the upper 5–15 m of bedrock in basement gneisses in eastern Sweden. Near Forsmark, pre-existing sub-horizontal fractures have been jacked open and filled with water-lain sediment, likely during the latest Weichselian glaciation. Despite extensive previous research, it is uncertain whether subglacial hydraulic jacking led to the generation of new fractures, in addition to reactivation of pre-existing ones. Re-analysis of historic photos from excavations near the Forsmark power plant indicates formation of two types of new fracture. Firstly, rock fragments were broken off the main fracture surfaces as existing fractures were jacked open. Secondly, fracture analysis shows that whilst few subvertical fractures occur above tight sub-horizontal fractures, a higher density of vertical fractures occurs above dilated sub-horizontal fractures, suggesting new formation. We apply a model of beam failure theory, borrowed from structural engineering, to constrain potential new fracture generation, using assumptions based on measured water pressure fluctuations from beneath the Greenland Ice Sheet. This modelling shows that beam failure is a plausible mechanism for the generation of new vertical fractures during a subglacial water fluctuation cycle under a range of realistic glaciological conditions. This implies that hydraulic jacking can result in further in situ disruption and brecciation of the shallow rock mass, decreasing the rock mass strength and increasing its hydraulic conductivity. Altogether, hydraulic jacking of existing fractures and the formation of new vertical fractures results in effective subglacial mechanical weathering of the shallow rock mass.
摘要:在瑞典东部基底片麻岩的基岩上部5–15m,局部出现密集的扩张裂缝网络。在Forsmark附近,先前存在的亚水平裂缝被顶开,并充满了水沉积,很可能是在最近的韦氏冰川作用期间。尽管之前进行了广泛的研究,但尚不确定冰下液压顶进是否会导致新裂缝的产生,以及原有裂缝的重新激活。对福斯马克发电厂附近挖掘的历史照片的重新分析表明,形成了两种类型的新裂缝。首先,当现有裂缝被顶开时,岩石碎片从主要裂缝表面破碎。其次,裂缝分析表明,虽然致密亚水平裂缝上方几乎没有垂直裂缝,但扩张亚水平裂缝之上出现了更高密度的垂直裂缝,这表明存在新的地层。我们借鉴结构工程中的梁破坏理论模型,使用基于格陵兰冰盖下实测水压波动的假设,来约束潜在的新裂缝生成。该模型表明,在一系列现实的冰川条件下,在冰下水波动周期中,梁破坏是产生新的垂直裂缝的一种合理机制。这意味着液压顶进会导致浅层岩体的进一步原位破坏和角砾化,降低岩体强度并增加其导水性。总之,现有裂缝的水力顶进和新的垂直裂缝的形成导致了浅层岩体的有效冰下机械风化。
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引用次数: 5
Searching for a nearest living equivalent for Bennettitales: a promising extinct plant group for stomatal proxy reconstructions of Mesozoic pCO2 寻找与Bennettitales相近的生物:一个有希望用于中生代pCO2气孔代用重建的已灭绝植物群
IF 1 4区 地球科学 Q2 Earth and Planetary Sciences
Gff
Pub Date : 2021-07-03 DOI: 10.1080/11035897.2021.1895304
M. Steinthorsdottir, C. Elliott‐Kingston, Mario Coiro, J. McElwain
ABSTRACT To understand Earth´s climate variability and improve predictions of future climate change, studying past climates is an important avenue to explore. A previously published record of pCO2, across the Triassic–Jurassic boundary (TJB, ~201 Ma) of East Greenland, showed that Bennettitales (Anamozamites and Pterophyllum) responded in parallel to the empirically proven pCO2-responders Ginkgoales, reducing their stomatal densities by half across the TJB, indicating a transient doubling of pCO2. The abundance of fossil Bennettitales in Mesozoic strata and natural history museum collections worldwide offers enormous potential for further stomatal proxy pCO2 reconstructions, but a suitable nearest living equivalent (NLE) should ideally first be identified for this extinct plant group. Using specimens from herbarium collections, three species of cycads, historically considered the best NLE, were tested for pCO2 response, as well as two species of tree ferns, grown in experimental growth chambers. None responded to changes in pCO2, and were consequently rejected as NLEs. Finally, two species of ferns were selected from the literature, and produced very similar pCO2 compared to Ginkgoales. However, these understory ferns are not appropriate NLEs for Bennettitales due to differences in habitat and a distant evolutionary relationship. Future work should test additional plant groups, in particular seed plants such as basal angiosperms and Gnetales, for suitability as NLE for Bennettitales in pCO2 reconstructions, for example through biogeochemical fingerprinting using infrared microspectroscopy. Until an appropriate NLE is identified, Bennettitales pCO2 can be reconstructed based on cross-calibration of stomatal densities with those of co-occurring pCO2 responders, such as Ginkgoales.
摘要为了了解地球的气候变化,提高对未来气候变化的预测,研究过去的气候是一条重要的探索途径。先前发表的一份关于东格陵兰三叠纪-侏罗纪边界(TJB,约201 Ma)pCO2的记录显示,Bennettitales(Anamozamites和Pterophyllum)与经验证明的pCO2响应Ginkgoales平行响应,在整个TJB将其气孔密度降低了一半,表明pCO2瞬时翻倍。世界各地中生代地层和自然历史博物馆收藏的Bennettitales化石为进一步重建气孔代理pCO2提供了巨大的潜力,但理想情况下,应该首先为这个已灭绝的植物群确定一个合适的最接近生命的等效物(NLE)。使用植物标本馆收集的标本,对三种苏铁(历史上被认为是最好的NLE)以及两种在实验生长室中生长的树蕨进行了pCO2反应测试。没有人对pCO2的变化做出反应,因此被拒绝为NLE。最后,从文献中选择了两种蕨类植物,与银杏叶相比,它们产生了非常相似的pCO2。然而,由于栖息地的差异和遥远的进化关系,这些林下蕨类植物不适合Bennettitales。未来的工作应该测试其他植物群,特别是种子植物,如基础被子植物和Gnetales,在pCO2重建中是否适合作为Bennettitales的NLE,例如通过使用红外显微光谱的生物地球化学指纹。在确定合适的NLE之前,Bennettitales pCO2可以基于气孔密度与共存pCO2应答者(如Ginkgoales)的气孔密度的交叉校准来重建。
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