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The Early Pliocene small mammals (Eulipotyphla, Rodentia, Lagomorpha) from Berești and Mălușteni (eastern Romania): a fresh look at old collections 来自Berești和Mălușteni(罗马尼亚东部)的上新世早期小型哺乳动物(真兽门,啮齿目,狐齿目):对旧藏品的新观察
IF 1.2 4区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2023-01-09 DOI: 10.1017/s175569102200024x
V. Crespo, Ș. Vasile, A. Petculescu, B. Rățoi, Bogdan S. Haiduc
The neighbouring sites of Berești and Mălușteni (Eastern Carpathian Foreland, eastern Romania) have yielded the most abundant and taxonomically diverse Pliocene vertebrate assemblages described so far from the entire country. Some of the small mammals found here were described as new taxa, and occasionally reassessed during the past one hundred years, but most of the material collected initially remained unrevised. Here, we provide a taxonomic revision of all the small mammal material (insectivores, rodents, and lagomorphs) that could be found in three main collections. The studied specimens were assigned to the insectivore families Desmanidae (Desmana verestchagini and Talpa sp.), and Erinaceidae (Erinaceus sp.); to the rodent families Muridae (Mimomys sp. or Promimomys sp.; Allocricetus sp.), Sciuridae (Spermophilus cf. nogaici), Spalacidae (Pliospalax macoveii), and Castoridae (Trogontherium minus, Castor fiber); and to the lagomorph families Leporidae (Trischizolagus dumitrescuae) and Ochotonidae (Ochotona ursui). Compared to the faunal assemblages described from Central-Eastern Europe, the identified taxa (some confirmed, others reassessed as synonyms) support an Early Pliocene age for the vertebrate assemblages from Berești and Mălușteni. Both faunal assemblages are assigned to the Ruscinian, with the faunas from Berești being considered geologically slightly older than the ones from Mălușteni.
邻近的Berești和Mălușteni(东喀尔巴阡前陆,罗马尼亚东部)发现了迄今为止在全国范围内描述的最丰富和分类最多样化的上新世脊椎动物组合。在这里发现的一些小型哺乳动物被描述为新的分类群,并且在过去的一百年中偶尔被重新评估,但最初收集的大多数材料仍然没有修改。在这里,我们提供了所有的小哺乳动物材料(食虫动物,啮齿动物和狐形动物),可以发现在三个主要收藏的分类修订。研究标本归属于食虫昆虫蛛科(Desmana verestchagini和Talpa sp.)和蚕科(Erinaceus sp.);到鼠科(Mimomys sp.或promomys sp.);异环蝇属(Allocricetus sp.)、麻蝇科(sperophilus cf. nogaici)、麻蝇科(Pliospalax macoveii)和蓖麻科(Trogontherium minus,蓖麻纤维);以及狐蝠科(狐蝠科)和狐蝠科(熊狐蝠科)。与中欧-东欧的动物群相比,已确定的分类群(一些得到确认,另一些被重新评估为同义词)支持Berești和Mălușteni的脊椎动物群的上新世早期。这两个动物群都属于俄罗斯,Berești的动物群在地质上被认为比Mălușteni的动物群要古老一些。
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引用次数: 1
Review of the Ordovician pelagic trilobite Ellipsotaphrus (Cyclopygoidea, Ellipsotaphridae) and its allies, with new discoveries from Girvan, Ayrshire 奥陶纪浮游三叶虫椭圆虫属(环盖虫亚目,椭圆虫科)及其盟友的研究综述,以及艾尔郡吉尔万的新发现
IF 1.2 4区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-12-01 DOI: 10.1017/S1755691022000263
J. Ingham, R. Fortey
ABSTRACT Following Fortey and Owens (1987), the Ordovician trilobite taxon Ellipsotaphrinae is established as valid, but is argued to have full family status within the Cyclopygoidea. It encompasses the existing genera Ellipsotaphrus, Girvanopyge Gamops and Circulocrania together with two new genera proposed herein, Arisemolobes and Synaptotaphrus. Typical ellipsotaphrid genera have a totally circumscribed ‘foreglabella’, incorporating extended S1 furrows and a portion of the occipital furrow. The known range of the family is Floian to Katian. Genera are conservative in form throughout their ranges and are widespread. All occur only in deeper water sediments with palaeooceanic access. Ellipsotaphrus monophthalmus and Ellipsotaphrus infaustus are reassessed and Ellipsotaphrus zhongguoensis, from the Katian of China, is regarded as a junior synonym of the Katian Girvan species Ellipsotaphrus pumilio. Girvanopyge [ = Cremastoglottos; Nanlingia; Waldminia] is demonstrated, partly on the basis of new material from the Katian of Girvan, to be an ellipsotaphrid cyclopygoid and not to have a close affinity to the remopleuridids, as had been clained. Girvanopyge barrandei, from the Katian of the Czech Republic, is synonymised with Girvanopyge caudata from China. Gamops is revived for forms showing a relationship to both Girvanopyge and Ellipsotaphrus. It encompasses three Czech species including the Dapingian Gamops triangulatus, which probably also occurs in correlative strata in South Wales. The systematic treatment is supported by new material from the Upper Ordovician of the Girvan district, and the relevant geology of this area is described in detail. New species proposed are: Arisemolobes zhouzhiyii, Synaptotaphrus oarion and Circulocrania ? dichaulax.
摘要继Fortey和Owens(1987)之后,奥陶纪三叶虫分类单元Ellipsotaphrinae被确定为有效分类单元,但被认为在环盖类中具有完整的家族地位。它包括现有的椭圆虫属、Girvanopyge Gamops属和Circulcrania属,以及本文提出的两个新属,Arisemolobes属和Synaptotaphrus属。典型的椭圆顶属有一个完全外切的“额眉间”,包括延伸的S1沟和一部分枕沟。已知的家族范围是Floian到Katian。Genera在其整个范围内的形式都是保守的,并且分布广泛。所有这些都只出现在具有古海洋通道的深水沉积物中。对中国加添地区的单眼椭圆虫和隐眼椭圆虫进行了重新评价,认为中果椭圆虫是加添Girvan种小椭圆虫的初级异名。Girvanopyge[=Cremastoglotos;Nanlingia;Waldminia]在一定程度上基于Girvan的Katian的新材料,被证明是一种椭圆形的环状体,而不是像人们所声称的那样与再胸膜体具有密切的亲和力。来自捷克共和国卡蒂亚的巴兰代Girvanopyge与来自中国的尾状Girvanopyge同义。Gamops因表现出与Girvanopyge和Ellipsotaphrus的关系而复活。它包括三个捷克物种,包括大平阶三角Gamops triangulatus,它可能也出现在南威尔士的相关地层中。Girvan地区上奥陶统的新材料支持了该系统的处理,并详细描述了该地区的相关地质情况。拟引进的新品种有:周志毅Ariemolobes zhouzhiyii、合龙Synaptotaphrus oarion和Circulcrania?二壳虫。
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引用次数: 0
TRE volume 113 issue 4 Cover and Back matter TRE第113卷第4期封面和封底
IF 1.2 4区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-12-01 DOI: 10.1017/s1755691023000129
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引用次数: 0
TRE volume 113 issue 4 Cover and Front matter TRE第113卷第4期封面和封面
IF 1.2 4区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-12-01 DOI: 10.1017/s1755691023000117
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引用次数: 0
Palaeo-climate and -topography of the continental orogen: Theoretical inversion with initial oxygen isotopes of ancient meteoric water 大陆造山带的古气候和地形:利用古代大气降水的初始氧同位素进行理论反演
IF 1.2 4区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-12-01 DOI: 10.1017/S1755691023000075
Chun-Sheng Wei, Zi‐Fu Zhao
ABSTRACT Ancient environments have been mostly reconstructed with exogenous records, yet the potential constraints from endogenous archives were less emphasised. It has been well known that the outer- and inner-spheres of the planetary Earth are naturally linked and/or interplayed each other among geospheres. As stable isotopes of the meteoric water are globally dependent upon precipitating environments, rocks and/or minerals hydrothermally altered by the meteoric water can thus imprint environmental information of continental settings. These valuable clues, however, have been intuitively and/or qualitatively inferred up to now. On the basis of an innovative procedure recently proposed for dealing with thermodynamic re-equilibration of oxygen isotopes between constituent minerals and water from fossil hydrothermal systems, ancient meteoric waters are theoretically inverted from the early Cretaceous post-collisional granitoid and Triassic gneissic country rocks across the Dabie orogen in central-eastern China. The initial oxygen isotopes of ancient meteoric water (i.e., $delta ^{18}O_W^i$ value hereafter) range from −11.01 ± 0.43 (one standard deviation, 1SD) to −7.61 ± 0.07‰ in this study, yet systematically and/or statistically deviating from modern local precipitation. These imply that either palaeoclimate could be colder than the present at least during the early Cretaceous or palaeoaltimetry has geographically varied across the Dabie orogen since the Triassic. Moreover, the lifetime of fossil hydrothermal systems is kinetically quantified to less than 1.2 million years (Myr) for the concurrent lowering of oxygen isotopes of hydrothermally altered rock-forming minerals through the surface-reaction oxygen exchange with ancient meteoric waters herein. Our results thus suggest that palaeoenvironments of the continental orogen can be scientifically and methodologically unearthed from endogenous archives and theoretical inversion of $delta ^{18}O_W^i$ values can be quantitatively applied beyond the Dabie orogen.
摘要古代环境大多是通过外源性记录重建的,但内源性档案的潜在约束却很少得到重视。众所周知,地球的外层和内层在地球圈之间自然相连和/或相互作用。由于大气降水的稳定同位素在全球范围内取决于沉淀环境,因此,被大气降水水热蚀变的岩石和/或矿物可以留下大陆环境的环境信息。然而,到目前为止,这些有价值的线索已经得到了直观和/或定性的推断。在最近提出的一种创新程序的基础上,处理组成矿物和化石热液系统水之间氧同位素的热力学再平衡,从理论上讲,古代大气降水是从中国中东部大别造山带上白垩纪早期碰撞后花岗质岩石和三叠纪片麻质围岩反演而来的。古代大气降水的初始氧同位素(即$delta^{18}O_W^i$值)在−11.01±0.43(一个标准差,1SD)到−7.61±0.07‰之间,但系统地和/或统计地偏离了现代局部降水。这意味着,至少在白垩纪早期,古气候可能比现在更冷,或者自三叠纪以来,大别造山带的古海拔测量在地理上发生了变化。此外,由于通过与古代大气降水的表面反应氧交换,水热蚀变岩石形成矿物的氧同位素同时降低,化石热液系统的寿命在动力学上被量化为小于120万年(Myr)。因此,我们的研究结果表明,大陆造山带的古环境可以从内生档案和$delta的理论反演中科学和方法地发掘出来^{18}O_W^i$值可以定量地应用于大别造山带以外的地区。
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引用次数: 1
Byronids and similar tubular fossils from the Devonian of the Barrandian area (Czech Republic) 巴兰地地区(捷克共和国)泥盆纪拜伦龙和类似管状化石
IF 1.2 4区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-12-01 DOI: 10.1017/S1755691023000099
M. Mergl, P. Kraft
ABSTRACT Byronids are problematic fossils of possible cnidarian affinity, only rarely reported from the Devonian, but never previously studied in the Barrandian area, Czech Republic. Two new species Prestephanoscyphus branzovensis sp. nov. and Prestephanoscyphus robustus sp. nov. are erected from the Lochkovian and the Eifelian, respectively. Four poorly known species referred to Byronia are described in open nomenclature from the Emsian and Eifelian. The new genus Parabyronia gen. nov. with the type species Parabyronia elegans sp. nov. is closely related to other sphenothallids but is distinguished by transverse ridges on its theca and short spines at the apertural end of the theca. Definite identification of phosphatic rings as the attachment structure of byronids was proved by new material of the Lochkovian age. A dwarf conularia Pidiconularia gen. nov. with the type species Pidiconularia tubulata sp. nov. is remarkable by its minute size and very fine ornament; its conulariid affinity is proved by subrectangular cross-section and four internal carinae. Microstructure of theca of Prestephanoscyphus is characterised by alternation of compact laminae of aligned columnar microcrystallites and chambered laminae with isometric microcrystallites of apatite. Accretionary growth of byronid theca and structure of the holdfast with basal opening for the adhesive pedal disc support their cnidarian affinity although they likely display the bilateral instead of tetramerous symmetry. The protective function of bilaterally symmetrical whorls of internal apophyses in Prestephanoscyphus is suggested and their role in strengthening of thecal wall or supporting function of gastric septa are disputed. The byronids are regarded as epibionts rather than attached to the rocks. They are representatives of benthic groups with phosphatic shells that declined with decreasing availability of phosphorus in seas and oceans.
摘要Byronids是一种可能与cnidarian亲缘关系有问题的化石,在泥盆纪很少有报道,但以前从未在捷克共和国的Barrandian地区进行过研究。两个新种Prestephanoscyphus branzovensis sp.nov.和Prestephanos cyphus robustus sp.nov..分别从Lochkovian和Eifelian建立。四个鲜为人知的物种被称为Byronia,以埃姆西亚和艾菲尔亚的开放命名法进行了描述。新属Parapyronia gen.nov.和模式种Parapyronnia elegans sp.nov.与其他蝶形体密切相关,但其特征是鞘上的横向脊和鞘开口端的短刺。罗奇科夫时代的新材料证明了磷酸环是拜伦子的附着结构。一个矮生圆锥虫Pidicocularia gen.nov.和模式种Pidicocleria tubulata sp.nov.以其微小的体积和非常精细的装饰而引人注目;其圆锥亲和性通过四个内部隆突和四个矩形下横截面得到证实。Prestephanoscyphus鞘的微观结构特征是排列的柱状微晶的致密层和磷灰石等轴晶的室状层交替。byronid鞘的增生生长和具有粘性踏板盘基底开口的固定器的结构支持了它们的cnidarian亲和力,尽管它们可能表现出双侧而不是四聚体对称性。提出了Prestephanoscyphus内质突两侧对称轮匝的保护作用,并对其在加强鞘壁或支持胃隔功能方面的作用存在争议。拜伦虫被认为是表生生物,而不是附着在岩石上。它们是具有磷壳的底栖生物群的代表,这些磷壳随着海洋中磷的可用性降低而减少。
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引用次数: 0
Tooth whorl structure, growth and function in a helicoprionid chondrichthyan Karpinskiprion (nom. nov.) (Eugeneodontiformes) with a revision of the family composition 螺旋体球粒陨石Karpinskiprion(nom.nov.)(Eugeneodontiformes)的牙齿轮生结构、生长和功能及其家族组成的修正
IF 1.2 4区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-12-01 DOI: 10.1017/S1755691022000251
O. A. Lebedev, W. Itano, Z. Johanson, A. Alekseev, Moya M. Smith, A. Ivanov, I. V. Novikov
ABSTRACT Restudy of Campyloprion annectans Eastman, 1902 from North America demonstrated that neither specimen included is diagnostic at the species level; thus, the species name is a nomen dubium. Since this species was designated as the type species of the genus, this requires suppression of the generic name also. Another species earlier assigned to Campyloprion, Campyloprion ivanovi Karpinsky, 1924 is used as a type for a newly established genus Karpinskiprion Lebedev et Itano gen. nov. The composition of the family Helicoprionidae Karpinsky, 1911 is reviewed, and a new family Helicampodontidae Itano et Lebedev fam. nov. is erected. A new specimen of Karpinskiprion ivanovi (Karpinsky, 1924) recently discovered in the Volgograd Region of Russia is the most complete Karpinskiprion specimen ever found. It unambiguously demonstrates the coiled nature of these tooth whorls and presents information on their developmental stages. During organogeny, cutting blades of the crown became reshaped, and basal spurs progressively elongated, forming a grater. Whorl growth occurred by addition of new crowns to the earlier mineralised base followed by later spur growth. In contrast to consistently uniform cutting blades, spurs are often malformed and bear traces of growth interruption. Both sides of the outer coil of the tooth whorl bear lifetime wear facets. The youngest (lingual) crowns are as yet unaffected by wear. The best-preserved facets show parallel radially directed scratch marks. The upper jaw dentition of Karpinskiprion is unknown, but we suggest that the faceted areas resulted from interaction with the antagonistic dental structures here. Three possible hypotheses for this interaction are suggested: (a) two opposing whorls acted as scissor blades, moving alternately from one side to another; (b) the lower tooth whorl fitted between paired parasymphyseal tooth whorls of the opposing jaw; or (c) the lower tooth whorl fitted into a dental pavement in the upper jaw.
摘要:对1902年来自北美的Campyloprion annectans Eastman的重新研究表明,这两个标本在物种水平上都不是诊断性的;因此,该种的名称是一个名称dubium。由于该物种被指定为该属的模式物种,这也需要抑制属名。另一个早期归属于Campyloprion的物种,Campyloprion-ivanovi-Karpinsky,1924被用作新建立的Karpinskiprion Lebedev et Itano gen.nov属的模式。回顾了Helicopionidae Karpinsky科的组成,1911,和一个新的Helicopiodontidae Itano et Lebedev fam。十一月成立。最近在俄罗斯伏尔加格勒地区发现的一个新的伊凡诺维卡平斯基虫标本(卡平斯基,1924年)是迄今为止发现的最完整的卡平斯基病毒标本。它明确地展示了这些齿轮匝的盘绕性质,并提供了关于其发育阶段的信息。在器官发生过程中,牙冠的切割刀片被重塑,基部刺逐渐拉长,形成一个磨碎器。通过在早期矿化基底上添加新冠,然后在后期进行刺状生长,从而实现了旋状生长。与一贯均匀的切割刀片相比,毛刺通常是畸形的,并带有生长中断的痕迹。齿轮圈的外圈两侧均具有寿命磨损面。最年轻的(舌侧)牙冠尚未受到磨损的影响。保存最完好的刻面显示平行的径向划痕。Karpinskiprion的上颚齿列尚不清楚,但我们认为刻面区域是与此处的拮抗性牙齿结构相互作用的结果。对于这种相互作用,提出了三种可能的假设:(a)两个相对的螺纹充当剪刀片,从一侧交替移动到另一侧;(b) 所述下齿轮圈配合在所述相对颌的成对副交感神经齿轮圈之间;或(c)安装到上颌中的牙齿铺面中的下牙螺纹。
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引用次数: 0
The remains of a large cercopithecid from the Lower Pleistocene locality of Karnezeika (southern Greece) 来自下更新世Karnezeika(希腊南部)地区的大型尾猿遗骸
IF 1.2 4区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-11-07 DOI: 10.1017/s1755691022000226
Panagiotis D. Sianis, Αthanassios Athanassiou, D. Kostopoulos, S. Roussiakis, N. Kargopoulos, G. Iliopoulos
The Lower Pleistocene Karnezeika locality, lies in the Peloponnese, southern Greece, and its fauna corresponds to the Middle Villafranchian biochronological unit (MN17). The recovered mammal assemblage includes, among others, a few remains of a large Cercopithecid. Herein, we describe this material, including an upper second molar, a partially preserved proximal radius and, possibly, an upper first incisor. The teeth show advanced stages of wear but retain their typical papionin characters, such as a strong lingual cleft and four bilophodont cusps in the molar. The general morphology and wear pattern of the teeth rules out the possibility that the remains belong to the genus Theropithecus, while the general size of the corresponding material excludes the possibility of a Macaca representative as well. On the contrary, the studied material better fits the size range of Paradolichopithecus. Even though this genus is likely represented in the Villafranchian of Europe by a single species, Par. arvernensis, the scarcity of the studied material imposes reservations and thus the Karnezeika papionin is referred at the moment to cf. Paradolichopithecus sp. As in the rest of Europe, the Paradolichopithecus record is rare in Greece, having been found in only two localities, Vatera and Dafnero. Despite its scarcity, the new material from Karnezeika indicates a wide distribution of this important taxon in the Greek peninsula.
下更新世Karnezeika地区位于希腊南部伯罗奔尼撒半岛,其动物群对应于中维拉弗兰奇生物地貌单元(MN17)。被发现的哺乳动物群落包括一些大型颈猿的遗骸。在此,我们描述了这种材料,包括上第二磨牙、部分保留的近端桡骨,以及可能的上第一切牙。牙齿表现出晚期磨损,但仍保持其典型的丘疹特征,如强烈的舌裂和臼齿中的四个双齿尖。牙齿的一般形态和磨损模式排除了遗骸属于Theropithecus属的可能性,而相应材料的一般大小也排除了Macaca代表的可能性。相反,所研究的材料更适合Paradolichopithecus的大小范围。尽管该属在欧洲的维拉弗兰奇亚可能由一个物种Par.avernensis代表,但所研究材料的稀缺性提出了保留意见,因此Karnezeika papionin目前被称为Paradolichopithecus sp.。与欧洲其他地区一样,Paradolichupithecus记录在希腊很罕见,只在两个地方发现,Vatera和Dafnero。尽管数量稀少,但来自Karnezeika的新材料表明,这一重要分类单元在希腊半岛分布广泛。
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引用次数: 1
A forgotten cirripedological gem: a new species of whale barnacle of the genus Cetopirus from the Pleistocene of the United States West Coast 被遗忘的河流学珍宝:来自美国西海岸更新世鲸豚属的一种新的鲸藤壶
IF 1.2 4区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-11-07 DOI: 10.1017/s1755691022000214
A. Collareta, M. Bosselaers, P. Holroyd, Ashley A. Dineen
A small lot of fossil whale barnacles from the Upper Pleistocene of California and the Middle Pleistocene (Chibanian) of Oregon (United States West Coast), described in a 1972 unpublished MA thesis, are formally described and illustrated herein. In that thesis, a new genus and species name were proposed; however, according to the International Code of Zoological Nomenclature, they have no taxonomic standing and are thus unavailable. Based on our reappraisal, two specimens in this lot belong to a new, extinct species that can be assigned to the purportedly extant genus Cetopirus. Cetopirus polysyrinx sp. nov. differs from congeners in that its secondary T-shaped flanges are multitubiferous internally, that is, they are perforated by a high number of irregularly-sized and irregularly-spaced tubules that result in a spongy aspect in transverse section. Whether or not this peculiar condition had any adaptive significance is difficult to determine. Considering that Cetopirus is currently known as an obligate epibiont of right whales (including the North Pacific form Eubalaena japonica (Lacépède 1818)), the host of C. polysyrinx sp. nov. was E. japonica or some other species of Eubalaena. The Plio-Pleistocene deposits of the Pacific coast of North America have yielded a rather idiosyncratic fossil whale barnacle fauna, inclusive of the genera Cetolepas, Cryptolepas and now Cetopirus, which seemingly contrasts with all other coeval assemblages worldwide, the latter being in turn dominated by Coronula spp.
1972年一篇未发表的MA论文中描述了大量来自加利福尼亚上更新世和俄勒冈州(美国西海岸)中更新世(Chibanian)的鲸鱼藤壶化石,本文对此进行了正式描述和说明。在这篇论文中,提出了一个新的属和种名称;然而,根据《国际动物命名法》,它们没有分类学地位,因此不可用。根据我们的重新评估,这批标本中的两个属于一个新的、已灭绝的物种,可以归属于据称现存的Cetopirus属。Cetopirus polysyrinx sp.nov.与同类的不同之处在于,它的次级T形凸缘内部是多管状的,也就是说,它们被大量大小不规则、间隔不规则的小管穿孔,导致横截面呈海绵状。这种特殊情况是否具有任何适应性意义尚难确定。考虑到Cetopirus目前被认为是露脊鲸的一种专性表生物(包括北太平洋形态的日本露脊鲸(Lacépède 1818)),C.polysyrinx sp.nov.的宿主是日本露脊鲸鱼或其他一些露脊鲸物种。北美洲太平洋海岸的上新世-更新世沉积物产生了一种相当独特的鲸鱼藤壶动物化石,包括鲸蜡属、隐壶属和现在的鲸蜡属,这似乎与世界各地的所有其他同时代组合形成了对比,后者反过来又以Coronaula属为主。
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引用次数: 1
Gigantoproductid shell spiral and microstructure of tertiary layer: evaluation as taxonomical characters 巨产品壳螺旋和三层显微结构:作为分类特征的评价
IF 1.2 4区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-10-21 DOI: 10.1017/s1755691022000196
J. R. MATEOS-CARRALAFUENTE, I. Coronado, P. Cózar, S. Rodríguez
Brachiopod taxonomy is based on descriptions of shell morphology and key characters, but diagenesis generally modifies or erases some of them, hindering brachiopod identification. Brachiopods that are taxonomically related usually present shells with similar appearance but can differ in size (i.e., Rhynchonellata). Some aspects of morphology – for example the angular measurement of the curvature of the shell or details of shell microstructure – could aid taxonomic identification. Gigantoproductids, which lack a robust taxonomy, have the largest shells among brachiopods and are ideal for this kind of study because of their gigantic size and morphological variability. Furthermore, they have a great abundance and worldwide distribution during the mid-Carboniferous. More than 700 samples have been collected from Sierra Morena (Spain), Montagne Noire (France) and Adarouch (Morocco) identifying up to six gigantoproductid genera: Globosoproductus, Semiplanus, Kansuella?, Latiproductus, Gigantoproductus and Datangia. Microstructural features from 170 thin sections belonging to gigantoproductid ventral valves have been studied, and six crystal morphologies have been distinguished within the tertiary layer: subhorizontal, imbricated, crenulated, acicular, short and long columnar morphologies. Moreover, 23 complete shells from all genera have been selected to investigate shell size and curvature. Results from this study emphasise that shell size, curvature and crystal shape are taxa-related. Finally, a remarkable morphological change in the gigantoproductid populations from the western Palaeo-Tethys occurred during the Viséan–Serpukhovian, from thin-shelled genera with subhorizontal morphology (Viséan) to thick-shelled genera with a tertiary layer consisting of long columnar crystals (Serpukhovian). This study proves that microstructure, maximum thickness and shell spiral characterisation are robust characters when applied to gigantoproductid taxonomy, but also have great potential in other brachiopod groups.
腕足类分类学是基于对外壳形态和关键特征的描述,但成岩作用通常会改变或抹去其中的一些,阻碍腕足类的鉴定。在分类学上有亲缘关系的腕足类通常有相似外观但大小不同的外壳(即Rhynchonellata)。形态学的某些方面——例如外壳曲率的角度测量或外壳微观结构的细节——可以帮助分类鉴定。巨型动物缺乏健全的分类学,在腕足类动物中外壳最大,由于其巨大的体型和形态变异性,是这类研究的理想对象。此外,在石炭纪中期,它们的数量非常丰富,分布在世界各地。从Sierra Morena(西班牙)、Montagne Noire(法国)和Adarouch(摩洛哥)采集了700多个样本,确定了多达6个巨大的生产属:Globosoproductus、Semiplanus、Kansuella?,Latiproductus、Gigantoproductus和Datangia。研究了170个巨大腹瓣薄片的显微结构特征,并在第三层内区分出六种晶体形态:近水平、叠瓦状、细齿状、针状、短柱状和长柱状。此外,从所有属中挑选了23个完整的贝壳来研究贝壳的大小和曲率。这项研究的结果强调,外壳大小、曲率和晶体形状是相关的分类群。最后,在维世-谢尔普霍夫阶期间,古特提斯西部的巨型生产种群发生了显著的形态变化,从具有亚水平形态的薄壳属(维世)到具有由长柱状晶体组成的第三层的厚壳属(谢尔普霍阶)。这项研究证明,微观结构、最大厚度和外壳螺旋特征在应用于大型动物分类学时是稳健的特征,但在其他腕足类中也有很大的潜力。
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Earth and Environmental Science Transactions of the Royal Society of Edinburgh
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