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Cephalopods from the Cretaceous-Paleogene (K-Pg) Boundary Interval on the Brazos River, Texas, and Extinction of the Ammonites 德克萨斯州布拉佐斯河白垩纪-古近纪(K-Pg)边界层的头足类动物和亚扪人的灭绝
IF 1.5 3区 环境科学与生态学 Q3 BIODIVERSITY CONSERVATION Pub Date : 2021-01-13 DOI: 10.1206/3964.1
J. Witts, N. Landman, M. Garb, Kayla M. Irizarry, E. Larina, N. Thibault, Mohammad Javad Razmjooei, T. Yancey, C. Myers
ABSTRACT We report on new collections of cephalopods (ammonites and nautilids) from the Cretaceous-Paleogene (K-Pg) successions of the Corsicana and Kincaid formations exposed along the Brazos River in Falls County, Texas. An abundant fauna of eight species comprising four genera of ammonites is described from the Corsicana Formation, including Discoscaphites mullinaxorum n. sp. The presence of abundant aptychi (probably lower jaws) of Discoscaphites and Eubaculites, as well as juvenile specimens, indicates a living population that experienced little postmortem drift. The lytoceratid genus Gaudryceras is also reported for the first time from the Brazos River area. Presence of the index taxon Discoscaphites iris (Conrad, 1858) indicates that the fauna belongs to the D. iris Range Zone, the highest ammonite range zone in North America. Correlation with new and existing microfossil data indicates that the fauna represents the uppermost Maastrichtian, and comparison with published records further suggests that this is the most diverse D. iris Zone fauna yet reported from the Gulf and Atlantic Coastal Plains of North America. Three ammonite genera are recorded from the basal units of the K-Pg event deposit at Brazos, which likely represents deposition in the immediate aftermath of the Chicxulub impact event. A single specimen of the nautilid Eutrephoceras is reported from the Danian Kincaid Formation, less than 300 kyr after the K-Pg boundary. These data provide new information on the differing fate of these cephalopod groups during the K-Pg mass extinction and add to the picture of diverse and abundant Maastrichtian ammonite faunas prior to the Chicxulub impact event.
摘要:我们报道了德克萨斯州福尔斯县Brazos河沿岸科西嘉那和金凯德地层白垩纪-古近纪(K-Pg)序列中新发现的头足类动物(菊石和鹦鹉螺)。科西嘉那组中有一个由八个物种组成的丰富动物群,包括四个属的菊石,包括斑蝶属(Discoscaphites mullinaxorum n.sp.)。斑蝶属和Eubaculites的大量aptychi(可能是下颚)以及幼年标本的存在表明,活的种群很少经历死后漂移。在布拉佐斯河地区也首次报道了lytocerated Gaudryceras属。索引分类单元Discoscaphites iris(Conrad,1858)的存在表明该动物群属于D.iris山脉带,这是北美最高的菊石山脉带。与新的和现有的微体化石数据的相关性表明,该动物群代表了最上层的马斯特里赫特阶,与已发表的记录的比较进一步表明,这是迄今为止北美海湾和大西洋海岸平原报告的最多样的D.iris带动物群。Brazos K-Pg事件矿床的基底单元中记录了三个菊石属,这可能代表了希克苏鲁伯撞击事件后的沉积。据报道,在K-Pg边界后不到300公里的Danian Kincaid地层中有一个鹦鹉螺Eutrephoceras的单一标本。这些数据为K-Pg大灭绝期间这些头足类的不同命运提供了新的信息,并为希克苏鲁伯撞击事件之前马斯特里赫特菊石动物群的多样性和丰富性增添了新的内容。
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引用次数: 5
Cretaceous Diversity of the Relict Genus Alavesia Waters and Arillo (Diptera: Empidoidea: Atelestidae) 白垩纪残属Alavesia Waters和Arillo的多样性(双翅目:翅甲总科:翅甲科)
IF 1.5 3区 环境科学与生态学 Q3 BIODIVERSITY CONSERVATION Pub Date : 2020-10-06 DOI: 10.1206/3961.1
B. Sinclair, D. Grimaldi
ABSTRACT Alavesia has been known on the basis of five species in Late Cretaceous amber from Spain and Myanmar and two extant species from Namibia. Very recently, another extant species has been found in southern Brazil. Here we present a surprising diversity of Alavesia flies from the mid-Cretaceous of Myanmar (Late Albian-Early Cenomanian, ca. 99 Ma), including nine new species: A. angusta, sp. nov., A. brevipennae, sp. nov., A. lanceolata, sp. nov., A. latala, sp. nov., A. longicornuta, sp. nov., A. magna, sp. nov., A. pankowskiorum, sp. nov., A. spinosa, sp. nov., and A. zigrasi, sp. nov. The following new synonymy is proposed: Neoalavesia Poinar and Vega, 2020 = Alavesia Waters and Arillo, 1999. Notes are provided on an unusual structure, leg furrows found on femora to varying degrees among species, sometimes on tibiae, as well as notes on swarming behavior. Alavesia has undergone extensive extinction, a finding that further supports the relict nature of this genus and helps to explain the distribution of the other four Recent genera of Atelestidae, found in south and north temperate regions. Relict arthropod taxa preserved in Burmese amber are briefly reviewed, which also are narrowly endemic to temperate regions today.
摘要阿拉维西亚是根据西班牙和缅甸晚白垩世琥珀中的五个物种和纳米比亚现存的两个物种而发现的。最近,在巴西南部发现了另一种现存物种。在这里,我们展示了来自缅甸白垩纪中期(晚阿尔比阶-早Cenomanian阶,约99 Ma)的Alavesia蝇的惊人多样性,包括九个新种:a.angusta,sp.nov.,a.brevipennae,sp.nova.,a.lanceolata,sp.nov,a.latala,sp.nv.,a.longiconuta,sp.nov.,a.magna,sp.niv.,a.pankowskiorum,sp.nv..,a.spinosa,sp.nev.和a.zigrasi,sp.nav。提出了以下新的同义词:Neoalavesia Poinar和Vega,2020=Alavesia Waters和Arillo,1999。提供了关于一种不寻常结构的注释,在不同物种的股骨上发现了不同程度的腿沟,有时在胫骨上,以及关于群集行为的注释。Alavesia已经经历了大范围的灭绝,这一发现进一步支持了该属的残余性质,并有助于解释在南温带和北温带发现的Atelestidae最近的其他四个属的分布。简要回顾了缅甸琥珀中保存的节肢动物遗迹分类群,这些分类群也是当今温带地区的特有种。
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引用次数: 5
First Known Extant Species of Alavesia (Diptera: Atelestidae) in the Neotropical Region: Alavesia leukoprosopa, sp. nov., from the Southern Atlantic Forest, Brazil 新热带地区已知的第一个现存物种:巴西南大西洋森林的白腹蛛属
IF 1.5 3区 环境科学与生态学 Q3 BIODIVERSITY CONSERVATION Pub Date : 2020-10-06 DOI: 10.1206/3962.1
Dalton De Souza Amorim, P. Riccardi, J. A. Rafael
ABSTRACT A first known extant Neotropical species of the atelestid genus Alavesia—Alavesia leukoprosopa, sp. nov.—is described from southeastern Brazil. The holotype (and only specimen so far) was collected with a Malaise trap in an area with strongly impacted semideciduous seasonal forest (dry forest) and open, entirely secondary vegetation. It was collected in mid spring, but has not been collected again so far. Alavesia leukoprosopa shares some derived features with the only two other extant species of the genus, known from the Brandberg Massif in Namibia, suggesting that all three extant species may compose together a small clade separate from the bulk of the Cretaceous diversity of Alavesia.
摘要:在巴西东南部发现了第一个已知的新热带无尾木犀属物种——白叶木犀(Alavesia leukoprospa,sp.nov.)。正模标本(也是迄今为止唯一的标本)是用Malaise陷阱在一个半干旱季节性森林(干燥森林)和开放的完全次生植被受到强烈影响的地区采集的。它是在仲春收集的,但到目前为止还没有被再次收集。白蜡木属与该属仅有的另外两个现存物种(已知于纳米比亚的Brandberg Massif)有一些衍生特征,这表明所有三个现存物种可能共同组成一个小分支,与白蜡木白垩纪的大部分多样性分离。
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引用次数: 3
On the Identity of Victoria's Mouse Opossum, Marmosa regina Thomas, 1898 论维多利亚的老鼠Opossum,Marmosa regina Thomas,1898
IF 1.5 3区 环境科学与生态学 Q3 BIODIVERSITY CONSERVATION Pub Date : 2020-09-10 DOI: 10.1206/3960.1
Thomas C. Giarla, R. Voss
ABSTRACT Phylogenetic analyses of molecular sequence data obtained from the holotype of Marmosa regina Thomas, 1898, together with a reassessment of its morphological characters indicate that this species does not belong to the subgenus Micoureus as previously believed. Instead, both molecular and phenotypic data are consistent with the hypothesis that M. regina is a senior synonym of M. isthmica Goldman, 1912, in the subgenus Exulomarmosa. Because replacing isthmica with regina would create nomenclatural confusion, we recommend maintaining current usage of the former name and suppressing usage of the latter.
摘要:从区域Marmosa的正模标本中获得的分子序列数据的系统发育分析,Thomas,1898,以及对其形态特征的重新评估表明,该物种不属于以前认为的Micoureus亚属。相反,分子和表型数据都与M.regina是M.isthmica Goldman(1912)在Exulomarmosa亚属中的高级异名的假设一致。因为用regina代替isthmica会造成命名混乱,我们建议保持前一个名称的当前用法,并抑制后一个名称。
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引用次数: 3
Taxonomic Revision of Camarochilus Harris (Hemiptera: Pachynomidae) 金针虫分类修订(半翅目:扁虫科)
IF 1.5 3区 环境科学与生态学 Q3 BIODIVERSITY CONSERVATION Pub Date : 2020-09-08 DOI: 10.1206/3959.1
C. Weirauch, D. Forero, R. Schuh
ABSTRACT The Neotropical genus Camarochilus Harris, 1930 (Hemiptera: Pachynomidae) was described based on two species from Panama and Brazil. The genus has not been revised since its original treatment, and specimens beyond the original type series have remained undocumented. Based on examination of 57 specimens, including holotypes or paratypes of the previously described species, we here revise Camarochilus and recognize 10 species, with eight described as new: C. fasciatus, n. sp., C. gilli, n. sp., C. globosus, n. sp., C. harrisi, n. sp., C. medius, n. sp., C. picturatus, n. sp., C. robustus, n. sp., and C. tenuis, n. sp. Habitus images are provided for all species, male and female genitalic structures are documented, and various morphological characters are illustrated using digital photography. Measurements are included for all species. Male genitalic characters including the degree of asymmetry and type of ornamentation of the pygophore and shape of parameres constitute important species-diagnostic features, in addition to coloration and measurements. Distribution maps illustrate that species occur from Bolivia to Honduras, a considerable expansion of the previously known species distribution ranges in this genus.
摘要根据巴拿马和巴西的两种记述了新热带Camarochilus Harris属(半翅目:厚翅虫科)。该属自其原始处理以来未被修订,并且超出原始类型系列的标本仍未记录。基于对57个标本的检查,包括先前描述的物种的全型或半型,我们在这里修改了Camarochilus,并识别出10种,其中8种被描述为新物种:本研究提供了所有物种的习性图像,记录了雄性和雌性的生殖器结构,并利用数码摄影技术对各种形态特征进行了说明。测量包括所有物种。除了颜色和测量外,男性生殖器特征,包括生殖器的不对称程度和装饰类型以及参数的形状,都是重要的物种诊断特征。分布图表明,物种分布在玻利维亚到洪都拉斯,这是该属以前已知的物种分布范围的一个相当大的扩展。
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引用次数: 0
Systematics of Neotropical Spiny Mice, Genus Neacomys Thomas, 1900 (Rodentia: Cricetidae), from Southeastern Amazonia, with Descriptions of Three New Species 亚马逊东南部新热带棘鼠属分类学(啮齿目:鼠科)及三新种记述
IF 1.5 3区 环境科学与生态学 Q3 BIODIVERSITY CONSERVATION Pub Date : 2020-09-05 DOI: 10.1206/3958.1
Thiago Borges Fernandes Semedo, M. N. D. Da Silva, E. Gutiérrez, D. C. Ferreira, Mario Da Silva Nunes, A. C. Mendes-Oliveira, I. Farias, R. Rossi
ABSTRACT Species of Neacomys are small cricetid rodents that occur in forested habitats of Central and South America, from eastern Panama to central Bolivia and central/western Brazil. In order to assess species diversity of this poorly known genus, we obtained cytochrome b gene sequences from the most comprehensive taxonomic and geographic sampling analyzed to date. We also conducted morphological analyses on a large series of specimens housed in 15 museums, including types of 10 out of 14 nominal taxa. Our analyses of the genetic data recovered 17 lineages clustered in four distinct clades. Among these lineages, 11 correspond to species currently recognized as valid, and the remaining six are putative new species. In southeastern Amazonia—the geographical scope of this report—four undescribed species were discovered, three of which are named herein: Neacomys marajoara, sp. nov., from the Island of Marajó, Pará state; Neacomys vossi, sp. nov., restricted to the Tapajós center of endemism (between the Tapajós and Xingu rivers); and Neacomys xingu, sp. nov., restricted to the Xingu center of endemism (between the Xingu and Araguaia/Tocantins rivers). The new species can be discriminated from other Neacomys species by the morphology of the nasal bones, zygomatic plate, interorbital region, subsquamosal fenestra, paraoccipital process, incisive foramina, auditory bullae, anterocone and anteroloph of the first upper molar, carotid circulation pattern, and karyotype. Our results substantially improve our understanding of the genus Neacomys by providing morphological, morphometric, and novel molecular insights about these poorly known rodents and demonstrate that the diversity of small Amazonian mammals is still poorly known, even in the relatively accessible southeastern part of the biome.
Neacomys是一种小型环状啮齿动物,分布于中美洲和南美洲的森林栖息地,从巴拿马东部到玻利维亚中部和巴西中西部。为了评估这一鲜为人知的属的物种多样性,我们从迄今为止分析过的最全面的分类和地理样本中获得了细胞色素b基因序列。我们还对15个博物馆收藏的大量标本进行了形态学分析,包括14个名义分类群中的10个类型。我们对遗传数据的分析恢复了17个谱系,这些谱系聚集在四个不同的分支中。在这些谱系中,11个对应于目前公认的有效物种,其余6个是假定的新物种。在亚马逊东南部——本报告的地理范围——发现了四个未描述的物种,其中三个在此命名:Neacomys marajoara, sp. nov.,来自Marajó, par州岛;Neacomys vossi, sp. nov.,局限于Tapajós特有种中心(Tapajós和新谷河之间);和Neacomys xinu, sp. nov.,局限于xinu的特有中心(在xinu和阿拉瓜亚/托坎廷斯河之间)。根据鼻骨、颧骨板、眶间区、鳞下孔、枕旁突、锐孔、听大泡、第一上磨牙前锥体和前窝、颈动脉循环模式和核型等形态学特征,可将新种与其他新种区分。我们的研究结果通过提供关于这些鲜为人知的啮齿动物的形态学、形态计量学和新的分子见解,大大提高了我们对Neacomys属的理解,并表明即使在相对容易接近的生物群系东南部,小型亚马逊哺乳动物的多样性仍然知之甚少。
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引用次数: 9
Eomakhaira molossus, A New Saber-Toothed Sparassodont (Metatheria: Thylacosmilinae) from the Early Oligocene (?Tinguirirican) Cachapoal Locality, Andean Main Range, Chile Eomakhaira molossus,智利安第斯山脉Cachapoal地区渐新世早期的一种新的剑齿雀形目(Metatheria:Thylacosmilinae)
IF 1.5 3区 环境科学与生态学 Q3 BIODIVERSITY CONSERVATION Pub Date : 2020-07-17 DOI: 10.1206/3957.1
Russell K. Engelman, J. J. Flynn, A. Wyss, D. Croft
ABSTRACT Thylacosmiline sparassodonts (previously recognized as thylacosmilids) are among the most iconic groups of endemic South American Cenozoic mammals due to their distinctive morphology and convergent resemblance to saber-toothed placental carnivores. However, the early evolution of this group and its relationship to other sparassodonts remains poorly understood, primarily because only highly specialized Neogene taxa such as Thylacosmilus, Anachlysictis, and Patagosmilus are well known. Here, we describe a new Paleogene sparassodont, Eomakhaira molossus, from the Cachapoal locality of central Chile, the first sparassodont reported from early Oligocene strata of the Abanico Formation. Eomakhaira shares features with both Neogene thylacosmilines and Paleogene “proborhyaenids,” and phylogenetic analyses recover this taxon as sister to the clade of Patagosmilus + Thylacosmilus. This broader clade, in turn, is nested within the group conventionally termed Proborhyaenidae. Our analyses support prior hypotheses of a close relationship between thylacosmilines and traditionally recognized proborhyaenids and provide the strongest evidence to date that thylacosmilines are proborhyaenids (i.e, the latter name as conventionally used refers to a paraphyletic group). To reflect the internestedness of these taxa, we propose use of Riggs' (1933) original name Thylacosmilinae for the less inclusive grouping and Proborhyaenidae for the more inclusive one. Saber teeth arose just once among metatherians (among thylacosmilines), perhaps reflecting a developmental constraint related to nonreplacement of canines in metatherians; hypselodonty may have relaxed this potential constraint in thylacosmilines. The occurrence of Eomakhaira in strata of early Oligocene age from the Chilean Andes demonstrates that the stratigraphic range of thylacosmilines spanned almost 30 million years, far surpassing those of saber-toothed placental lineages.
摘要:Thylacosmiline sparassodons(以前被认为是乙基粘粒动物)是南美新生代特有哺乳动物中最具标志性的类群之一,因为它们具有独特的形态,与剑齿胎盘食肉动物趋同相似。然而,该类群的早期进化及其与其他雀形目的关系仍知之甚少,主要是因为只有高度特化的新第三纪分类群,如Thylacosmilus、Anachlysectis和Patagosmilus是众所周知的。在这里,我们描述了一种来自智利中部Cachapoal地区的新的古近系裂头龙,Eomakhaira molossus,这是阿巴尼科组渐新世早期地层中报道的第一种裂头龙。Eomakhaira与新第三纪的ethyacosmilines和古第三纪“proborhyaenids”都有共同的特征,系统发育分析将该分类单元恢复为Patagosmilus+Thylacosmilus分支的姐妹。反过来,这个更广泛的分支嵌套在传统上被称为Proborhyaenidae的群体中。我们的分析支持了先前关于甲基粘蛋白和传统上公认的透明质类之间密切关系的假设,并提供了迄今为止最有力的证据,证明甲基粘蛋白是透明质类(即,传统上使用的后一个名称指代一个异基因群)。为了反映这些分类群的国际性,我们建议使用Riggs(1933)的原名Thylacosmilinae作为不太包容的类群,使用Proborhyaenidae作为更包容的类群。剑齿在metatherians中只出现过一次(在乙基cosmilines中),这可能反映了与Metatherianes中犬科动物不安置有关的发育限制;hypselodonty可能已经放松了甲基粘粒蛋白的这种潜在限制。Eomakhaira在智利安第斯山脉渐新世早期地层中的出现表明,类囊动物的地层范围跨越了近3000万年,远远超过了剑齿胎盘谱系。
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引用次数: 9
Reconstructing the Specimens and History of Howe Quarry (Upper Jurassic Morrison Formation; Wyoming) 重建豪采石场的标本和历史(怀俄明州上侏罗纪Morrison组)
IF 1.5 3区 环境科学与生态学 Q3 BIODIVERSITY CONSERVATION Pub Date : 2020-06-24 DOI: 10.1206/3956.1
E. Tschopp, C. Mehling, M. Norell
ABSTRACT In 1934, Barnum Brown of the American Museum of Natural History in New York (AMNH) led a large-scale dinosaur excavation in northern Wyoming, where he had found bones two years earlier. Initially, Brown expected to excavate two skeletons of sauropod dinosaurs, but soon after opening the quarry, the team realized that the site far exceeded their expectations: in the end, they unearthed approximately 3000 bones within six months and sent approximately 144 crates back to the AMNH. Due to the enormous number of bones, the site became world famous as a dinosaur graveyard, and media from all over the United States and abroad reported on the expedition. Soon after, however, the collection shifted away from a curatorial focus. What followed was a history of neglect: inappropriate storage conditions, water and fire damage, collapsing wooden boxes, and deteriorating plaster jackets. Relocation and further excavation of the quarry by the Sauriermuseum Aathal (Switzerland; SMA) from 1989 to 1991 confirmed earlier finds of skin impressions and resulted in the recognition of a novel diplodocid sauropod: Kaatedocus siberi. Given that the amount of bones found by the AMNH far exceeded those found by the SMA, a new project was started in 2017 to reassess the state and scientific value of the historic collections at AMNH. Although most of the bones are heavily fragmented, preliminary results show that overall preservation is still exceptional. Here, we reconstruct the history of the excavation, as well as past conservation and preparation procedures, and provide a report of current efforts to prepare, conserve, and catalog the material that has remained in storage since the 1930s. These current efforts show that historic collections, even after decades of neglect, can be of great and unexpected value, both for research and scientific outreach.
1934年,纽约美国自然历史博物馆(AMNH)的巴纳姆·布朗(Barnum Brown)在怀俄明州北部领导了一次大规模的恐龙发掘活动,他两年前在那里发现了恐龙骨头。最初,布朗希望挖掘出两具蜥脚类恐龙的骨骼,但在采石场开放后不久,团队意识到这个遗址远远超出了他们的预期:最终,他们在六个月内挖掘出了大约3000具骨骼,并将大约144个箱子送回了美国国家博物馆。由于大量的骨头,该遗址成为世界闻名的恐龙墓地,美国各地和国外的媒体都报道了这次探险。然而,不久之后,这个系列就从策展的焦点转移了。随之而来的是被忽视的历史:不适当的储存条件,水和火的破坏,倒塌的木箱,和老化的石膏外套。由saurermuseum Aathal(瑞士)重新安置和进一步挖掘采石场;1989年至1991年的一项研究证实了早期发现的皮肤印痕,并确认了一种新的梁龙科蜥脚类动物:Kaatedocus siberi。鉴于AMNH发现的骨头数量远远超过SMA发现的骨头数量,2017年开始了一项新的项目,以重新评估AMNH历史收藏品的状态和科学价值。虽然大部分骨头都严重破碎,但初步结果表明,整体保存情况仍然很好。在这里,我们重建了挖掘的历史,以及过去的保护和准备程序,并提供了一份报告,介绍了自20世纪30年代以来一直保存在仓库中的材料的准备、保护和编目工作。目前的这些努力表明,即使经过几十年的忽视,历史收藏品在研究和科学推广方面也可能具有巨大而意想不到的价值。
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引用次数: 4
A Revision of Philander (Marsupialia: Didelphidae), Part 2: Phylogenetic Relationships and Morphological Diagnosis of P. nigratus Thomas, 1923 Philander(有袋动物:Didelphidae)的修订,第2部分:黑腹蛛的系统发育关系和形态学诊断Thomas,1923
IF 1.5 3区 环境科学与生态学 Q3 BIODIVERSITY CONSERVATION Pub Date : 2020-06-24 DOI: 10.1206/3955.1
R. Voss, Thomas C. Giarla
ABSTRACT Newly available molecular sequences and morphological data suggest that Philander nigratus Thomas, 1923, is a valid species. Currently known from just eight specimens collected in the Peruvian departments of Junín and Ayacucho, P. nigratus does not appear to be closely related to either of the congeneric taxa with which it was previously synonymized.
新获得的分子序列和形态学数据表明,Philander nigratus Thomas, 1923是一个有效的种。目前仅从秘鲁Junín和Ayacucho部门收集的8个标本中了解到,P. nigratus似乎与之前被称为同属分类群的任何一个都没有密切关系。
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引用次数: 3
A New Genus of Late Cretaceous Angel Shark (Elasmobranchii; Squatinidae), with Comments on Squatinid Phylogeny 晚白垩世天使鲨一新属;下蹲科),并对下蹲科系统发育的评述
IF 1.5 3区 环境科学与生态学 Q3 BIODIVERSITY CONSERVATION Pub Date : 2020-06-16 DOI: 10.1206/3954.1
J. Maisey, Dana J. Ehret, J. Denton
ABSTRACT Three-dimensional Late Cretaceous elasmobranch endoskeletal elements (including palatoquadrates, ceratohyals, braincase fragments, and a series of anterior vertebrae) are described from the Late Cretaceous University of Alabama Harrell Station Paleontological Site (HSPS), Dallas County, Alabama. The material is referred to the extant elasmobranch Family Squatinidae on the basis of several distinctive morphological features. It also exhibits features not shared by any modern or fossil Squatina species or the extinct Late Jurassic squatinid Pseudorhina. A new genus and species is erected, despite there being some uncertainty regarding potential synonymy with existing nominal species previously founded on isolated fossil teeth (curiously, no squatinid teeth have been documented from the HSPS). A preliminary phylogenetic analysis suggests that the new genus falls on the squatinid stem, phylogenetically closer to Squatina than Pseudorhina. The craniovertebral articulation in the new genus exhibits features considered convergent with modern batomorphs (skates and rays), including absence of contact between the posterior basicranium and first vertebral centrum, and a notochordal canal which fails to reach the parachordal basicranium. Supporting evidence that similarities in the craniovertebral articulation of squatinoids and batomorphs are convergent rather than synapomorphic (as “hypnosqualeans”) is presented by an undescribed Early Jurassic batomorph, in which an occipital hemicentrum articulates with the first vertebral centrum as in all modern sharklike (selachimorph) elasmobranchs. The fossil suggests instead that the batomorph synarcual evolved by fusion of the anterior basiventral and basidorsal cartilages prior to the reduction of the anterior centra and loss of the occipital hemicentrum, not afterward as predicted by the hypnosqualean hypothesis.
在阿拉巴马州达拉斯县的晚白垩世阿拉巴马大学哈雷尔站古生物遗址(HSPS)中,描述了三维晚白垩世elasmobranch内骨骼元素(包括腭方体、角状体、脑壳碎片和一系列前椎骨)。根据几个独特的形态特征,该材料被指为现存的板颌科蹲蝇科。它还展示了任何现代或化石下蹲物种或已灭绝的晚侏罗世下蹲物种所没有的特征。一个新的属和种被建立,尽管有一些不确定的潜在的同义词与现有的名义物种先前建立在孤立的化石牙齿(奇怪的是,没有蹲下人的牙齿从HSPS记录)。初步的系统发育分析表明,新属落在蹲属茎上,系统发育上更接近于蹲属而不是假冠属。新属的颅椎关节表现出被认为与现代蝙蝠兽(鳐和鳐)趋同的特征,包括后基颅和第一椎体之间缺乏接触,以及一条不能到达副基颅的脊索管。早侏罗世的一种未描述的蝙蝠形提供了支持证据,证明蹲鲨类和蝙蝠形的颅椎关节的相似性是趋同的,而不是突触性的(如“睡角鲨类”),其中一个枕骨半中心与第一个椎体关节相连,就像所有现代鲨鱼类(蛛形)的蛛形一样。相反,化石表明,在前中央减少和枕骨半中心丢失之前,由前基腹软骨和前基背软骨融合进化而来的,而不是像催眠角鲨假说所预测的那样,在此之后。
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引用次数: 3
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American Museum Novitates
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