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Bivalves of superfamily Galeommatoidea (Mollusca, Bivalvia) from western South Africa, with observations on commensal relationships and habitats 南非西部 Galeommatoidea 超科双壳类动物(软体动物,双壳目),以及对共生关系和栖息地的观察
IF 1.3 3区 生物学 Q2 ZOOLOGY Pub Date : 2024-07-22 DOI: 10.3897/zookeys.1207.124517
P. Valentich-Scott, Charles Griffiths, J. Landschoff, Ruiqi Li, Jingchun Li
The Galeommatoidea are a diverse but little-studied group of small bivalves, well known for the symbiotic relationships many species have with a range of invertebrate taxa. Four species collected from the Western Cape region of South Africa were examined and illustrated, providing new details on their habitat preferences, and depicting the mantle structure of live specimens for the first time. Brachiomya ducentiunussp. nov., is described herein, and an additional record of Montacuta substriata (Montagu, 1808) is reported from South Africa. Brachiomya ducentiunus and Montacuta substriata have obligate symbiotic relationships with different burrowing echinoids, while Kellia becki (WH Turton, 1932) and Melliteryx mactroides (Hanley, 1857) are free-living. DNA data and phylogenetic analyses are provided for three of the species.
Galeommatoidea 是一个种类繁多但鲜有研究的小型双壳类群,以许多物种与一系列无脊椎动物类群的共生关系而闻名。研究人员对从南非西开普地区采集的四个物种进行了研究并绘制了图解,提供了关于其栖息地偏好的新细节,并首次描绘了活体标本的外壳结构。本文描述了 Brachiomya ducentiunussp.Brachiomya ducentiunus 和 Montacuta substriata 与不同的穴居回声虫有强制性共生关系,而 Kellia becki (WH Turton, 1932) 和 Melliteryx mactroides (Hanley, 1857) 则是自由生活的。本文提供了其中三个物种的 DNA 数据和系统发育分析。
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引用次数: 0
Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae) Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)
IF 1.3 3区 生物学 Q2 ZOOLOGY Pub Date : 2024-07-22 DOI: 10.3897/zookeys.1207.117225
Robert B. Angus, Jason F. Maté, Elizabeth M. Angus, David Král
The Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 are revised using a combination of chromosome analysis, molecular phylogenetics and morphological statistical analysis. Sixteen species are recognised, one of which is shown to comprise two subspecies. Based mainly on the morphology of the aedeagal endophallus and the phylogenetic analysis, they are placed in two groups: the. niger group, apparently monophyletic, comprising Aphodius (Liothorax) niger Illiger, 1798, A. (L.) muscorum (Ádám, 1994), stat. rest., A. (L.) felixsp. nov., A. (L.) bellumgerenssp. nov., A. (L.) bameulisp. nov., A. (L.) krellisp. nov., A. (L.) isikdagensis (Balthasar, 1953), A. (L.) albertisp. nov. and A. (L.) wilsonae Maté et Angus, 2005, stat. rest.; and the plagiatus group, almost certainly paraphyletic, comprising A. (L.) plagiatus (Linnaeus, 1767), including A. (L.) p. plagiatus and A. (L.) p. sinoplagiatussubsp. nov., A. (L.) rodrigoisp. nov., A. (L.) discoidesA. Schmidt, 1916, stat. rest., A. (L.) rutilipennis (Baudi di Selve, 1870), stat. rest., A. (L.) chellalasp. nov., A. (L.) kraatzi Harold, 1868, and A. (L.) rusakovi Gusakov, 2004. A key to the species is given as well as details of their morphology, distributions, and habitats.
结合染色体分析、分子系统学和形态统计分析,对 Aphodius Hellwig, 1798 亚属 Liothorax Motschulsky, 1860 的古北欧种进行了修订。共确认了 16 个种,其中一个种包括两个亚种。主要根据雌蕊柄内口的形态和系统发生学分析,它们被归入两个类群:尼格类,显然是单系的,包括 Aphodius (Liothorax) niger Illiger, 1798, A. (L.) muscorum (Ádám, 1994), stat、A. (L.) felixsp. nov., A. (L.) bellumgerenssp. nov., A. (L.) bameulisp. nov., A. (L.) krellisp. nov.、(L.) isikdagensis (Balthasar, 1953), A. (L.) albertisp. 新种和 A. (L.) wilsonae Maté et Angus, 2005, stat、新种,A. (L.) rodrigoisp.本文给出了这些物种的检索表,并详细介绍了它们的形态、分布和栖息地。
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引用次数: 0
Amphibian survey of Ko Pha-gnan in Surat Thani Province, Thailand. 泰国素叻他尼府帕南岛两栖动物调查。
IF 1.3 3区 生物学 Q2 ZOOLOGY Pub Date : 2024-07-19 eCollection Date: 2024-01-01 DOI: 10.3897/zookeys.1207.116758
Dawn R Cook-Price, Sunchai Makchai, Sasitorn Hasin, Pongthep Suwanwaree

Insular amphibian species are often overlooked, rendering them vulnerable to habitat encroachment and other anthropogenic threats. The aim of this study was to compile a comprehensive list of amphibian species on Ko Pha-ngan in Surat Thani Province, Thailand. Data were collected via transect surveys and drift line fence traps in three different habitat types from February 2021 to September 2023. Our efforts detected 12 unique amphibian species in each of the three habitat types. The highest number of detections was observed in the Ko Pha-ngan-Than Sadet National Park protected areas. The common tree frog (Polypedatesleucomystax) and the common Asian toad (Duttaphrynusmelanostictus) were the two most abundantly found species on the island, whereas the Koh Tao caecilian (Ichthyophiskohtaoensis) and the newly described false Doria's fanged frog (Limnonectespseudodoriae) where the least commonly found species. In addition, Microhylaheymonsi and Fejervaryalimnocharis tadpoles were observed developing in high-salinity water bodies. Many species have shown a high tolerance in human-dominated landscapes. This study sheds light on the need for additional monitoring to better understand the dynamics of endemic species in addition to the impact tourism-driven development and habitat destruction has on a species with an insularly finite habitat.

海岛两栖动物物种经常被忽视,导致它们很容易受到栖息地侵占和其他人为威胁的影响。本研究的目的是编制一份泰国素叻他尼府帕岸岛两栖动物物种综合名录。从 2021 年 2 月到 2023 年 9 月,我们在三种不同的栖息地类型中通过横断面调查和漂移线围栏诱捕器收集了数据。我们在三种栖息地类型中的每一种都发现了 12 种独特的两栖动物。发现数量最多的是高帕岸-丹萨德国家公园保护区。常见的树蛙(Polypedatesleucomystax)和常见的亚洲蟾蜍(Duttaphrynusmelanostictus)是岛上发现最多的两种两栖动物,而涛岛蛙(Ichthyophiskohtaoensis)和新描述的假多利亚獠蛙(Limnonectespseudodoriae)则是发现最少的物种。此外,在高盐度水体中还观察到蝌蚪发育的微型蛙(Microhylaheymonsi)和蝾螈(Fejervaryalimnocharis)。许多物种在人类主导的景观中表现出很强的耐受性。这项研究表明,除了旅游业驱动的开发和栖息地破坏对栖息地有限的物种造成的影响外,还需要进行更多监测,以更好地了解特有物种的动态。
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引用次数: 0
A new species of Neotropiconyttus Kirkaldy (Hemiptera, Reduviidae) in a cacao plantation from the Colombian Napo Province, including a key to species, and taxonomic notes of the genus. 哥伦比亚纳波省可可种植园中的一个新物种 Neotropiconyttus Kirkaldy(半翅目,Reduviidae),包括物种检索表和该属的分类说明。
IF 1.3 3区 生物学 Q2 ZOOLOGY Pub Date : 2024-07-19 eCollection Date: 2024-01-01 DOI: 10.3897/zookeys.1207.120663
Jean Gamboa, Hélcio R Gil-Santana, Armando Gamboa-Tabares, Eidy Martínez-Viuche, Francisco Serna

A new species of the genus Neotropiconyttus Kirkaldy, 1909 (Hemiptera: Heteroptera: Reduviidae: Harpactorinae: Harpactorini) is described and illustrated. Neotropiconyttusarmandoi Gamboa & Gil-Santana, sp. nov. represents the first record of the genus for the Province of Napo in Colombia, and the first description of a male individual in the genus. The male specimen representing the new species was collected on a leaf of cacao (Theobromacacao L.-Malvaceae). Its remarkable similarity in external coloration and structure with that of the true bug Monaloniondissimulatum Distant, 1883 (Hemiptera: Miridae) inhabiting cacao agroforestry systems suggests that the new species could be part of a mimetic complex that incorporates phytophagous and predator bugs. Comments and figures of type specimens of Neotropiconyttusalboannulatus (Stål, 1855) and Neotropiconyttusdama (Burmeister, 1838), and a key to the species of the genus are also provided.

本研究描述了 Neotropiconyttus Kirkaldy, 1909 属的一个新种(半翅目:异翅目:红绣线菊科:绣线菊属:绣线菊科),并绘制了插图。Neotropiconyttusarmandoi Gamboa & Gil-Santana,sp. nov.是该属在哥伦比亚纳波省的首次记录,也是对该属雄性个体的首次描述。代表该新种的雄性标本采集于可可(Theobromacacao L.-Malvaceae)的一片叶子上。它的外部颜色和结构与栖息在可可农林系统中的真蝽 Monaloniondissimulatum Distant, 1883 (Hemiptera: Miridae) 非常相似,这表明该新种可能是一个包含植食性蝽和捕食性蝽的拟态复合体的一部分。此外,还提供了 Neotropiconyttusalboannulatus (Stål, 1855) 和 Neotropiconyttusdama (Burmeister, 1838) 的模式标本的注释和图像,以及该属物种的检索表。
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引用次数: 0
Taxonomic considerations of selected Western Palaearctic Mordellidae Latreille, 1802 (Coleoptera, Tenebrionoidea) 某些西古北Mordellidae Latreille, 1802(鞘翅目,Tenebrionoidea)的分类考量
IF 1.3 3区 生物学 Q2 ZOOLOGY Pub Date : 2024-07-12 DOI: 10.3897/zookeys.1207.119398
E. Ruzzier, A. Di Giulio
The present contribution is devoted to the review of some species of Mordellidae belonging to the Italian fauna described by Mariano Zuccarello Patti and Mario Enrico Franciscolo. New taxonomic combinations are proposed: Mediimorda argyropleura (Franciscolo, 1942), comb. nov., Mordella quomoi Franciscolo, 1942, comb. rev., and Mordellokoiles grandii Franciscolo, 1942, stat. nov.Mediimorda argyropleura, Mordella quomoi and Mordellokoiles grandii are considered species that are not part of the Italian fauna. In addition, given the impossibility of identifying the species based on its original description and the destruction of type material, Mordella aradasiana Patti, 1840 is treated as nomen dubium.
本论文主要是对马里亚诺-祖卡雷洛-帕蒂(Mariano Zuccarello Patti)和马里奥-恩里科-弗朗西斯科洛(Mario Enrico Franciscolo)描述的意大利动物区系中的一些莫代尔科物种进行回顾。本文提出了新的分类组合:Mediimorda argyropleura (Franciscolo, 1942), comb.此外,由于无法根据原始描述确定该物种,且模式标本已被毁坏,因此 Mordella aradasiana Patti, 1840 被视为疑似物种。
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引用次数: 0
Systematic revision of the South American “Nuncia” (Opiliones, Laniatores, Triaenonychidae) 南美洲 "Nuncia"(Opiliones, Laniatores, Triaenonychidae)的系统修订
IF 1.3 3区 生物学 Q2 ZOOLOGY Pub Date : 2024-07-12 DOI: 10.3897/zookeys.1207.120068
Willians Porto, Shahan Derkarabetian, Gaston Giribet, A. Pérez-González
The genus Nuncia has long been the most speciose within the Opiliones family Triaenonychidae, comprising 63 species and subspecies distributed across New Zealand and South America. Recent molecular studies utilizing Sanger sequencing and ultraconserved elements (UCEs) have indicated that this genus is not monophyletic, and true Nuncia are actually confined to New Zealand. Here, the morphology of all South American triaenonychids is re-examined and DNA sequence data compiled from three markers (18S rRNA, 28S rRNA and cytochrome c oxidase subunit I) for a large number of triaenonychid species, including specimens from all areas with species currently and formerly classified in Nuncia to reassess their phylogenetic position. Based on our findings we 1) revalidate the genus Chilenuncia (Muñoz-Cuevas, 1971) nom. rest.; 2) describe five new genera: Fresiaxgen. nov., Mistraliagen. nov., Laftrachiagen. nov., Lautariagen. nov., Nerudiellagen. nov.; 3) redescribe five species: Fresiax spinulosacomb. nov., Mistralia verrucosacomb. nov., Chilenuncia chilensiscomb. nov., Chilenuncia rostratacomb. nov., Nerudiella americanacomb. nov.; and 4) describe 22 new species of South American triaenonychids: Fresiax conicasp. nov., Fresiax fraysp. nov., Fresiax mauryisp. nov., Fresiax pichicuysp. nov., Mistralia ramirezisp. nov., Laftrachia robinsp. nov., Lautaria ceacheisp. nov., Nerudiella cachaisp. nov., Nerudiella caramavidasp. nov., Nerudiella cautinsp. nov., Nerudiella choapasp. nov., Nerudiella curisp. nov., Nerudiella goroisp. nov., Nerudiella jaimeisp. nov., Nerudiella mallecosp. nov., Nerudiella pencosp. nov., Nerudiella pichisp. nov., Nerudiella portaisp. nov., Nerudiella quenessp. nov., Nerudiella vilchessp. nov., Nerudiella wekufesp. nov., and Nerudiella zapallarsp. nov. Furthermore, we provide detailed illustrations of all the South American species belonging to these lineages formerly classified in Nuncia.
一直以来,Nuncia属都是Opiliones科Triaenonychidae属中物种最多的,共有63个种和亚种,分布在新西兰和南美洲。最近利用桑格测序和超保守元素(UCEs)进行的分子研究表明,该属并非单系,真正的Nuncia实际上仅限于新西兰。在此,我们重新研究了所有南美洲三尖杉属植物的形态学,并通过三个标记(18S rRNA、28S rRNA 和细胞色素 c 氧化酶亚单位 I)为大量三尖杉属物种编制了 DNA 序列数据,其中包括目前和以前被归入 Nuncia 的物种的所有地区的标本,以重新评估它们的系统发育位置。根据我们的研究结果,我们:1)重新确认了 Chilenuncia 属(Muñoz-Cuevas,1971 年)的名称;2)描述了五个新属:nov.、Mistraliagen.nov.、Laftrachiagen.nov.、Lautariagen.nov.、Nerudiellagen.nov.;3)重新描述了五个物种:nov.、Chilenuncia chilensiscomb.nov.、Chilenuncia rostratacomb.nov.、Nerudiella americanacomb.nov.;以及 4)描述了 22 个南美洲三叶虫新种:Fresiax conicasp.nov.、Fresiax fraysp.nov.、Fresiax mauryisp.nov.、Fresiax pichicuysp.nov.、Mistralia ramirezisp.nov.、Laftrachia robinsp.nov.、Lautaria ceacheisp.nov.、Nerudiella cachaisp.nov.、Nerudiella caramavidasp.nov.、Nerudiella cautinsp.nov.、Nerudiella choapasp.nov、Nerudiella curisp.新种, Nerudiella goroisp.新种, Nerudiella jaimeisp.新种, Nerudiella mallecosp.新种, Nerudiella pencosp.新种, Nerudiella pichisp.新种、nov.、Nerudiella pencosp.nov.、Nerudiella pichisp.nov.、Nerudiella portaisp.nov.、Nerudiella quenessp.nov.、Nerudiella vilchessp.nov.、Nerudiella wekufesp.nov.和 Nerudiella zapallarsp.此外,我们还提供了以前归入 Nuncia 的属于这些种系的所有南美洲物种的详细插图。
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引用次数: 0
Low-cost, high-volume imaging for entomological digitization 用于昆虫学数字化的低成本、高容量成像技术
IF 1.3 3区 生物学 Q2 ZOOLOGY Pub Date : 2024-07-11 DOI: 10.3897/zookeys.1206.123670
Dirk Steinke, Jaclyn T. A. McKeown, Allison Zyba, Joschka McLeod, Corey Feng, P. Hebert
Large-scale digitization of natural history collections requires automation of image acquisition and processing. Reflecting this fact, various approaches, some highly sophisticated, have been developed to support imaging of museum specimens. However, most of these systems are complex and expensive, restricting their deployment. Here we describe a simple, inexpensive technique for imaging arthropods larger than 5 mm. By mounting a digital SLR camera on a CNC (computer numerical control) motor-drive rig, we created a system that captures high-resolution z-axis stacked images (6960 × 4640 pixels) of 95 specimens in 30 minutes. This system can be assembled inexpensively ($1000 USD without a camera) and it is easy to set-up and maintain. By coupling low cost with high production capacity, it represents a solution for digitizing any natural history collection.
自然历史藏品的大规模数字化需要图像采集和处理的自动化。为反映这一事实,人们开发了各种方法来支持博物馆标本的成像,其中一些方法非常复杂。然而,这些系统大多复杂而昂贵,限制了它们的应用。在此,我们介绍一种简单、廉价的技术,用于对大于 5 毫米的节肢动物进行成像。通过将数码单反相机安装在 CNC(计算机数控)电机驱动装置上,我们创建了一个系统,可在 30 分钟内捕捉 95 个标本的高分辨率 Z 轴堆叠图像(6960 × 4640 像素)。该系统组装成本低廉(1000 美元,不含摄像头),易于安装和维护。通过将低成本与高产能结合起来,它为任何自然历史藏品的数字化提供了解决方案。
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引用次数: 0
The tiniest violin: the male of Loxosceles vallenar (Araneae, Sicariidae) 最细小的小提琴:Loxosceles vallenar(鹤形目,蝎科)的雄鸟
IF 1.3 3区 生物学 Q2 ZOOLOGY Pub Date : 2024-07-11 DOI: 10.3897/zookeys.1206.122469
I. Magalhães, H. Iuri, A. D. Brescovit, J. Pizarro-Araya
In recent years, several endemic species of Loxosceles, violin spiders, have been described from the North-Central Chile biodiversity hotspot, some of which have ambiguous placement within the species groups of the genus. In a recent expedition to the Atacama region, we collected male specimens representing new records of two recently described species: Loxosceles vicentei Taucare-Ríos, Brescovit & Villablanca, 2022 and Loxosceles vallenar Brescovit, Taucare-Ríos, Magalhaes & Santos, 2017 (Araneae, Sicariidae). Males of the latter are hitherto unknown and are here described for the first time. Examination of the morphology of these species revealed characters such as an embolic keel and digitiform median receptacles, which suggest they do not belong in the laeta species group, but rather in the spadicea species group, which is briefly re-diagnosed. With carapace lengths smaller than 2 mm, the newly discovered males of L. vallenar are the tiniest members of the genus. In addition, males of this species bear strong macrosetae in the clypeus, a sexually dimorphic character not previously reported in Loxosceles.
近年来,智利中北部生物多样性热点地区描述了几个当地特有的小提琴蜘蛛(Loxosceles)物种,其中一些物种在该属物种组中的位置不明确。在最近一次对阿塔卡马地区的考察中,我们采集到了雄性标本,代表了最近描述的两个物种的新记录:Loxosceles vicentei Taucare-Ríos, Brescovit & Villablanca, 2022 和 Loxosceles vallenar Brescovit, Taucare-Ríos, Magalhaes & Santos, 2017(鹤形目,蝎科)。后者的雄性迄今为止尚不为人所知,本文对其进行了首次描述。对这些物种形态的研究发现了栓龙骨和指状中间托等特征,这表明它们不属于laeta物种组,而是属于spadicea物种组,现对其进行简要的重新诊断。新发现的雄性 L. vallenar 的躯体长度小于 2 毫米,是该属中最小的成员。此外,该物种的雄性鳞片上有很强的大网眼,这是以前在 Loxosceles 中从未报道过的性二态特征。
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引用次数: 0
First record of subgenus Synaldis Foerster (Hymenoptera, Braconidae, Alysiinae, Dinotrema Foerster) from Chile, with description of ten new species 智利首次记录到 Synaldis Foerster 亚属(膜翅目,腕足动物,Alysiinae,Dinotrema Foerster),并描述了 10 个新种
IF 1.3 3区 生物学 Q2 ZOOLOGY Pub Date : 2024-07-10 DOI: 10.3897/zookeys.1206.124515
Franciélle Dias de Oliveira, A. Penteado-Dias
Synaldis is a taxon within the Aspilota group with a contentious taxonomic history, currently classified as a subgenus of the genus Dinotrema. Species of Synaldis were only documented in the Neotropical region in 2017, and until then, the Neotropical fauna of this subgenus was represented by five species from Brazil. In this study, Synaldis is reported for the first time in Chile, with the description and illustration of ten new species, namely: Dinotrema (Synaldis) acarinareolatumsp. nov., D. (S.) brunneumsp. nov., D. (S.) chilensesp. nov., D. (S.) daltonisp. nov., D. (S.) flavumsp. nov., D. (S.) latusdentertiumsp. nov., D. (S.) perisfelipoisp. nov., D. (S.) pilosicaudatumsp. nov., D. (S.) puyehuesp. nov., and D. (S.) veraesp. nov. The studied specimens were collected during expeditions to southern Chile, in the Valdivian temperate rainforest at Parque Nacional de Puyehue. This study also includes a dichotomous identification key for Neotropical species of Synaldis, as well as a discussion of the primary morphological characters used to distinguish species within the Neotropical and Nearctic regions.
Synaldis是Aspilota类群中的一个分类群,其分类历史颇具争议,目前被归类为Dinotrema属的一个亚属。2017 年才在新热带地区记录到 Synaldis 的物种,在此之前,该亚属的新热带动物区系只有来自巴西的 5 个物种。本研究首次报道了智利的 Synaldis,并描述和说明了 10 个新物种,即Dinotrema(Synaldis)acarinareolatumsp.nov.、D. (S.)brunneumsp.nov.、D. (S.)chilensesp.nov.、D. (S.)daltonisp.nov.、D. (S.)flavumsp.nov、D.(S.)latusdentertiumsp.nov.、D.(S.)perisfelipoisp.nov.、D.(S.)pilosicaudatumsp.nov.、D.(S.)puyehuesp.nov.和 D.(S.)veraesp.nov.所研究的标本是在智利南部普耶胡国家公园的瓦尔迪温带雨林中采集的。本研究还包括 Synaldis 新热带物种的二分识别钥匙,以及用于区分新热带和近北极地区物种的主要形态特征的讨论。
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引用次数: 0
A new species of the genus Luciogobius Gill, 1859 (Teleostei, Oxudercidae) from Taiwan. 来自台湾的 Luciogobius Gill, 1859 属新种(Teleostei, Oxudercidae)。
IF 1.3 3区 生物学 Q2 ZOOLOGY Pub Date : 2024-07-09 eCollection Date: 2024-01-01 DOI: 10.3897/zookeys.1206.118757
Kuan-Hsun Chen, Te-Yu Liao

A new species, Luciogobiusopisthoproctus sp. nov., is described based on 18 specimens collected from Daxi Creek (Yilan) and Babian Creek (Taitung) in Taiwan. The new species is characterized by having a yellowish body with scattered spots on the sides, a black blotch on the caudal fin, the absence of free pectoral-fin rays, and more than 40 vertebrae. The new species can be distinguished from congeners by the following combination of characters: AAA distance (anus to anal-fin origin) shorter than twice the body depth at anus, 4.2-7.2% of standard length (SL); pre-anus length 80.0-92.8% of pre-anal-fin length; snout length 39.7-62.7% of AAA distance; abdominal vertebrae 20-22; caudal vertebrae 20-22; first anal-fin pterygiophore usually inserted behind the second haemal spine.

本研究根据在台湾宜兰大溪及台东八面溪所采集的18件标本,描述了一个新种-Luciogobiusopisthoproctus sp.nov.。新种的特征是体色淡黄,体侧有零星斑点,尾鳍上有黑色斑点,无游离胸鳍鳍条,脊椎骨超过 40 节。该新种可通过以下特征组合与同类区分开来:AAA距离(肛门到臀鳍起源)短于肛门处体深的两倍,为标准体长(SL)的 4.2-7.2%;前肛门长度为前臀鳍长度的 80.0-92.8%;吻长为 AAA 距离的 39.7-62.7%;腹椎 20-22;尾椎 20-22;第一臀鳍翼管通常插入第二血棘后方。
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引用次数: 0
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