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Functional morphology of the stem in the Lower Palaeozoic crinoid Macrostylocrinus Hall from Scotland 苏格兰下古生界板鳃亚纲棘皮动物 Macrostylocrinus Hall 茎的功能形态
IF 0.7 4区 地球科学 Q4 GEOLOGY Pub Date : 2023-12-11 DOI: 10.1144/sjg2023-021
Stephen K. Donovan
Two species of the closely-related monobathrid crinoid from the Lower Palaeozoic of Scotland, namely Macrostylocrinus cirrifer Ramsbottom (Upper Ordovician, Katian) and Macrostylocrinus silurocirrifer Brower (Lower Silurian, Telychian), are similar in having elongate, unbranched radices proximally. These were not cirri, as suggested by their names, but were radices which were more or less inflexible, lacking contractile tissues. The function of these radices was uncertain. In the absence of contractile tissues, they could not have been for grasping other upright structures and crinoid do not need help to balance, their posture being maintained by mutable collagenous tissues. It is possible, but unlikely, that they may have acted to direct feeding currents towards the crown. Most probably, in an analogy to the post-Palaeozoic isocrinids, the stem acted like a ‘conveyor belt’, the proximal, radicular and upright part being carried away from the cup as further columnals are inserted, eventually forming a distal, recumbent attachment structure. The elongate radices would have stabilised the dististele, but, unlike isocrinids, the arms of Macrostylocrinus spp. were not adapted for crawling and thus escaping predators. Both M. silurocirrifer (type species) and M. cirrifer are included in Macrostylocrinus ( Scotimacrostylocrinus ) subgen. nov.
苏格兰下古生界的两个物种,即Macrostylocrinus cirrifer Ramsbottom(上奥陶世,卡蒂期)和Macrostylocrinus silurocirrifer Brower(下志留纪,泰利奇期),是与之密切相关的单叶棘皮动物,它们的相似之处在于下部都有细长、不分枝的辐射状突起。正如它们的名字所暗示的那样,这些根状茎并不是卷须,而是或多或少缺乏弹性、缺乏收缩组织的根状茎。这些放射线的功能尚不确定。在没有收缩组织的情况下,它们不可能用来抓住其他直立的结构,而且海龙类不需要帮助来保持平衡,它们的姿势是由可变异的胶原组织维持的。有可能,但可能性不大,它们的作用可能是将食流引向冠部。最有可能的情况是,与后古生代的异长类类似,茎干就像一条 "传送带",当更多的柱状体插入时,近端、放射状和直立的部分就会被带离杯状体,最终形成一个远端、后倾的附着结构。细长的辐射状突起可以稳定远端,但与等轴类不同的是,Macrostylocrinus属的臂不适合爬行,因此无法躲避捕食者。M.silurocirrifer(模式种)和M.cirrifer都被归入Macrostylocrinus ( Scotimacrostylocrinus ) subgen.
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引用次数: 0
Dr John Grant Malcolmson and a reconciliation of the Middle Devonian Lethen Bar and Lethen House fish bearing nodule localities 约翰-格兰特-马尔科姆森博士与中泥盆世勒森巴尔和勒森豪斯含鱼结核地点的核对工作
IF 0.7 4区 地球科学 Q4 GEOLOGY Pub Date : 2023-11-28 DOI: 10.1144/sjg2023-011
R. G. Davidson, N. H. Trewin, J. Armstrong, S. R. Waters
The area known as Lethen Bar and Clune, SE of Nairn, Scotland, is a classic Middle Devonian locality which has yielded nodules or concretions, some of which contain fossil fishes with the highest quality of preservation. The locality was largely centred on farm limestone quarries situated around the perimeter of an isolated outlier of the main fishbed. It was first described in the 19 th century, although the upsurge in collecting fossil fishes only occurred some twelve years after the quarries were first mentioned in the scientific literature. Our knowledge of the provenance of the locality is based on very limited accounts, which have never been scientifically tested; these accounts also contain apparent anomalies that have never been adequately addressed. Based on these anomalies, the author of a paper published in 1983 proposed that the locality had been untraceable since the late 19 th century and that the outcrop was quarried out. In 2005, the present authors recorded the first scientifically detailed stratigraphical section of the fishbed, followed in 2021-2023 by detailed field surveys and by a re-appraisal of the literature. This work has resolved the 19 th century anomalies and enabled us to confirm the locations of old quarries, to give affirmation to two previously unrecognised sites, and to show that the outcrop is still present. Thematic collection: This article is part of the Geology of Scotland collection available at: https://www.lyellcollection.org/topic/collections/geology-of-scotland
位于苏格兰奈恩东南部的莱森巴尔和克鲁恩地区是典型的中泥盆纪地点,这里出产结核或凝块,其中一些含有保存质量极高的鱼类化石。该地点主要以农场石灰岩采石场为中心,这些采石场位于主鱼床一个孤立的外围。尽管在采石场首次被科学文献提及约 12 年后,收集鱼类化石的热潮才兴起,但 19 世纪就首次对该地点进行了描述。我们对该地点来源的了解是基于非常有限的描述,这些描述从未经过科学检验;这些描述还包含明显的异常现象,但从未得到充分解决。基于这些异常现象,1983 年发表的一篇论文的作者提出,该地点自 19 世纪晚期以来就已无迹可寻,露头已被采光。2005 年,本文作者首次对鱼床进行了科学详细的地层剖面记录,随后在 2021-2023 年进行了详细的实地调查,并对文献进行了重新评估。这项工作解决了 19 世纪的异常现象,使我们能够确认旧采石场的位置,确认两个以前未确认的遗址,并表明露头仍然存在。 专题文集: 本文是《苏格兰地质》文集的一部分,可从以下网址获取: https://www.lyellcollection.org/topic/collections/geology-of-scotland
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引用次数: 0
A revision of the ‘coelophysoid-grade’ theropod specimen from the Lower Jurassic of the Isle of Skye (Scotland) 苏格兰斯凯岛下侏罗纪“腔体级”兽脚亚目标本的修订
4区 地球科学 Q4 GEOLOGY Pub Date : 2023-11-09 DOI: 10.1144/sjg2023-012
Martín D. Ezcurra, Daniel Marke, Stig A. Walsh, Stephen L. Brusatte
The broadest diversification of early predatory dinosaurs is represented by the ‘coelophysoid-grade’ neotheropods, but their Hettangian–Sinemurian ( c. 191–201 Ma) record is scarce worldwide. More information is needed to shed light on the evolution of this dinosaur group after the end-Triassic mass extinction ( c. 201 Ma). Here we revisit the anatomy and phylogeny of one of these earliest Jurassic neotheropod specimens, an isolated partial tibia from the lower Sinemurian of the Isle of Skye (Scotland) that was previously identified as probably closely related to Liliensternus liliensterni and coelophysids. However, we found that the Skye specimen is positioned in the branch leading to Averostra (Ceratosauria + Tetanurae), in a polytomy with Sarcosaurus woodi from the late Hettangian–lower Sinemurian of central England and a clade composed of Tachiraptor admirabilis and Averostra. The morphology of the Skye specimen is congruent with that of referred specimens of Sarcosaurus woodi , but because it probably represents a skeletally immature specimen, we assign it to cf. Sarcosaurus woodi . The Skye specimen increases the number of averostran-line neotheropod specimens recorded in the Lower Jurassic of Europe and current evidence indicates that these forms, and not coelophysoids, were relatively common in this part of the world at that time. Supplementary material : Character list and dataset for phylogenetic analysis and 3D model of the tibia are available at https://doi.org/10.6084/m9.figshare.c.6863016 Thematic collection: This article is part of the Palaeontology of Scotland collection available at: https://www.lyellcollection.org/topic/collections/palaeontology-of-scotland
早期掠食性恐龙最广泛的多样性是“腔骨类”新足类,但它们的hettangan - sinemurian(约191-201 Ma)记录在世界范围内是罕见的。在三叠纪末大灭绝(c. 201 Ma)之后,需要更多的信息来阐明这一恐龙群的进化。在这里,我们重新审视了这些最早的侏罗纪新足类标本之一的解剖和系统发育,这是一块来自苏格兰斯凯岛下Sinemurian的分离的部分胫骨,之前被认为可能与Liliensternus liliensterni和腔体类密切相关。然而,我们发现Skye标本与来自英格兰中部Hettangian-lower Sinemurian的woodsarcosaurus和一个由速盗龙和Averostra组成的分支(角鼻龙+破伤风科)位于一个分支中。Skye标本的形态与参考标本一致,但由于它可能代表一个骨骼未成熟的标本,我们将其定为cf. Sarcosaurus woodi。Skye标本增加了在欧洲下侏罗纪记录到的averostranline新足类标本的数量,目前的证据表明,这些形式,而不是腔骨类,在当时的世界这一地区相对常见。补充资料:用于系统发育分析的特征列表和数据集以及胫骨的3D模型可在https://doi.org/10.6084/m9.figshare.c.6863016上获得。主题集合:本文是苏格兰古生物学集合的一部分,可在https://www.lyellcollection.org/topic/collections/palaeontology-of-scotland上获得
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引用次数: 0
Arthropleura trackway ( Diplichnites cuithensis ) from the Carboniferous, Serpukhovikan, Limestone Coal Formation, Clackmannan Group, Linn Park, Glasgow 格拉斯哥林恩公园Clackmannan群石炭纪Serpukhovikan石灰岩煤组的关节胸膜足迹(Diplichnites cuithensis)
4区 地球科学 Q4 GEOLOGY Pub Date : 2023-11-06 DOI: 10.1144/sjg2021-019
James O. Buckman, Simon J. Cuthbert, Paul G. Polson
The Arthropleurid trackway Diplichnites cuithensis has previously been described from Scotland from the Upper Carboniferous Serpukhovikan Limestone Coal Formation on the Isle of Arran, and the Lower Carboniferous Visean Pittenweem and Anstruther Formations on the East Fife coast. Here we describe a new west coast single trackway from the Serpukhovikan Limestone Coal Formation of Glasgow's, Linn Park. The trackway occurs associated with simple horizontal burrows assignable to Planolites? , vertical openings of Arenicolites, examples of Taenidium barretti (formerly Beaconites barretti ), and irregular large scale bioturbation or possibly rootlet casts. The trace fossils and sedimentary structures (including trough cross-bedding and flaser bedding) indicate a fluvial sandbar or plain environment, possibly of estuarine origin, locally colonized by plants. Diplichnites cuithensis (and other Diplichnites species) commonly occur associated with the burrow Taenidium barretti . The latter is known to have been widespread globally throughout the Carboniferous, and is a common component of fluviatile sequences within the Lower Carboniferous succession of NW Ireland. This suggests that previously undocumented older Scottish Carboniferous examples of both Diplichnites ichnospecies and Taenidium barretti may also be present, assuming that suitable environments persisted and are currently adequately exposed.
先前在苏格兰的Arran岛的上石炭世Serpukhovikan石灰石煤组和East Fife海岸的下石炭世Visean Pittenweem和Anstruther组中描述过Arthropleurid trackway Diplichnites cuithensis。在这里,我们描述了一条新的西海岸单轨,来自格拉斯哥的林恩公园的Serpukhovikan石灰石煤组。这条轨迹与Planolites的简单水平洞穴有关?,砂粒石的垂直开口,Taenidium barretti(以前的Beaconites barretti)的例子,不规则的大规模生物扰动或可能的根状铸造。遗迹化石和沉积构造(包括槽状交错层理和闪光层理)表明其为河流沙洲或平原环境,可能起源于河口,局部被植物殖民。cuithensis(和其他Diplichnites物种)通常与穴居Taenidium barretti相关。众所周知,后者在整个石炭世的全球范围内广泛存在,并且是西北爱尔兰下石炭世演替中的流动层序的共同组成部分。这表明,假设合适的环境持续存在,并且目前已充分暴露,那么以前未记载的苏格兰石炭纪更古老的双长石属鱼种和巴雷蒂Taenidium barretti也可能存在。
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引用次数: 0
Dipnoan diversity in the early Pennsylvanian of Scotland: new lungfish from the Lower Coal Measures of North Lanarkshire 苏格兰宾夕法尼亚早期的肺鱼多样性:来自北拉纳克郡下煤系的新肺鱼
IF 0.7 4区 地球科学 Q4 GEOLOGY Pub Date : 2023-07-06 DOI: 10.1144/sjg2023-006
Francis M. Elliott, T. Challands, T. Smithson
Only two lungfish have been recorded in the Scottish Coal Measures in the past one hundred and fifty years: Ctenodus and Sagenodus . Here we describe a suite of new lungfish specimens collected from sites in the Scottish Central Coalfield that represent a least four taxa: Sagenodus ; Conchopoma ; and two new forms Braccodus kerri gen. et sp. nov and Lanarkodus clarki gen. et sp. nov. These are part of an extensive vertebrate fauna recently discovered in colliery waste from mining the Upper and Lower Drumgray Coal. These coals lie within the Communis Chronozone and are of Langsettian age. The specimens are much smaller than those found previously in the Scottish Coal Measures and represent fish between 60 and 300 mm long. The basihyal tooth plates of Conchopoma are the first record of this genus in the Pennsylvanian of Europe. Lanarkodus clarki has a heterodont dentition not previously described from the Pennsylvanian. All the new material is preserved in thin, laminated shales, suggesting a small lake environment rather than the typical coal swamp. These new discoveries demonstrate that Pennsylvanian lungfish were more diverse than previously realised and add to growing evidence that the rate of lungfish evolution did not decline significantly after the Devonian, and remained high throughout the Carboniferous. Thematic collection: This article is part of the The Palaeontology of Scotland collection available at: https://www.lyellcollection.org/topic/collections/palaeontology-of-scotland
在过去的150年里,苏格兰煤系中只记录了两种肺鱼:Ctenodus和Sagenodus。在这里,我们描述了从苏格兰中央煤田采集的一组新的肺鱼标本,这些标本至少代表了四个分类群:Sagenodus;Conchopoma;以及两种新形式的Braccodus kerri gen.et sp.nov和Lanarkodus clarki gen.et sp.nov。它们是最近在德拉姆格雷上下煤矿开采的煤矿废物中发现的广泛脊椎动物群的一部分。这些煤位于Communis时间带内,属于朗赛特时代。这些标本比之前在苏格兰煤系中发现的标本小得多,代表了60至300毫米长的鱼类。Conchopoma的基牙板是该属在欧洲宾夕法尼亚纪的第一个记录。克拉基拉纳科杜斯有一个宾夕法尼亚人以前没有描述过的异齿列。所有新材料都保存在薄层页岩中,这表明这里是一个小湖泊环境,而不是典型的煤沼泽。这些新发现表明,宾夕法尼亚州的肺鱼比以前意识到的更为多样,并为越来越多的证据提供了证据,证明肺鱼的进化率在泥盆纪之后没有显著下降,在整个石炭纪都保持较高水平。主题收藏:本文是苏格兰古生物学收藏的一部分,可在以下网站获取:https://www.lyellcollection.org/topic/collections/palaeontology-of-scotland
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引用次数: 0
On a new species of Rhizodopsis from the Carboniferous of Scotland 苏格兰石炭纪一新种根瘤菌的研究
IF 0.7 4区 地球科学 Q4 GEOLOGY Pub Date : 2023-07-05 DOI: 10.1144/sjg2023-008
Francis M. Elliott
A new species of Rhizodopsis is described from material recovered from several Carboniferous locations within the Midland Valley of Scotland. Individual dermal skull bones were obtained from derelict coal waste tips at Wester Bracco, North Lanarkshire from shales originally overlying the Drumgray Coal. Individual dermal skull bones are well preserved, allowing for detailed description and reconstructions of the skull roof and cheek. Rhizodopsis rankini sp. nov. is distinguished by the presence of a lobed opercular, a deeply embayed postparietal shield between the supratemporal and tabular bones, and enlarged lateral extrascapulars. Rhizodopsis is here included in the Megalichthyidae, and an emended diagnosis of the family is given. The composition of the genus Rhizodopsis is reviewed. Except for R hizodopsis hanbuchi, Rhizodopsis rankini and Rhizodopsis sauroides , all other species are based solely on scales. The validity of these scale-based species is reassessed and all but Rhizodopsis granulatus and Rhizodopsis mazonius are considered to be valid species. Thematic collection: This article is part of the The Palaeontology of Scotland collection available at: https://www.lyellcollection.org/topic/collections/palaeontology-of-scotland
从苏格兰米德兰山谷的几个石炭纪地点回收的材料中描述了一种新的根茎opsis。单个真皮颅骨是从北拉纳克郡韦斯特布拉科的废弃煤渣尖上获得的,这些煤渣来自最初覆盖在Drumgray煤上的页岩。单个真皮颅骨保存完好,允许详细描述和重建头骨顶部和脸颊。兰氏根opsis rankini sp. 11 .的特征是在颞骨和板骨之间有裂片状的眼状、深嵌的顶骨后盾状和增大的外侧肩胛外肌。根茎藓属在这里包括在巨型鱼纲,并修订诊断的家庭是给出。综述了根霉属植物的组成。除汉布氏根瘤菌、兰氏根瘤菌和索氏根瘤菌外,其余种均以鳞片为基础。对这些基于尺度的种的有效性进行了重新评估,除细粒根opsis和马祖根opsis外,其余种均被认为是有效种。专题收藏:这篇文章是苏格兰古生物学收藏的一部分,可在:https://www.lyellcollection.org/topic/collections/palaeontology-of-scotland
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引用次数: 0
The stable isotope (C,O,S) record of Palaeoproterozoic marbles, Scotland 苏格兰古元古代大理岩的稳定同位素(C,O,S)记录
IF 0.7 4区 地球科学 Q4 GEOLOGY Pub Date : 2023-07-05 DOI: 10.1144/sjg2023-009
J. Parnell, A. Boyce, J. Armstrong, A. Schito, D. Muirhead
Palaeoproterozoic marbles occur widely in North West Scotland. The isotopically heavy carbonate carbon (δ 13 C >3 ‰) in marbles that characterizes the worldwide Lomagundi-Jatuli Event (2.3 to 2.05 Ga) is recognized in each of the Laurentian Foreland, the Moine Nappe and the Sgurr Beag Nappe, over a 150 km transect across the Caledonian thrust belt. A light oxygen isotope composition distinguishes marbles which have been sheared and retrogressed by ingress of meteoric water, possibly during both Laxfordian and Caledonian orogenesis. The shearing of marbles also contributed to graphite formation (mean δ 13 C -7.2 ‰). Pyrite in the marbles contains isotopically heavy sulphur, typical of Palaeoproterozoic diagenetic sulphides precipitated from low-sulphate seawater. These data show that the ∼2 Ga marbles in Scotland are a high-quality archive of information on their depositional and post-depositional history. The data emphasize a continuum of a Palaeoproterozoic marble-graphite-sulphide bearing assemblage from Eastern Canada and Greenland through Scotland to Scandinavia.
古元古代大理石广泛分布于苏格兰西北部。大理岩中的同位素重碳酸盐碳(δ13C>3‰)是世界范围内Lomagundi-Jatuli事件(2.3至2.05 Ga)的特征,在整个喀里多尼亚逆冲带150公里的横断面上,在劳伦阶前陆、莫因纳佩和斯古尔比格纳佩各有发现。轻氧同位素组成区分了可能在拉克斯福德造山运动和喀里多尼亚造山运动期间,由于大气降水的进入而剪切和后退的大理石。弹珠的剪切也有助于石墨的形成(平均δ13C-7.2‰)。大理岩中的黄铁矿含有同位素重硫,这是典型的古元古代成岩硫化物,由低硫酸盐海水沉淀而成。这些数据表明,苏格兰的~2Ga大理石是关于其沉积和沉积后历史的高质量信息档案。这些数据强调了从加拿大东部和格陵兰岛到苏格兰再到斯堪的纳维亚半岛的古元古代大理岩-石墨-硫化物组合的连续性。
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引用次数: 0
Trace element minerals from carbonatite-related fluids, The Aird, Scotland 来自碳酸盐岩相关流体的微量元素矿物,爱尔兰,苏格兰
IF 0.7 4区 地球科学 Q4 GEOLOGY Pub Date : 2023-06-02 DOI: 10.1144/sjg2021-015
E. Heptinstall, J. Parnell, A. Boyce, J. Still
Fenitised rocks in Neoproterozoic country rocks adjacent to the Great Glen Fault, Scotland, contain calcite veins with mineral inclusions. The minerals include the rare earth vanadate wakefieldite, sulphur-rich monazite, scandium-rich ilmenite, and manganese oxides rich in barium and lead. Each of these phases is recorded in carbonatites elsewhere. An affinity with carbonatites is further suggested by the carbon isotope composition of the calcite veins, and anomalously high levels of manganese and strontium in the veins. These data add weight to a model in which Scotland northwest of the Great Glen Fault was affected by carbonatite-related fluids during the Caledonian Orogeny. Thematic collection: This article is part of the Early Career Research collection available at: https://www.lyellcollection.org/topic/collections/early-career-research
苏格兰大格伦断裂带附近新元古代乡村岩石中的粉碎化岩石含有方解石脉和矿物包裹体。这些矿物包括稀土钒酸钠矿、富硫独居石、富钪钛铁矿和富钡、富铅的锰氧化物。这些相中的每一个都在别处的碳酸盐岩中有记录。方解石脉的碳同位素组成和脉中异常高水平的锰和锶进一步表明方解石与碳酸盐的亲和关系。这些数据为大格伦断层西北部的苏格兰在加里东造山运动期间受到碳酸盐岩相关流体影响的模型增加了分量。主题收集:本文是早期职业研究收集的一部分:https://www.lyellcollection.org/topic/collections/early-career-research
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引用次数: 0
Aspects of Aktuo-Paläontologie of the rocky beaches of the eastern Isle of Mull, UK 英国马尔岛东部岩石海滩Aktuo-Paläontologie的各个方面
IF 0.7 4区 地球科学 Q4 GEOLOGY Pub Date : 2023-04-18 DOI: 10.1144/sjg2023-004
S. Donovan
Conglomerates are, commonly, only poorly fossiliferous at best. Yet beaches with common lithic clasts can be used to model the taphonomy of fossils in conglomeratic settings. Four beaches on the east coast of the Isle of Mull, Inner Hebrides, are clast-rich, with lithic pebbles, cobbles and boulders, but poor in shells, many of which are poorly preserved. There is ample evidence of shells being bored and encrusted, yet many or most of these were infested after death of the host. Of the ‘boring trinity’ – Caulostrepsis Clarke, Entobia Bronn and Gastrochaenolites Leymerie – so typical of the Trypanites Ichnofacies around Britain's coasts, only the last ichnogenus was not present, most likely due to the absence of suitable mobile substrates (such as limestone cobbles and oysters). Encrusters including Balanus , serpulids and spirorbids, show different patterns of preservation, probably due to multiple factors. Bored wood ( Apectoichnus ) was only found at one locality, which may be due to hydrodynamic sorting. Whelk shells show a range of patterns of breakage, most probably due to mechanical damage. But conglomerates commonly preserve fossil snails either complete or not at all. The results from these sites suggest that they represent an intermediate condition rarely preserved in the rock record.
一般来说,砾岩最多只能形成很差的化石。然而,具有共同岩屑碎屑的海滩可以用来模拟砾岩环境中的化石地貌学。内赫布里底群岛(Inner Hebrides)马尔岛(Isle of Mull)东海岸的四个海滩上,碎屑丰富,有鹅卵石、鹅卵石和巨石,但贝壳很少,其中许多保存得很差。有充分的证据表明,这些贝壳被钻穿和结壳,但其中许多或大部分在寄主死后被侵染。在“无聊的三位一体”——Caulostrepsis Clarke, Entobia Bronn和Gastrochaenolites Leymerie——是英国海岸周围典型的锥虫鱼相,只有最后一个鱼属没有出现,很可能是由于缺乏合适的可移动基质(如石灰石鹅卵石和牡蛎)。包括Balanus, serpulids和spirorbids在内的壳体呈现出不同的保存模式,可能是由于多种因素的影响。钻孔木(Apectoichnus)只在一个地方发现,这可能是由于水动力分选。海螺壳呈现出一系列破碎的模式,很可能是由于机械损伤。但是,砾岩通常保存完整的蜗牛化石,或者根本不保存。这些遗址的结果表明,它们代表了一种中间状态,很少在岩石记录中保存下来。
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引用次数: 0
New postcranial remains from the Lealt Shale Formation of the Isle of Skye, Scotland, showcase hidden pterosaur diversity in the Middle Jurassic 苏格兰斯凯岛Lealt页岩组的新颅后遗迹展示了侏罗纪中期隐藏的翼龙多样性
IF 0.7 4区 地球科学 Q4 GEOLOGY Pub Date : 2023-04-18 DOI: 10.1144/sjg2023-001
N. Jagielska, T. Challands, Michael O'Sullivan, D. Ross, N. Fraser, M. Wilkinson, S. Brusatte
The transition between Early to Middle Jurassic was significant in pterosaur evolution, when these volant reptiles exploded in diversity alongside dinosaurs and other animals. It has long been thought, however, that pterosaurs did not develop large wingspans until after the Jurassic, a notion challenged by the recent discovery of Dearc sgiathanach in the Bathonian-aged Lealt Shale Formation of the Isle of Skye, Scotland, whose holotype specimen had an estimated wingspan greater than 2.5 meters. We here report the discovery of a new pterosaur specimen from the Lealt Shale Formation, comprising a tibiotarsus, metatarsal, pedal phalanges, and caudal vertebrae. The elongate tail vertebrae with ossified processes indicate the specimen is a non-pterodactyloid pterosaur, albeit its fragmentary nature makes it difficult to determine whether it belongs to a new taxon. Its metatarsal and caudal vertebrae are considerably larger than corresponding bones in the Dearc holotype, indicating that it belonged to an even larger individual, thus demonstrating that pterosaurs with broad wingspans were not anomalous in the Middle Jurassic. The growing Middle Jurassic pterosaur record of Scotland and England, although mostly represented by isolated and fragmentary fossils, reveals a high diversity of clades, long obscured by the lack of well-preserved skeletons.
早侏罗纪到中侏罗纪之间的过渡在翼龙的进化中意义重大,当时这些volant爬行动物与恐龙和其他动物一起出现了多样性。然而,长期以来人们一直认为,翼龙直到侏罗纪之后才发展出大的翼展,最近在苏格兰斯凯岛巴通尼亚时代的Lealt Shale组发现的Dearc sgiathanach对这一概念提出了质疑,该翼龙的正模标本估计翼展超过2.5米。我们在这里报告了在Lealt Shale组发现的一个新的翼龙标本,包括胫骨、跖骨、脚趾骨和尾椎骨。具有骨化突的细长尾椎骨表明该标本是一种非翼龙类翼龙,尽管其零碎的性质使其难以确定是否属于一个新的分类单元。它的跖骨和尾椎骨比Dearc正模标本中的相应骨骼大得多,这表明它属于一个更大的个体,从而表明翼展较宽的翼龙在侏罗纪中期并不异常。苏格兰和英格兰不断增长的中侏罗纪翼龙记录,尽管主要以孤立和零碎的化石为代表,但揭示了分支的高度多样性,长期以来由于缺乏保存完好的骨骼而被掩盖。
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Scottish Journal of Geology
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