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A security game approach for strategic conservation against poaching considering food web complexities 考虑到食物网的复杂性,一种针对偷猎的战略保护的安全博弈方法
IF 3.5 3区 环境科学与生态学 Q2 ECOLOGY Pub Date : 2021-12-01 DOI: 10.1016/j.ecocom.2021.100970
Zohreh S. Gatmiry , Ashkan Hafezalkotob , Morteza Khakzar bafruei , Roya Soltani

Mitigating the poaching pressure on food webs under multiple constraints (including financial and ecological ones) remains an open problem within conservation. Within this field, mathematically modeling the effects of poaching threats on managerial decision-making is a novel approach. To fill this scientific gap, the present paper uses a security game approach to model the interactions between an environmental manager (defender) and a group of profit-seeking pursuit poachers (attackers) who target species which are nodes of the food web. Based upon the non-cooperative Stackelberg game, the objective of the defender (as leader) is to keep the food web at or near equilibrium through optimally manipulating the populations of an optimal subset of species. In contrast, each attacker strives to maximize monetary profit by hunting an optimal population size of the selected species. The model is validated by a numerical example examining the food web of the endangered Persian leopard (Panthera pardus saxicolor), which lives in Golestan National Park (GNP), Iran. The model provides an overarching biotic intervention strategy to keep the (1 predator-4 prey) food web near equilibrium, while only 2 prey species (the urial (Ovis vignei) and the red deer (Cervus elaphus)) are directly threatened by poachers. The examination revealed that both species population data and poaching data should be taken into account to set effective multi-species conservation prioritization levels. In a sensitivity analysis approach, it was found that, despite the fact that red deer is endangered and preferred by poachers, the deterrent penalty measure should be 1.5 times greater for poaching urial than red deer. The output analysis illustrated that, in order to bring the urial deterrent penalty measure closer to the red deer one, enforcement measures should be about 2.5 times stricter for poaching urial than red deer. The results specifically yield insight into how to optimally conserve a food web equilibrium under poaching pressure and within several constraints.

在多重限制(包括财政和生态限制)下,减轻偷猎对食物网的压力仍然是保护工作中的一个悬而未决的问题。在这一领域,用数学方法模拟偷猎威胁对管理决策的影响是一种新颖的方法。为了填补这一科学空白,本文使用安全博弈方法来模拟环境管理者(保护者)和一群追求利润的偷猎者(攻击者)之间的相互作用,偷猎者以食物网的节点物种为目标。基于非合作的Stackelberg博弈,防御者(作为领导者)的目标是通过最优操纵物种的最优子集的数量来保持食物网处于或接近平衡。相比之下,每个攻击者都努力通过狩猎选定物种的最佳种群规模来最大化货币利润。该模型通过对生活在伊朗Golestan国家公园(GNP)的濒危波斯豹(Panthera pardus saxicolor)食物网的数值分析得到了验证。该模型提供了一个总体的生物干预策略,以保持(1捕食者-4猎物)食物网接近平衡,而只有2种猎物物种(乌拉尔(Ovis vignei)和马鹿(Cervus elaphus))受到偷猎者的直接威胁。研究表明,在确定有效的多物种保护优先级时,应同时考虑物种种群数据和偷猎数据。通过敏感性分析方法发现,尽管马鹿是濒危物种,也是偷猎者的首选,但对偷猎马鹿的威慑惩罚措施应该是马鹿的1.5倍。输出分析表明,为了使尿液威慑处罚措施更接近马鹿,偷猎尿液的执法措施应该比马鹿严格2.5倍左右。结果特别产生洞察如何最佳地保护偷猎压力下的食物网平衡和在几个限制。
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引用次数: 2
Explicit solution of fractional order atmosphere-soil-land plant carbon cycle system 分数阶大气-土壤-土地植物碳循环系统的显式解
IF 3.5 3区 环境科学与生态学 Q2 ECOLOGY Pub Date : 2021-12-01 DOI: 10.1016/j.ecocom.2021.100966
Tanfer Tanriverdi , Haci Mehmet Baskonus , Adnan Ahmad Mahmud , Kalsum Abdulrahman Muhamad

In this paper, the dynamical behaviours and mathematics of the fractional order atmosphere-soil-land plant carbon cycle system involving the time dependent variable of carbon flux in atmosphere, the carbon flux of soil, and the carbon flux of animals and plants are qualitatively and numerically investigated. Explicit solutions in terms of the Mittag-Leffler functions to the terrestrial carbon cycle system around the equilibrium point are first time reported by applying Laplace transform of Caputo fractional derivative. The graphs of obtained solutions the time dependent variable of carbon flux in atmosphere, the carbon flux of soil and the carbon flux of animals and plants are plotted against each other. Explicit solutions to original system and stability of the fractional order linearized system around the equilibrium point are graphically compared as well.

本文以大气碳通量、土壤碳通量和动植物碳通量为时变变量,对分数阶大气-土壤-土地-植物碳循环系统的动力学行为和数学进行了定性和数值研究。利用Caputo分数阶导数的拉普拉斯变换,首次报道了平衡点附近陆地碳循环系统的Mittag-Leffler函数的显式解。得到了大气碳通量、土壤碳通量和动植物碳通量随时间变化的解的相互关系图。并对原系统的显式解和分数阶线性化系统在平衡点附近的稳定性进行了图解比较。
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引用次数: 18
The fox who cried wolf: A keywords and literature trend analysis on the phenomenon of mesopredator release 喊狼来了的狐狸:中掠食者释放现象的关键词及文献趋势分析
IF 3.5 3区 环境科学与生态学 Q2 ECOLOGY Pub Date : 2021-12-01 DOI: 10.1016/j.ecocom.2021.100963
Laura Saggiomo , Valentina Bar , Bruno Esattore

Human activities severely impact the distribution and behaviour of apex predators in numerous terrestrial and aquatic ecosystems, with cascading effects on several species. Mesopredator outbreaks attributable to the removal of an apex predator have often been recorded and described in the literature as “mesopredator release”. During recent decades several examples of the phenomenon have been observed and studied in many different parts of the world. In this paper, we quantitatively reviewed the existing literature on mesopredator release using two software packages (VOSviewer and CiteSpace) to investigate patterns and trends in author keywords through occurrences and temporal analyses, and creating relative network maps. The results showed that even though the general scientific interest in mesopredator release has increased in recent decades, the vast majority of studies focus on canid species, leaving many other species or entire taxa (e.g., reptiles) understudied and under-described. The connection between invasive species and mesopredator release has only recently been more extensively explored and also the effects of apex predators declining in aquatic ecosystems are still only partially investigated. Due to the increasing effect of biological invasions, overfishing, and either the decline or the rise of apex predators in different parts of the world, we expect an even higher increase in interest and number of published documents on the subject. We also encourage widening the research focus beyond canids to include other important taxa.

人类活动严重影响了许多陆地和水生生态系统中顶端捕食者的分布和行为,并对一些物种产生了级联效应。由于顶端捕食者的移除而导致的中掠食者爆发经常被记录并在文献中描述为“中掠食者释放”。近几十年来,在世界上许多不同的地方观察和研究了这种现象的几个例子。本文利用VOSviewer和CiteSpace两种软件,通过对作者关键词的频次分析和时间分析,对现有文献进行了定量梳理,并绘制了相关网络图。结果表明,尽管近几十年来对中掠食性动物释放的普遍科学兴趣有所增加,但绝大多数研究都集中在犬科动物身上,而对许多其他物种或整个分类群(如爬行动物)的研究和描述不足。入侵物种与中捕食者释放之间的关系直到最近才得到更广泛的探索,而且对水生生态系统中顶端捕食者减少的影响也只进行了部分研究。由于生物入侵、过度捕捞以及世界各地顶级食肉动物的减少或增加的影响越来越大,我们预计对这一主题的兴趣和发表的文件数量会有更高的增长。我们也鼓励将研究重点从犬科扩展到其他重要的分类群。
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引用次数: 1
Study on the long-term and short-term effects of globalization and population aging on ecological footprint in OECD countries 经济合作与发展组织国家经济全球化和人口老龄化对生态足迹的长期和短期影响研究
IF 3.5 3区 环境科学与生态学 Q2 ECOLOGY Pub Date : 2021-09-01 DOI: 10.1016/j.ecocom.2021.100946
Xiyue Yang , Nan Li , Hailin Mu , Ming Zhang , Jingru Pang , Mahmood Ahmad

Population aging has become a global phenomenon. Whereas, the ecological consequences of population aging are rarely addressed in current research. In this context, this study contributes to the existing literature by providing new empirical evidence on how population aging along with globalization, economic growth, energy consumption, natural resource rent, and human capital affect ecological footprint for selected 27 Organization for Economic Cooperation and Development (OECD) countries during 1970–2017. This study utilizes an advanced econometric approach, Pooled Mean Group (PMG) estimator for empirical estimation, that allows heterogeneity in the slope parameters and dependencies across countries. The long-term results disclose that globalization (overall) decreases the ecological footprint. On the other hand, financial and political globalization poses a favorable impact on environmental quality, while economic and social globalization is found to increase environmental degradation. Population aging has a statistically significant negative effect on the ecological footprint, but its non-linear term increases the ecological footprint. Additionally, economic growth, energy consumption, and natural resource rent exacerbate environmental deterioration. In contrast, human capital decreases ecological footprint. Based on the empirical results, important policy implications have been provided.

人口老龄化已经成为一个全球性的现象。然而,目前的研究很少涉及人口老龄化的生态后果。在此背景下,本研究通过对1970-2017年27个经济合作与发展组织(OECD)成员国人口老龄化与全球化、经济增长、能源消耗、自然资源租金和人力资本对生态足迹的影响提供新的实证证据,对现有文献做出了贡献。本研究采用了一种先进的计量经济学方法,即混合平均组(PMG)估计器进行经验估计,允许各国斜率参数和依赖关系的异质性。长期结果表明,全球化(总体上)减少了生态足迹。另一方面,金融和政治全球化对环境质量产生了有利影响,而经济和社会全球化则加剧了环境恶化。人口老龄化对生态足迹具有显著的负向影响,但其非线性项增加了生态足迹。此外,经济增长、能源消耗和自然资源租金加剧了环境恶化。相反,人力资本减少了生态足迹。在实证结果的基础上,提出了重要的政策启示。
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引用次数: 38
Bubbling and hydra effect in a population system with Allee effect 具有Allee效应的种群系统中的气泡和水螅效应
IF 3.5 3区 环境科学与生态学 Q2 ECOLOGY Pub Date : 2021-09-01 DOI: 10.1016/j.ecocom.2021.100939
Koushik Garain , Partha Sarathi Mandal

We present a continuous time predator-prey model and predator’s growth subjected to component Allee effect. The model also includes density dependent mortality of predator. We investigate our model both analytically and numerically, and highlighted the effect of density independent mortality and Allee effect. In our system, we find that a fixed point representing the extinction of predator is always a stable point. When coexistence equilibria exists our system is bistable. We have observed that tristability is possible for our model that includes two stable co-existence fixed point. The most important phenomena which we have observed are hydra effect and cascading effect. Due to component Allee effect in predator the system shows multiple hydra effect. We discuss the phenomenon of bubbling, which indicates increasing and decreasing of amplitudes of cycles. We have presented one-parametric as well as two-parametric bifurcation diagram and also all possible bifurcations that the system could go through.

我们提出了一个连续时间捕食者-被捕食者模型,以及受分量Allee效应影响的捕食者的生长。该模型还考虑了捕食者的密度依赖性死亡率。我们对我们的模型进行了分析和数值研究,并强调了密度无关死亡率和Allee效应的影响。在我们的系统中,我们发现一个表示捕食者灭绝的不动点总是一个稳定点。当共存平衡存在时,系统是双稳态的。我们已经观察到包含两个稳定共存不动点的模型是可能的三稳定性。我们观察到的最重要的现象是水螅效应和级联效应。由于捕食者中存在分量折线效应,系统呈现多重水螅效应。我们讨论了鼓泡现象,它表示周期振幅的增减。我们给出了单参数分岔图和双参数分岔图以及系统可能经历的所有可能的分岔图。
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引用次数: 8
Ecosystemology: A new approach toward a taxonomy of ecosystems 生态系统:研究生态系统分类学的新方法
IF 3.5 3区 环境科学与生态学 Q2 ECOLOGY Pub Date : 2021-09-01 DOI: 10.1016/j.ecocom.2021.100945
Bruno Senterre , Porter P. Lowry II , Ehoarn Bidault , Tariq Stévart

Over the last several years, the IUCN Red List approach for assessing the risk of extinction faced by species has been adapted into a Red List of Ecosystems methodology. This endeavor faces several important challenges, including how to define the types of ecosystems to which the Red List criteria are applied, and how to manage information on the geographic distribution of ecosystems in an open, transparent, and standardized manner linking mapping, typology, and field studies. We propose a fundamentally novel approach that differs from currently available ecosystem typologies in three important aspects by (1) offering a new way of conceptualizing types of ecosystems, (2) providing an explicit method for communicating the conceptualized ecosystems and how they are circumscribed, and (3) developing technical tools for managing the resulting conceptual model. Firstly, ecosystem types are defined by studying biogeoclimatic gradients using an approach that is both modular (in which combinations of ecological factors are studied at a given scale) and hierarchical (involving relative spatial and temporal scales in which local/site gradients are dependent on bioclimatic/regional gradients). This avoids the problem of classes that are not mutually exclusive and enables the classification of all types of ecosystems, including for example marshes on rocky outcrops in superhumid tropical montane areas. Secondly, the names of ecosystem species are linked to a nomenclatural type defined by a ‘type site’ or ‘biotype’, adopting a principle that makes clear a given author's notion of an ecosystem type even if the accompanying name and description are partial or imperfect, or when the ecosystem type is delimited too broadly according to the interpretation of another author. Ecosystem names are structured as a descriptive diagnosis based on a standardized set of characters and character states. This typological approach for facilitating the naming and comparison of ecosystem circumscriptions is thus truly taxonomic in nature. Thirdly, in order to facilitate the use and application of the conceptual approach presented here, we translate it into a practical tool by developing a smartphone-based system to collect data for observing and describing virtual ecosystem specimens in the field, along with the "Bio" database, which manages ecosystem data and also enables tracking synonymies using an open system that entails assigning determinavits to biotypes.

在过去几年中,世界自然保护联盟评估物种灭绝风险的红色名录方法已被改编为生态系统红色名录方法。这一努力面临着几个重要的挑战,包括如何定义适用红色名录标准的生态系统类型,以及如何以开放、透明和标准化的方式管理生态系统地理分布的信息,将制图、类型学和实地研究联系起来。我们提出了一种全新的方法,它与现有的生态系统类型学在三个重要方面有所不同:(1)提供了一种概念化生态系统类型的新方法;(2)提供了一种明确的方法来交流概念化生态系统及其如何被限制;(3)开发管理由此产生的概念模型的技术工具。首先,生态系统类型是通过使用模块化(在给定尺度上研究生态因子的组合)和分层(涉及相对时空尺度,其中局部/站点梯度依赖于生物气候/区域梯度)的方法来研究生物地理气候梯度来定义的。这样就避免了不相互排斥的分类问题,并使所有类型的生态系统能够分类,例如包括在超潮湿的热带山区的岩石露头上的沼泽。其次,生态系统物种的名称与由“类型地点”或“生物类型”定义的命名类型相关联,采用一种原则,即使随附的名称和描述是部分或不完整的,或者根据另一位作者的解释,生态系统类型的界限过于宽泛,也能明确给定作者对生态系统类型的概念。生态系统名称的结构是基于一组标准化的字符和字符状态的描述性诊断。因此,这种便于命名和比较生态系统界限的类型学方法在本质上是真正的分类学方法。第三,为了促进这里提出的概念方法的使用和应用,我们将其转化为一个实用的工具,通过开发一个基于智能手机的系统来收集数据,用于观察和描述野外的虚拟生态系统标本,以及“生物”数据库,该数据库管理生态系统数据,并使用一个开放的系统来跟踪同义词,该系统需要为生物类型分配决定性因素。
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引用次数: 2
How seasonal variations in birth and transmission rates impact population dynamics in a basic SIR model 在基本SIR模型中,出生和传播率的季节变化如何影响种群动态
IF 3.5 3区 环境科学与生态学 Q2 ECOLOGY Pub Date : 2021-09-01 DOI: 10.1016/j.ecocom.2021.100949
Charlotte Ward, Alex Best

The changing climate is expected to alter the timings of key events in species life-histories. These shifts are likely to have important consequences for infectious disease dynamics, as the distribution and abundance of host species will lead to a different environment for parasites. Previous work has shown how seasonality in single host traits - most commonly the reproduction rate or transmission rate - can lead to an array of complex epidemiological dynamics, including chaos and multiple-stable states, with changes to the timing and amplitude of the seasonal peaks often driving drastic changes in behaviour. However, more than one life-history trait is likely to be seasonal, and changing environmental conditions may impact each of them in different ways, yet there have been few studies of host-parasite dynamics that include more than one seasonal trait. Here we examine a Susceptible-Infected-Recovered epidemiological model in which both reproduction and transmission exhibit seasonal fluctuations. We examine how the amplitude and timing of these seasonal peaks impact disease dynamics. We show that the relative timing of the two events is key, with the most stable dynamics when births peak a few months before transmission. We also show that chaotic dynamics become more likely when transmission in particular has a high amplitude, and when baseline transmission and virulence are high. Our results emphasise the importance of seasonality and timing of host life-history events to disease dynamics.

气候变化预计会改变物种生活史中关键事件发生的时间。这些变化可能对传染病动态产生重要影响,因为宿主物种的分布和丰度将导致寄生虫的不同环境。以前的工作已经表明,单个宿主特征的季节性——最常见的是繁殖率或传播率——如何导致一系列复杂的流行病学动态,包括混乱和多重稳定状态,季节性高峰的时间和幅度的变化通常会导致行为的剧烈变化。然而,不止一种生活史特征可能是季节性的,不断变化的环境条件可能以不同的方式影响它们中的每一种,但很少有包括多种季节性特征的宿主-寄生虫动力学研究。在这里,我们研究了一个易感-感染-康复的流行病学模型,其中繁殖和传播都表现出季节性波动。我们研究了这些季节性高峰的幅度和时间如何影响疾病动态。我们表明,这两个事件的相对时间是关键,在传播前几个月出生高峰时,动态最稳定。我们还表明,当传播特别是具有高振幅时,以及基线传播和毒力高时,更有可能出现混沌动力学。我们的研究结果强调了宿主生活史事件的季节性和时间对疾病动力学的重要性。
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引用次数: 3
The co-circulation of two infectious diseases and the impact of vaccination against one of them 两种传染病的共同流行以及针对其中一种传染病接种疫苗的影响
IF 3.5 3区 环境科学与生态学 Q2 ECOLOGY Pub Date : 2021-09-01 DOI: 10.1016/j.ecocom.2021.100941
G.F. Puga , L.H.A. Monteiro

An epidemiological model based on probabilistic cellular automaton is proposed to investigate the dynamics of two co-circulating infections. In the model, one of these two diseases compromises the immune response to future infections; however, there is vaccine against this immunosuppressive disease. The goal is to evaluate the impact of the vaccination coverage on the prevalence and on the cumulative deaths associated with both contagious diseases. The performed numerical simulations highlight the importance of vaccination on decreasing morbidity and mortality. The results are discussed from a public health standpoint, by taking into account outbreaks of measles and COVID-19.

提出了一种基于概率元胞自动机的流行病学模型来研究两种共循环感染的动力学。在该模型中,这两种疾病中的一种损害了对未来感染的免疫反应;然而,有针对这种免疫抑制疾病的疫苗。目标是评估疫苗接种覆盖率对这两种传染病的流行率和与之相关的累积死亡人数的影响。所进行的数值模拟突出了疫苗接种对降低发病率和死亡率的重要性。研究结果从公共卫生的角度进行了讨论,考虑了麻疹和COVID-19的爆发。
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引用次数: 2
Human population induced urban developments and their effects on temperature rise: a nonlinear mathematical model 人口对城市发展的影响及其对气温上升的影响:一个非线性数学模型
IF 3.5 3区 环境科学与生态学 Q2 ECOLOGY Pub Date : 2021-09-01 DOI: 10.1016/j.ecocom.2021.100947
Abhinav Tandon, Prachi Rai Verma

Excessive infrastructural developments, driven by urbanization, have not only brought destruction of forests, but also exacerbated the temperature of cities (or towns) due to formation of urban heat islands. Keeping such an urban system in mind, a nonlinear dynamical model is formulated in the proposed work in terms of system of differential equations. The model, comprising of forest resources, human population, urban infrastructural developments and temperature as system variables, is formulated on the assumption that infrastructural developments, induced through human population, escalate temperature of the region at the cost of deforestation. The derived model is mathematically analyzed for qualitative properties of its equilibrium solutions, extending from their existences to stabilities. Further, to demonstrate the impact of parametric variations on dynamical behavior, the system is also investigated for transcritical and Hopf - bifurcations. Quantitative analysis is also being executed with available numerical data to substantiate qualitative findings and to determine sensitiveness of equilibrium values of model outcomes towards system parameters. The results reveal that any of the parameters, which directly or indirectly, responsible for escalation in temperature of the region can put the system in a state of periodic oscillations, arises through Hopf - bifurcation. Therefore, it is suggested to control urban infrastructural developments through implementation of government strategies, which should include check over illegal encroachment of forested land for infrastructural developments.

在城市化的推动下,过度的基础设施建设不仅带来了森林的破坏,而且由于城市热岛的形成,导致城市(或城镇)的温度升高。考虑到这样一个城市系统,在提出的工作中,以微分方程组的形式制定了非线性动力学模型。该模型由森林资源、人口、城市基础设施发展和温度作为系统变量组成,其假设是由人口引起的基础设施发展以森林砍伐为代价使该地区的温度升高。用数学方法分析了该模型平衡解的定性性质,从存在性扩展到稳定性。此外,为了证明参数变化对动力学行为的影响,还研究了系统的跨临界和Hopf分岔。还利用现有的数值数据进行定量分析,以证实定性结果,并确定模型结果的平衡值对系统参数的敏感性。结果表明,任何直接或间接导致区域温度升高的参数都可以通过Hopf分岔产生,从而使系统处于周期振荡状态。因此,建议通过实施政府战略来控制城市基础设施的发展,其中应包括检查非法侵占林地进行基础设施建设。
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引用次数: 4
High-order interactions maintain or enhance structural robustness of a coffee agroecosystem network 高阶相互作用维持或增强咖啡农业生态系统网络的结构稳健性
IF 3.5 3区 环境科学与生态学 Q2 ECOLOGY Pub Date : 2021-09-01 DOI: 10.1016/j.ecocom.2021.100951
Cecilia González González , Emilio Mora Van Cauwelaert , Denis Boyer , Ivette Perfecto , John Vandermeer , Mariana Benítez

The capacity of highly diverse systems to prevail has proven difficult to explain. In addition to methodological issues, the inherent complexity of ecosystems and issues like multicausality, non-linearity and context-specificity make it hard to establish general and unidirectional explanations. Nevertheless, in recent years, high order interactions have been increasingly discussed as a mechanism that benefits the functioning of highly diverse ecosystems and may add to the mechanisms that explain their persistence. Until now, this idea has been explored by means of hypothetical simulated networks. Here, we test this idea using an updated and empirically documented network for a coffee agroecosystem. We identify potentially key nodes and measure network robustness in the face of node removal with and without incorporation of high order interactions. We find that the system's robustness is either increased or unaffected by the addition of high order interactions, in contrast with randomized counterparts with similar structural characteristics. We also propose a method for representing networks with high order interactions as ordinary graphs and a method for measuring their robustness.

事实证明,很难解释高度多样化的系统为何能够盛行。除了方法论问题外,生态系统固有的复杂性以及多因果性、非线性和环境特异性等问题使得很难建立一般和单向的解释。然而,近年来,高阶相互作用作为一种有利于高度多样化生态系统功能的机制被越来越多地讨论,并可能增加解释其持久性的机制。到目前为止,这个想法已经通过假设的模拟网络进行了探索。在这里,我们使用更新和经验记录的咖啡农业生态系统网络来测试这一想法。我们识别潜在的关键节点,并测量面对节点移除时的网络鲁棒性,无论是否包含高阶交互。我们发现,与具有相似结构特征的随机系统相比,系统的鲁棒性要么增加,要么不受高阶相互作用的影响。我们还提出了一种将具有高阶交互作用的网络表示为普通图的方法和一种测量其鲁棒性的方法。
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引用次数: 0
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Ecological Complexity
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