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Landscape pattern changes across alpine shrub meadows gradient in warm-season pastures on the Qinghai-Tibet Plateau 青藏高原暖季牧草高寒灌丛草甸梯度景观格局变化
IF 3.5 3区 环境科学与生态学 Q2 ECOLOGY Pub Date : 2022-03-01 DOI: 10.1016/j.ecocom.2022.100979
Dawen Qian , Qian Li , Bo Fan , Xiaowei Guo , Yangong Du , Guangmin Cao

Grassland degradation has been one of the major ecological concerns on the Qinghai-Tibet Plateau (QTP) in recent years, but the degradation of alpine shrub meadows, and in particular the changes in its surface landscape pattern, has been less well assessed. This study selected a warm-season pasture on the QTP as a study area, and used an unmanned aerial vehicle (UAV) to collect aerial photographs along the degradation gradient from late June to early July 2018. We then classified the surface landscape as alpine shrub, alpine meadow, bare soil and plateau pika hole and analyzed the landscape pattern changes at different degradation levels. The results showed that the alpine shrub and alpine meadow dominated landscape degraded to a pattern of alpine meadow and bare soil dominance and pika hole pervasiveness, during which vegetation cover declined and the overall landscape pattern tended to fragment. Landscape pattern characteristics related to the area, density, connectivity and boundaries respond more clearly to the shrub degradation, with moderate degradation being the key stage at which the surface landscape pattern changes dramatically. Our study demonstrates a potential application of UAV technology in the study of grassland degradation. Future research should focus on the status, mechanisms and ecological effects of alpine shrub meadows degradation and the quantitative relationships between surface landscape patterns and ecological functions.

近年来,草地退化已成为青藏高原主要的生态问题之一,但对高寒灌丛草甸退化,特别是其地表景观格局变化的评价却较少。以青藏高原暖季牧草为研究区,利用无人机于2018年6月下旬至7月上旬沿退化梯度进行航拍。将地表景观划分为高寒灌丛、高寒草甸、裸土和高原鼠兔洞,分析了不同退化程度下的景观格局变化。结果表明:高寒灌丛和高寒草甸为主的景观格局逐渐退化为高寒草甸和裸土为主、鼠洞普遍的格局,植被覆盖减少,整体景观格局趋于破碎化;与面积、密度、连通性和边界相关的景观格局特征对灌木退化的响应更为明显,其中中度退化是地表景观格局发生剧烈变化的关键阶段。我们的研究展示了无人机技术在草地退化研究中的潜在应用。未来的研究重点应放在高寒灌丛草甸退化的现状、机制、生态效应以及地表景观格局与生态功能的定量关系等方面。
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引用次数: 1
Interactions between changes in land cover and potential of ecosystem services in Lithuania at temporal and spatial scale 立陶宛土地覆盖变化与生态系统服务潜力的时空相互作用
IF 3.5 3区 环境科学与生态学 Q2 ECOLOGY Pub Date : 2022-03-01 DOI: 10.1016/j.ecocom.2022.100984
Gintarė Sujetovienė, Giedrius Dabašinskas

The growing interest in ecosystem services is mainly related to land use changes. The aim of the study is to analyse spatial-temporal changes in the capacity to supply of ES in Lithuania based on land use changes in 1990–2018. The results show some balance between loss and gains of ecosystem services capacity due to land use changes. Decrease in heterogenous agricultural areas had negative impact on provision of ecosystem integrity and services. Considerable increase in scrubland and herbaceous vegetation areas significantly increased the ecosystem service potential. The conversion of former agricultural land to less intensively managed ecosystems enhance the potential of valuable habitats for biodiversity and ecosystem services associated with natural grasslands, moors and heathland, transitional woodland shrubs. The urbanization process along with increase in urban fabric areas had little effect on ESs potential since artificial vegetated areas had compensated the loss of ESs due to increased areas of urbanized ecosystems. Despite the area of open spaces slightly increased, this led to decrease of provision of ESs. Temporal changes in overall ESs capacity indicated an increase in Lithuania over the last two decades. Given the observed dynamic context of land cover, the structure of ecosystem services may face potential threats from land use change due to urban development and agricultural activities.

对生态系统服务日益增长的兴趣主要与土地利用变化有关。该研究的目的是基于1990-2018年土地利用变化分析立陶宛ES供应能力的时空变化。结果表明,由于土地利用变化,生态系统服务能力的损益之间存在一定的平衡。异质性农业区的减少对生态系统完整性和服务的提供产生了负面影响。灌丛和草本植被面积的显著增加显著提高了生态系统服务潜力。将以前的农业用地转变为集约管理较少的生态系统,增加了与天然草原、沼泽和石楠地、过渡林地灌木有关的生物多样性和生态系统服务的宝贵生境的潜力。城市化进程和城市肌构面积的增加对生态环境潜力的影响不大,因为人工植被面积弥补了城市化生态系统面积增加所造成的生态环境潜力损失。虽然休憩用地的面积略有增加,但这导致社会福利的供应减少。总ESs容量的时间变化表明立陶宛在过去二十年中有所增加。鉴于观测到的土地覆盖动态背景,城市发展和农业活动导致的土地利用变化可能对生态系统服务结构构成潜在威胁。
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引用次数: 0
Is landscape fragmentation always detrimental for species conservation? The case of the Iberian lynx in central Spain 景观破碎化是否总是不利于物种保护?西班牙中部伊比利亚猞猁的案例
IF 3.5 3区 环境科学与生态学 Q2 ECOLOGY Pub Date : 2022-03-01 DOI: 10.1016/j.ecocom.2022.100985
PEDRO ALFAYA, CARLOS T.L. DE PABLO, GERMÁN ALONSO

The patch-corridor-matrix is the most commonly used model when dealing with landscape characterization studies, but it shows relevant limitations to detect landscape heterogeneity. Other authors have used a functional approach, since it is well known that nutrient, mineral and energy flows exist among ecosystems. These flows can be perceived in boundaries between different landcovers, making possible the identification of spatial units sharing a common pattern of ecological interactions known as mosaics. While the influence of each mosaic over a certain species has been previously addressed, no attention has been given to the intra-mosaic variation. The aim of this research is to assess the influence of functional diversity and connectivity, on the habitat suitability of the Iberian lynx (Lynx pardinus). For this, we built two GLMs to test if these features show a differential effect on lynxes’ habitat suitability depending on the mosaic. Both GLMs built show that the influence of these landscape features on lynxes’ habitat suitability depends on the landscape spatial organization and landcover composition, suggesting that there is no unique response of a species to changes in landscape diversity and/or connectivity. Thus, we conclude that considering both landscape and species features would allow to a better integration of land management strategies and conservation actions, which could favor species adaptation to highly human-modified landscapes.

斑块-廊道-矩阵是景观表征研究中最常用的模型,但在检测景观异质性方面存在一定的局限性。其他作者使用了功能方法,因为众所周知,生态系统之间存在养分、矿物质和能量流动。这些流动可以在不同土地覆盖之间的边界上被感知,从而有可能识别共享一种称为马赛克的共同生态相互作用模式的空间单元。虽然每一种马赛克对某一物种的影响以前已经解决了,但没有注意到马赛克内部的变化。本研究旨在评估伊比利亚猞猁(lynx pardinus)功能多样性和连通性对生境适宜性的影响。为此,我们建立了两个glm来测试这些特征是否显示出不同马赛克对猞猁栖息地适宜性的不同影响。结果表明,景观特征对山猫生境适宜性的影响取决于景观空间组织和土地覆盖组成,表明物种对景观多样性和/或连通性的变化没有独特的响应。因此,我们得出结论,考虑景观和物种特征将有助于更好地整合土地管理策略和保护行动,从而有利于物种适应高度人为改变的景观。
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引用次数: 1
Optimal control of hybrid variable-order fractional coronavirus (2019-nCov) mathematical model; numerical treatments 混合变阶分数型冠状病毒(2019-nCov)数学模型的最优控制数值的治疗方法
IF 3.5 3区 环境科学与生态学 Q2 ECOLOGY Pub Date : 2022-03-01 DOI: 10.1016/j.ecocom.2022.100983
N.H. Sweilam , S.M. AL-Mekhlafi , T.M. Al-Ajami

A novel coronavirus is a serious global issue and has a negative impact on the economy of Egypt. According to the publicly reported data, the first case of the novel corona virus in Egypt was reported on 14 February 2020. Total of 96753 cases were recorded in Egypt from the beginning of the pandemic until the eighteenth of August, where 96, 581 individuals were Egyptians and 172 were foreigners. Recently, many mathematical models have been considered to better understand coronavirus infection. Most of these models are based on classical integer-order derivatives which can not capture the fading memory and crossover behavior found in many biological phenomena. Therefore, we study the coronavirus disease in this paper by exploring the dynamics of COVID-19 infection using new variable-order fractional derivatives. This paper presents an optimal control problem of the hybrid variable-order fractional model of Coronavirus. The variable-order fractional operator is modified by an auxiliary parameter in order to satisfy the dimensional matching between the both sides of the resultant variable-order fractional equations. Existence, uniqueness, boundedness, positivity, local and global stability of the solutions are proved. Two control variables are considered to reduce the transmission of infection into healthy people. To approximate the new hybrid variable-order operator, Grünwald-Letnikov approximation is used. Finite difference method with a hybrid variable-order operator and generalized fourth order Runge-Kutta method are used to solve the optimality system. Numerical examples and comparative studies for testing the applicability of the utilized methods and to show the simplicity of these approximation approaches are presented. Moreover, by using the proposed methods we can concluded that, the model given in this paper describes well the confirmed real data given by WHO about Egypt.

新型冠状病毒是一个严重的全球问题,对埃及的经济产生了负面影响。根据公开报告的数据,埃及于2020年2月14日报告了首例新型冠状病毒病例。从大流行开始到8月18日,埃及共记录了96753例病例,其中96,581人是埃及人,172人是外国人。最近,人们认为许多数学模型可以更好地理解冠状病毒感染。这些模型大多是基于经典的整阶导数,不能捕捉到许多生物现象中的衰落记忆和交叉行为。因此,我们利用新的变阶分数阶导数来探索COVID-19感染的动态,以此来研究冠状病毒病。提出了冠状病毒变阶分数阶混合模型的最优控制问题。通过一个辅助参数对变阶分数算子进行修改,以满足所得到的变阶分数方程两边的维数匹配。证明了解的存在性、唯一性、有界性、正性、局部稳定性和全局稳定性。考虑两个控制变量来减少感染向健康人的传播。为了逼近新的混合变阶算子,采用了gr nwald- letnikov近似。采用混合变阶算子有限差分法和广义四阶龙格-库塔法求解最优性系统。为验证所采用方法的适用性和说明这些近似方法的简洁性,给出了数值算例和对比研究。此外,利用所提出的方法,我们可以得出结论,本文给出的模型很好地描述了世界卫生组织关于埃及的确认真实数据。
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引用次数: 3
Dynamics and bifurcations of a discrete-time prey-predator model with Allee effect on the prey population 具有Allee效应的离散食饵-捕食模型的动力学和分岔
IF 3.5 3区 环境科学与生态学 Q2 ECOLOGY Pub Date : 2021-12-01 DOI: 10.1016/j.ecocom.2021.100962
Z. Eskandari , J. Alidousti , Z. Avazzadeh , J.A. Tenreiro Machado

This paper studies the dynamic behavior of a discrete-time prey-predator model. It is shown that this model undergoes codimension one and codimension two bifurcations such as transcritical, flip (period-doubling), Neimark-Sacker and strong resonances 1:2, 1:3 and 1:4. The bifurcation analysis is based on the numerical normal form method and the bifurcation scenario around the bifurcation point is determined by their critical normal form coefficients. The advantage of this method is that there is no need to calculate the center manifold and to convert the linear part of the map to a Jordan form. The bifurcation curves of fixed points under variation of one and two parameters are obtained, and the codimensions one and the two bifurcations on the corresponding curves are computed.

研究了离散时间捕食-食饵模型的动态行为。结果表明,该模型经历了余维一分岔和余维二分岔,如跨临界分岔、翻转(周期加倍)分岔、neimmark - sacker分岔和强共振1:2、1:3和1:4分岔。分岔分析基于数值范式方法,分岔点周围的分岔场景由其临界范式系数决定。这种方法的优点是不需要计算中心流形,也不需要将映射的线性部分转换为约旦形式。得到了不动点在一个和两个参数变化下的分岔曲线,并计算了相应曲线上的余维数1和两个分岔。
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引用次数: 11
Complex dynamics of a stochastic uni-directional consumer-resource mutualism system 随机单向消费-资源共生系统的复杂动力学
IF 3.5 3区 环境科学与生态学 Q2 ECOLOGY Pub Date : 2021-12-01 DOI: 10.1016/j.ecocom.2021.100965
Rong Liu , Guirong Liu

Like predation and competition, mutualism is recognized as a consumer-resource interaction, which includes bi-directional and uni-directional mutualisms. In this paper, we firstly propose a stochastic uni-directional consumer-resource system of two species in which the consumer has both positive and negative effects on the resource, while the resource has only a positive effect on the consumer. We then mathematically analyze the system, to demonstrate the existence, uniqueness, asymptotic pathwise behavior and stochastically ultimately boundedness of the global positive solution, and to establish sufficient conditions for the global attractivity and the existence of ergodic stationary distribution of the system. We also establish sufficient conditions for the extinction and persistence in mean of the resource, the consumer or the entire system. Numerical simulations are carried out to demonstrate the analytical results.

与掠夺和竞争一样,互惠互利被认为是一种消费者-资源的互动,包括双向互惠和单向互惠。本文首先提出了一种具有两物种的随机单向消费者-资源系统,其中消费者对资源既有积极作用,也有消极作用,而资源对消费者只有积极作用。然后对系统进行数学分析,证明了系统全局正解的存在唯一性、渐近路径行为和随机最终有界性,并建立了系统全局吸引性和遍历平稳分布存在性的充分条件。我们还从资源、消费者或整个系统的角度建立了灭绝和持续的充分条件。数值模拟验证了分析结果。
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引用次数: 0
The impact of feral domestic cats on native bird populations. Predictive modelling approach on a country scale 野猫对本地鸟类种群的影响。国家范围内的预测建模方法
IF 3.5 3区 环境科学与生态学 Q2 ECOLOGY Pub Date : 2021-12-01 DOI: 10.1016/j.ecocom.2021.100964
Jakub Z. Kosicki

Ecological interactions between native species are often disturbed by invasive species. However, to understand their impact on wild native animal populations on a country scale it is necessary to develop a predictive model. Therefore, I followed the species density distribution modelling approach to explore how feral domestic cats (Felis catus) along with environmental predictors determined densities of two bird species, the Yellowhammer (Emberiza citrinella) and the Yellow Wagtail (Motacilla flava) on the whole area of Poland. As a modelling method, I used the Generalised Additive Model to develop two models for each of the two bird species: The first with the feral cat density as an additional predictor, and the second without it. As a result, I demonstrated the negative impact of cat density on native bird populations, illustrated by reduced density of the two studied species in their preferred habitats, in which cats reached a high density. Although it cannot be explicitly asserted that cats lead to a local extinction of the two bird species, these predators should not be underestimated. In many locations feral populations are fed with new individuals, and they do not follow the same internal mechanisms regulating their population as the native bird fauna. Thus, on a large spatial scale species density distribution models of birds should include cats’ population size as an additional predictor when this predator's environmental preferences overlap with preferences of the studied target groups.

原生物种之间的生态相互作用常常受到外来入侵物种的干扰。然而,要了解它们对国家范围内野生本地动物种群的影响,有必要建立一个预测模型。因此,我遵循物种密度分布建模方法,探索野生家猫(Felis catus)如何与环境预测因子一起决定两种鸟类的密度,黄鹀(Emberiza citrinella)和黄鹡起(Motacilla flava)在整个波兰地区。作为建模方法,我使用广义加性模型为两种鸟类分别开发了两个模型:第一个将野猫密度作为额外的预测因子,第二个没有。结果,我证明了猫的密度对本地鸟类种群的负面影响,这可以从两种被研究物种在其首选栖息地的密度下降来说明,在这些栖息地,猫的密度达到了很高。虽然不能明确断言猫导致了这两种鸟类的局部灭绝,但这些食肉动物不应被低估。在许多地方,野生种群被新个体喂养,它们不遵循与本地鸟类种群相同的调节种群的内部机制。因此,在大的空间尺度上,当捕食者的环境偏好与研究目标群体的偏好重叠时,鸟类物种密度分布模型应该包括猫的种群大小作为额外的预测因子。
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引用次数: 3
Empirical assessment of the relation between ecological connectivity and land complexity based on information-theoretic metrics 基于信息测度的生态连通性与土地复杂性关系实证评价
IF 3.5 3区 环境科学与生态学 Q2 ECOLOGY Pub Date : 2021-12-01 DOI: 10.1016/j.ecocom.2021.100969
Derya Gülçin

Habitat fragmentation and connectivity loss pose significant threats to biodiversity at both local and landscape levels. Strategies to increase ecological connectivity and preserve strong connectivity are important for dealing with the potential threat of habitat degradation. Various metrics have been used to measure (i.e., quantify) landscape composition and configuration in landscape ecology. However, their relationship with ecological connectivity must be understood to interpret landscape patterns comprehensively. In the present study, correlations between ecological connectivity and land complexity are examined based on information-theory metrics. Two primary questions are explored: (1) to what extent are landscape mosaic measures of entropy correlated with ecological connectivity, with landscape gradient-based measures, and with each other? (2) are landscape gradient-based entropy measures correlated with ecological connectivity more than discrete entropy measures? Results show that all information theoretic metrics are statistically significant (p < 0.05) for modelling ecological connectivity. Among categorically-based indices, the relationship between ECI and joint entropy was the most significant, while a generalized additive model indicated that Boltzmann entropy could predict the ecological connectivity index, explaining ∼60% of the variance. Therefore, configurational entropy can be used for improving ecological connectivity models.

栖息地破碎化和连通性丧失对地方和景观层面的生物多样性构成重大威胁。加强生态连通性和保持强连通性的战略对于应对栖息地退化的潜在威胁至关重要。在景观生态学中,各种各样的指标被用来衡量(即量化)景观的组成和配置。然而,要全面地解释景观格局,必须了解它们与生态连通性的关系。在本研究中,生态连通性和土地复杂性之间的相关性是基于信息理论的度量。本文探讨了两个主要问题:(1)景观熵的马赛克度量与生态连通性、基于景观梯度的度量以及彼此之间的相关性在多大程度上?(2)基于景观梯度的熵测度是否比离散熵测度与生态连通性更相关?结果表明,所有信息论指标均具有统计学显著性(p <0.05)模拟生态连通性。在基于分类的指数中,ECI与联合熵之间的关系最为显著,而广义加性模型表明,Boltzmann熵可以预测生态连通性指数,解释了60%的方差。因此,构型熵可以用于改进生态连通性模型。
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引用次数: 3
For whom is it more beneficial to stop interactions with defectors: Cooperators or defectors? 停止与叛逃者的交往对谁更有利:合作者还是叛逃者?
IF 3.5 3区 环境科学与生态学 Q2 ECOLOGY Pub Date : 2021-12-01 DOI: 10.1016/j.ecocom.2021.100968
Shun Kurokawa

Cooperation is a mysterious evolutionary phenomenon and its mechanisms require elucidation. When cooperators can stop interactions with defectors, the evolution of cooperation becomes possible; this is one mechanism that facilitates the evolution of cooperation. Here, stopping interactions with defectors is beneficial not only for cooperators but also for defectors. The question then arises, for whom is stopping interactions with defectors more beneficial: cooperators or defectors? By utilizing evolutionary game theory, I addressed this question using a two-player game involving four strategies: (1) cooperators who stop the interaction if the current partner is a defector, (2) cooperators who attempt to maintain a relationship with anyone, (3) defectors who stop the interaction if the current partner is a defector, and (4) defectors who attempt to maintain a relationship with anyone. Our results show that, at equilibrium, the ratio of cooperators who stop the interaction if the current partner is a defector to cooperators who attempt to maintain a relationship with anyone is larger than the ratio of defectors who stop the interaction if the current partner is a defector to defectors who attempt to maintain a relationship with anyone. Thus, cooperators rather than defectors are more likely to stop interactions with defectors at equilibrium. This result is consistent with a previous experimental study in which a positive correlation was detected between the degree of individuals’ cooperativeness and how accurately the individuals recognize whether other individuals are cooperators or defectors. Thus, the theoretical work presented in this study provides relevant insights into the natural phenomena of cooperation and recognition.

合作是一种神秘的进化现象,其机制有待阐明。当合作者可以停止与叛逃者的互动时,合作的进化成为可能;这是促进合作进化的一种机制。在这里,停止与脱北者的互动不仅对合作者有利,对脱北者也有利。那么问题来了,停止与叛逃者的互动对谁更有利:合作者还是叛逃者?利用进化博弈论,我用一个涉及四种策略的双人博弈来解决这个问题:(1)如果当前合作伙伴是一个叛逃者,合作者会停止互动;(2)合作者试图与任何人保持关系;(3)叛逃者如果当前合作伙伴是一个叛逃者,叛逃者会停止互动;(4)叛逃者试图与任何人保持关系。我们的结果表明,在均衡状态下,如果当前合作伙伴是一个叛逃者,那么合作者停止互动的比例与试图与任何人保持关系的合作者的比例大于如果当前合作伙伴是一个叛逃者,那么叛逃者停止互动的比例与试图与任何人保持关系的叛逃者的比例。因此,合作者比叛逃者更有可能在平衡状态下停止与叛逃者的互动。这一结果与之前的一项实验研究一致,该研究发现,个体的合作程度与个体识别其他个体是合作者还是叛逃者的准确性之间存在正相关关系。因此,本研究的理论工作为合作与认同的自然现象提供了相关的见解。
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引用次数: 2
Tick bite risk resulting from spatially heterogeneous hazard, exposure and coping capacity 空间异质性危害、暴露和应对能力导致的蜱叮咬风险
IF 3.5 3区 环境科学与生态学 Q2 ECOLOGY Pub Date : 2021-12-01 DOI: 10.1016/j.ecocom.2021.100967
S.O. Vanwambeke , P.H.T. Schimit

Tick-borne diseases have long been mainly associated with forests, the primary habitat for Ixodes ricinus, where they are mostly found. However, increasing evidence shows that humans also often get bitten in gardens, parks and other habitats generally associated with lower vegetation and tick density. Therefore, to understand the risk of infection from a tick bite and thus of potential subsequent infection, it is necessary to separate the factors of risk: hazard and vulnerability, here detailed as exposure and coping capacity, and to examine their spatial heterogeneity. This paper proposes a spatially explicit model for human movement through the entire landscape and forest visits to investigate the three components of risk. The population and its movements are set spatially in three study case landscapes extracted from Wallonia, south Belgium. Parameters that are challenging to estimate, such as the probability of a person getting bitten in various environments and the probability of inspecting one’s body to remove ticks, are analyzed in a wide range of combinations. Results show that, while bites are densest in the forest, they happen across the landscape at levels comparable when summed. When coping capacity is modified, as it could be through raising awareness and improving uptake of protective measures, the most at-risk group can change, and the riskier landscape can become the periurban or the rural landscape. This model offers a platform to investigate the respective contributions of hazard, exposure, and people’s capacity to cope with the hazard. It would benefit from empirical input parameters measured more specifically for its purpose.

长期以来,蜱传疾病主要与森林有关,森林是蓖麻蜱的主要栖息地,它们大多在那里被发现。然而,越来越多的证据表明,人类也经常在花园、公园和其他通常与低植被和蜱虫密度相关的栖息地被咬伤。因此,为了了解蜱叮咬的感染风险以及潜在的后续感染风险,有必要分离风险因素:危害和脆弱性,这里详细描述为暴露和应对能力,并研究它们的空间异质性。本文提出了一个人类在整个景观和森林中活动的空间明确模型,以调查风险的三个组成部分。人口及其流动在比利时南部瓦隆尼亚提取的三个研究案例景观中进行空间设置。一些难以估计的参数,比如一个人在各种环境中被咬的概率,以及检查身体以清除蜱虫的概率,都是在广泛的组合中进行分析的。结果表明,虽然咬伤在森林中最密集,但它们在整个景观中发生的水平与总和相当。当应对能力得到改善时,比如可以通过提高认识和采取更好的保护措施,风险最高的群体就会发生变化,风险较高的景观就会变成城郊或农村景观。该模型提供了一个平台来调查危害、暴露和人们应对危害的能力各自的贡献。它将受益于更具体地为此目的而测量的经验输入参数。
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引用次数: 5
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Ecological Complexity
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