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New material of the frog Hungarobatrachus szukacsi Szentesi & Venczel, 2010, from the Santonian of Hungary, supports its neobatrachian affinities and reveals a Gondwanan influence on the European Late Cretaceous anuran fauna 匈牙利Santonian蛙Hungarobatrachus szukacsi Szentesi&Venczel,2010的新材料支持了其新巴塔拉纪的亲缘关系,并揭示了冈瓦纳大陆对欧洲晚白垩世无核动物群的影响
IF 1.8 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2021-04-08 DOI: 10.5252/geodiversitas2021v43a7
M. Venczel, Zoltán Szentesi, J. Gardner
ABSTRACT The Late Cretaceous anuran Hungarobatrachus szukacsi Szentesi & Venczel, 2010 was erected for isolated ilia and tibio-fibulae from the Santonian-age Iharkút locality, in northwestern Hungary. On the strength of ilial features, H. szukacsi was interpreted as a neobatrachian and possible ranoid, making it the only pre-Cenozoic occurrence for both clades in Laurasia. New ilia and the first examples of skull bones (incomplete frontoparietals, squamosals, maxillae, and angulosplenials) from the type locality provide new insights into the taxonomic distinctiveness, osteology, and evolutionary history of H. szukacsi. In addition to its diagnostic ilia (e.g., dorsal crest tall and ornamented laterally with prominent ridges; extensive interiliac tubercle developed across entire medial surface of acetabular region), H. szukacsi is characterized further by having a moderately hyperossified skull exhibiting such traits as frontoparietals, squamosals, and maxillae externally covered with prominent pit-and-ridge and weakly developed tuberculate ornament (i.e., exostosis), frontoparietals solidly fused along midline, frontoparietals expanded posterolaterally to form a broad squamosal process, squamosals expanded anteroposteriorly to form a plate-like lamella alaris, and maxilla articulating posteriorly with the quadratojugal to form a solid bony ‘cheek'. The first cladistic analysis to include H. szukacsi corroborates its neobatrachian status, but consistently places it among hyloids, rather than ranoids as originally proposed. Indications of hyloids on the African continent and in Madagascar during the Late Cretaceous, suggest that the ancestor of H. szukacsi may have dispersed from Africa, across the proto-Mediterranean and into Europe, prior to the Santonian.
摘要:晚白垩世anuran Hungarobatrachus szukacsi Szentesi&Venczel,2010是为匈牙利西北部Santonian时代Iharkút地区的孤立髂骨和胫腓骨建造的。根据髂骨特征的强度,H.szukacsi被解释为新巴塔拉奇阶和可能的ranoid,使其成为劳拉西亚两个分支在新生代前唯一出现的分支。新髂骨和第一批来自该类型地区的颅骨(不完整的额顶骨、鳞片骨、上颌骨和角脾骨)为H.szukacsi的分类独特性、骨学和进化史提供了新的见解。除了诊断性髂骨(例如,背嵴高大,侧面有突出的嵴;在髋臼区的整个内侧表面发育有广泛的髂骨间结节),H.szukacsi的特征还在于具有中等程度的高分类颅骨,表现出额顶骨、鳞片、,上颌骨外部覆盖有突出的窝脊和发育较弱的结节状装饰物(即外突),额顶沿中线牢固融合,额顶向后扩展形成宽阔的鳞片突,鳞片前后扩展形成板状翼片,上颌骨与颈方向后铰接,形成一个坚固的骨质“脸颊”。第一个包含H.szukacsi的支序分析证实了它的新巴塔拉奇阶地位,但始终将其归入亚门,而不是最初提出的ranoid。白垩纪晚期非洲大陆和马达加斯加的类人猿迹象表明,H.szukacsi的祖先可能在Santonian之前就已经从非洲分散开来,穿过原地中海,进入欧洲。
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引用次数: 5
Les Thalattosuchia jurassiques de Normandie des collections Vautier et Morière : contexte historique et redécouverte des plastotypes Vautier和Morière收藏的诺曼底侏罗纪thalattosuchia:塑性模型的历史背景和重新发现
IF 1.8 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2021-03-25 DOI: 10.5252/geodiversitas2021v43a6
A. Brignon
RÉSUMÉ Abel Vautier (1794-1863) avait constitué à Caen, un musée privé éclectique dans lequel était conservée une importante collection paléontologique. Il avait fait l'acquisition de plusieurs spécimens remarquables de reptiles marins jurassiques de Normandie (Ichthyosauria, Plesiosauria et Thalattosuchia). Après sa mort, ces spécimens furent achetés en novembre 1863 par Jacques-Amand Eudes-Deslongchamps et intégrèrent en 1935 les collections de la Faculté des Sciences de Caen. Une autre personnalité caennaise, Pierre-Gilles Morière (1817-1888), avait également collecté plusieurs spécimens importants de Thalattosuchia dans le Jurassique de Normandie. Contrairement à Vautier, Morière ne cherchait pas à constituer une collection personnelle et déposait directement ses trouvailles à la Faculté de Caen. L'ensemble des pièces collectées par Vautier et Morière subirent néanmoins le même sort et furent détruites dans les bombardements de Caen en 1944. Cet article retrace l'histoire des spécimens les plus emblématiques de ces collections, à savoir notamment, l'holotype de « Metriorhynchus » blainvillei (J.-A. Eudes-Deslongchamps, 1867) trouvé en 1844 dans le Callovien inférieur de Sannerville (Calvados), l'holotype de Deslongchampsina larteti (J.-A. Eudes-Deslongchamps, 1866) découvert avant 1853 dans le Calcaire de Caen (Bathonien moyen) à Fleury-sur-Orne, l'holotype de « Metriorhynchus » brachyrhynchus (J.-A. Eudes-Deslongchamps, 1867) trouvé en 1864 dans le Callovien au Mesnil de Bavent (Calvados) et enfin un crâne entier de Teleidosaurus calvadosii (J.-A. Eudes-Deslongchamps, 1866), désigné ici lectotype de l'espèce, découvert en 1864 dans le Calcaire de Caen (Bathonien moyen) à Fleury-sur-Orne. Eudes-Deslongchamps avait fait faire, avec le soutien d'Henri Milne Edwards et du ministère de l'Instruction publique, des moulages en plâtre de ces spécimens. Ces plâtres furent réalisés entre 1866 et 1868 par Jean-Benjamin Stahl, chef de l'atelier de moulage du Muséum national d'Histoire naturelle. Ils furent distribués à différents musées et facultés des sciences en France. Des plastotypes de « Metriorhynchus » blainvillei, Deslongchampsina larteti, « Metriorhynchus » brachyrhynchus et de Teleidosaurus calvadosii dont l'origine était oubliée depuis longtemps ont pu ainsi être retrouvés dans plusieurs collections publiques françaises.
阿贝尔·沃蒂尔(1794-1863)在卡昂建立了一个折衷的私人博物馆,其中保存了大量古生物学藏品。他获得了诺曼底侏罗纪海洋爬行动物(鱼龙、Plesiosauria和Thalattosuchia)的几个杰出标本。1863年11月,雅克·阿曼德·尤德斯·德斯朗尚(Jacques Amand Eudes Deslongchamps)在他去世后购买了这些标本,并于1935年成为卡昂科学院收藏的一部分。另一位卡昂人皮埃尔·吉勒斯·莫里埃(1817-1888)也在诺曼底侏罗纪收集了一些重要的Thalattosuchia标本。与沃蒂尔不同,莫里埃并没有试图建立个人收藏,而是将他的发现直接存放在卡昂学院。然而,沃蒂尔和莫里埃收集的所有物品都遭遇了同样的命运,并在1944年卡昂爆炸中被摧毁。本文追溯了这些藏品中最具代表性的标本的历史,特别是1844年在Sannerville(Calvados)的下Callovian发现的“Metriorhynchus”Blainvillei(J.-A.Eudes Deslongchamps,1867年)的正型,以及1853年之前在奥恩河畔弗勒里的卡昂石灰岩(中巴托尼亚)中发现的Deslongchampsina larteti(J.-A.Eudes Deslongchamps,1866年)正型,“Metriorhynchus”Brachyrhynchus(J.-A.Eudes Deslongchamps,1867年)的正型于1864年在Mesnil de Bavent(Calvados)的Callovian中发现,最后是Teleidosaurus calvadosii(J.-A.Eudes Deslongchamps,1866年)的整个头骨,这里被指定为该物种的选择型,于1864年间在奥恩河畔弗勒里的卡昂石灰岩(中巴托尼亚)中发现。在亨利·米尔恩·爱德华兹(Henri Milne Edwards)和公共教育部的支持下,尤德斯·德斯朗尚(Eudes Deslongchamps)制作了这些标本的石膏模型。这些石膏是由国家自然历史博物馆铸造车间负责人让·本杰明·斯塔尔于1866年至1868年间制作的。它们被分发给法国的不同博物馆和科学院。“Metriorhynchus”Blainvillei、Deslongchampsina larteti、“Metriorhynchus”Brachyrhynchus和Teleidosaurus calvadosii的塑形模型,其起源早已被遗忘,已在法国公共收藏中找到。
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引用次数: 1
Middle Eocene vertebrate fauna from the Aridal Formation, Sabkha of Gueran, southwestern Morocco 摩洛哥西南部Gueran Sabkha地区Aridal组中始新世脊椎动物区系
IF 1.8 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2021-03-11 DOI: 10.5252/geodiversitas2021v43a5
Samir Zouhri, P. Gingerich, Bouziane Khalloufi, E. Bourdon, S. Adnet, S. Jouve, Najia Elboudali, Ayoub Amane, J. Rage, Rodolphe Tabuce, France de Lapparent de Broin
ABSTRACT In the Sahara Desert of southwestern Morocco, the Aridal Formation of Gueran is known for the world's richest Bartonian archaic whale assemblage, which includes both protocetids and basilosaurids. Gueran has also yielded another rich and diverse vertebrate fauna described in detail herein —The chondrichthyan assemblage of twelve species is quite similar to that of the Midawara Formation (Egypt). Actinopterygians include Siluriformes, Percomorpha and rostra of Cylindracanthus Leidy, 1856. Turtles are attributed to at least three indetermined species: two marine cryptodires – a cheloniid and a dermochelyid, and a possible littoral pleurodire, as found in Ad-Dakhla (Morocco) and Fayum (Egypt). The crocodylians comprise at least two longirostrine taxa, including a gavialoid that resembles the late Eocene-early Oligocene Eogavialis africanum Andrews, 1901 from Egypt. The second form is too fragmentary to be identified more precisely than Crocodyliformes indet. Two snake vertebrae belong to Pterosphenus cf. schweinfurthi Andrews, 1901. Two other incomplete snake vertebrae probably belong to Paleophiidae as well. Seabird remains belong to a gigantic soaring pseudo-toothed bird (Pelagornithidae) and constitute the earliest occurrence of the genus Pelagornis sp. Lartet, 1857. This material extends the fossil record of Pelagornis back in time by at least 10 million years. Based on their size and enamel microstructure, mammal dental fragments are attributed to the proboscidean ?Barytherium sp. The Bartonian age of the fauna, initially based on an archaeocete cetacean assemblage, is also supported by chondrichthyans. Affinities of the Gueran faunal assemblage are analyzed in comparison with those from other middle and upper Eocene deposits of North Africa and elsewhere.
在摩洛哥西南部的撒哈拉沙漠,Gueran的Aridal组以世界上最丰富的巴尔顿古鲸组合而闻名,其中包括原鲸类和龙王鲸类。Gueran还发现了另一种丰富多样的脊椎动物区系,本文详细描述了12种软骨鱼的组合,与埃及Midawara组非常相似。放线菌科包括Siluriformes, Percomorpha和rostra of canthus Leidy, 1856。海龟至少属于三种不确定的物种:两种海洋隐龟类——一种龟类和一种皮龟类,以及一种可能的滨海胸龟,在摩洛哥的Ad-Dakhla和埃及的Fayum发现。鳄鱼至少包括两个长颈类群,包括一个类似始新世晚期-渐新世早期非洲古鳄鱼(Eogavialis africanum Andrews, 1901年)的类鳄鱼。第二种形式太零碎,不能比鳄鱼形更精确地识别。两条蛇椎骨属于Pterosphenus . cf. schweinfurthi Andrews, 1901。另外两条不完整的蛇椎骨可能也属于古蛇科。海鸟遗骸属于一种巨大的翱翔的伪齿鸟(Pelagornithidae),构成了Pelagornis sp. Lartet, 1857年的最早发现。这些材料将伯拉哥龙的化石记录往前推了至少1000万年。根据它们的大小和牙釉质微观结构,哺乳动物的牙齿碎片被归为proboscidean Barytherium sp.。Bartonian时代的动物群,最初是基于一个太古鲸类组合,也得到了软骨鱼类的支持。通过与北非和其他地区始新世中上沉积的动物组合进行比较,分析了圭兰动物组合的亲缘性。
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引用次数: 5
Le Jurassique du Jebel Ressas (Tunisie). Identification du Sinémurien supérieur dans la Formation Zaghouan. Précisions stratigraphiques et paléobiogéographiques sur le Sinémurien de Tunisie Jebel Ressas的侏罗纪(突尼斯)。Zaghouan地层中高级sinemurian的鉴定。突尼斯Sinemurian的地层和古生物地理细节
IF 1.8 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2021-02-18 DOI: 10.5252/geodiversitas2021v43a4
P. Fauré, Nejla Sekatni, Mohamed Sabri Arfaoui, R. Alouani
RÉSUMÉ La révision de la succession lithostratigraphique du Jurassique du Jebel Ressas (Dorsale tunisienne, Tunisie) permet d'identifier toutes les formations déjà définies par ailleurs dans la Dorsale tunisienne. La Formation Zaghouan y est nouvellement datée du Sinémurien supérieur par des ammonites appartenant aux chronozones à Obtusum (sous-chronozone à Stellare) et à Raricostatum (sous-chronozone à Raricostatum). Six taxons d'Ammonitina appartenant aux genres Asteroceras (A. gr. saltriense (Parona, 1896), A. cf. meridionalis Dommergues, Meister & Mettraux, 1990), Arnioceras (A. gr. ceratitoides (Quenstedt, 1849) – rejectum Fucini, 1902), Epophioceras (E. cf. landrioti (d'Orbigny, 1850)) et Paltechioceras (P. bavaricum (Böse, 1894), P. cf. charpentieri (Schafhäult, 1847)) sont décrits et figurés. Ces ammonites permettent de corréler la « Formation Zaghouan » avec les « Niveaux condensés du sommet de la Formation Oust », déjà bien connus pour leur riche ammonitofaune sinémurienne et dont les âges sont ici réévalués. Les affinités paléobiogéographiques des ammonites du Sinémurien de Tunisie sont discutées.
通过对Jebel Ressas侏罗纪(突尼斯背侧,突尼斯)岩石地层演替的修正,可以确定突尼斯背侧其他地方已经确定的所有地层。Zaghouan组新确定为上西尼莫里亚时期,菊石属于钝时带(亚时带带Stellare)和Raricostatum(亚时带带Raricostatum)。六个类群d’Ammonitina Asteroceras所属流派(a·斯莱saltriense Parona(1896)、a .参看meridionalis Dommergues Meister & Mettraux, 1990)、Arnioceras (a·斯莱ceratitoides (Quenstedt—1849)、rejectum Fucini 1902)、Epophioceras(1850))(参看landrioti orbigny先生阁下和p bavaricum Paltechioceras(1894)、p (B o se,见charpentieri Schafh外护理,1847)和描述)的大小。这些菊石使我们有可能将“扎古万组”与“乌斯特组顶部的浓缩水平”联系起来,乌斯特组已经以其丰富的西尼莫亚植物群而闻名,并在这里重新评估其年龄。讨论了突尼斯西尼莫里亚菊石的古生物地理亲缘关系。
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引用次数: 2
The freshwater and terrestrial turtles from Monte Pila and Fuenmayor (La Rioja, northern Spain): new data on the lower Miocene turtle diversity of the Ebro Basin 来自Monte Pila和Fuenmayor(西班牙北部拉里奥哈)的淡水和陆生龟:埃布罗盆地中新世龟多样性的新数据
IF 1.8 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2021-02-11 DOI: 10.5252/geodiversitas2021v43a3
A. Pérez‐García, Oier Suárez-Hernando, J. Hernández, S. García, X. Murelaga
ABSTRACT Abundant turtle fossils from two lower Miocene localities in which this lineage had not previously been studied are described herein. The localities are Monte Pila and Fuenmayor, which are 10 km apart, both located in the Autonomous Community of La Rioja (northern Spain), in the Ebro Basin. Information about the turtles from the lower Miocene levels in this basin was until now restricted to the finds made in the MN3 biozone of the Bardenas Reales of Navarre. Several turtle taxa had been recognized there. However, the material attributed to each of them was scarce. Monte Pila and Fuenmayor are located in biozone MN2. The remains of turtles are abundant in both sites. A terrestrial lineage, recognized at Monte Pila, is not represented in the Bardenas Reales fauna of Navarre. It represents the first record of Titanochelon Pérez-García & Vlachos, 2014 in La Rioja and is one of the oldest member of this genus, being assigned to Titanochelon cf. bolivari (Hernández-Pacheco, 1917). The only lineage of turtles identified in Fuenmayor, and the most abundant in Monte Pila, is Ptychogasterinae. All material of this clade of freshwater turtles identified there is compatible with a single form, for which a high range of intraspecific variability is recognized. It is attributed to Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola & Astibia, 1999, this species having been recognized, until now, as exclusive to the Bardenas Reales of Navarre.
摘要本文描述了来自中新世下两个地区的丰富海龟化石,这些地区以前从未对该谱系进行过研究。地点是相距10公里的Monte Pila和Fuenmayor,均位于埃布罗盆地的拉里奥哈自治区(西班牙北部)。到目前为止,该盆地中新世较低水平的海龟信息仅限于在纳瓦尔Bardenas Reales的MN3生物区发现的海龟。在那里已经发现了几个海龟分类群。然而,属于他们每个人的材料都很少。Monte Pila和Fuenmayor位于MN2生物区。这两处遗址都有大量的海龟遗骸。在蒙特皮拉发现的陆地谱系在纳瓦拉的Bardenas Reales动物群中没有代表性。它代表了Titanochelon Pérez-García和Vlachos于2014年在拉里奥哈的第一个记录,是该属最古老的成员之一,被分配给Titanocholon cf.bolivari(Hernández-Pacheco,1917)。在Fuenmayor发现的唯一一个海龟谱系,也是在Monte Pila发现的最丰富的海龟谱系,是Ptychogasterinae。在那里发现的淡水龟分支的所有材料都与一种单一的形式兼容,这种形式具有高度的种内变异性。它被认为是Ptychogaster(Temnoclemmys)(参见bardenensis Murelaga,Laparent de Broin,Pereda Suberbiola&Astibia,1999),该物种迄今为止一直被认为是纳瓦尔Bardenas Reales的专属物种。
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引用次数: 1
Review of the Late Jurassic erymoid lobsters (Crustacea: Decapoda) 晚侏罗世杏鲍目龙虾研究进展(甲壳纲:十足目)
IF 1.8 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2021-01-28 DOI: 10.5252/geodiversitas2021v43a2
J. Devillez, S. Charbonnier
ABSTRACT Erymoid lobsters (Crustacea, Decapoda, Erymoidea) are an important component of Mesozoic crustacean faunas in Europe, especially during the Jurassic. With 36 species reported, these lobsters reach their highest diversity during the Late Jurassic. After the review presented here, 23 species belonging to Eryma Meyer, 1840 (11 species), Palaeastacus Bell, 1850 (2 species), Pustulina Quenstedt, 1857 (2 species) and Stenodactylina Beurlen, 1928 (8 species) remain valid. One new species is described: Stenodactylina shotoverigiganti n. sp., and Eryma pseudoventrosa Beurlen, 1928 is integrated to Stenodactylina. We also notice the oldest representative of Enoploclytia M'Coy, 1849, known by a single specimen unidentified at specific level. Eryma ventrosum (Meyer, 1835) is the most common species in Western Europe, and may be seen as emblematic of the Middle-Late Jurassic. Moreover, the lithographic limestones of Germany yield an exceptionally diversified erymoid fauna, with four genera (Eryma, Palaeastacus, Pustulina, Stenodactylina) and 11 species listed. All the Late Jurassic representatives of Palaeastacus were found in this lithology. Finally, the examination of some specimens allows the observation of the strong effects of the decortication on the ornamentation of the erymoids and the resulting taxonomic issues.
摘要乙状龙虾(甲壳纲、十足目、乙状目)是欧洲中生代甲壳类动物群的重要组成部分,尤其是在侏罗纪。据报道,这些龙虾有36种,在侏罗纪晚期达到了最高的多样性。经过本文的综述,属于Eryma Meyer,1840(11种),Paleastacus Bell,1850(2种),Pustulina Quenstedt,1857(2种,)和Stenodacylina Beurlen,1928(8种)的23种物种仍然有效。描述了一个新种:Stenodactylina shotoverigianti n.sp.和Eryma pseudo-ventrosa Beurlen,1928整合到Stenodacylina。我们还注意到最古老的伊诺普洛克莱蒂亚M'Coy代表,1849年,通过一个特定级别的不明标本已知。文氏Eryma ventrosum(Meyer,1835)是西欧最常见的物种,可能被视为侏罗纪中晚期的象征。此外,德国的石版石灰石产生了异常多样化的梨状动物群,共有四个属(Eryma、Paleastacus、Pustulina、Stenodactylina)和11个物种。Palaistacus的所有晚侏罗世代表都是在该岩性中发现的。最后,通过对一些标本的检查,可以观察到剥皮对筛骨纹饰的强烈影响以及由此产生的分类学问题。
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引用次数: 4
A nearly complete skeleton of the oldest definitive erycine boid (Messel, Germany) 一具几乎完整的最古老的确定的狼人骨架(德国梅塞尔)
IF 1.8 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2021-01-14 DOI: 10.5252/geodiversitas2021v43a1
Krister T. Smith, A. Scanferla
ABSTRACT A nearly complete skeleton of an erycine boid is described from the Ypresian-Lutetian (early-middle Eocene) site of Messel, Germany, for which we propose the name Rageryx schmidi n. gen., n. sp. The animal had a total length of c. 52 cm, with c. 258 vertebrae. In skull proportions it is similar to ungaliophiine boids, especially Ungaliophis, and to Tropidophis. The proportions and distinctive accessory processes of the distal caudal vertebrae that are common to all living erycine boids are present in the specimen, although the processes are not as elaborate as in many extant species. The premaxilla was not protruded to form a wedge-shaped snout, and the nasofrontal joint does not appear to show any special buttressing, unlike in many burrowing snake species. Furthermore, the inner ear lacks adaptations to an actively burrowing mode of life. We conclude that the animal, while it was probably secretive, was not fossorial. Phylogenetic analyses using maximum parsimony and Bayesian inference place Rageryx schmidi n. gen., n. sp. unambiguously on the stem of the North American clade (Lichanura + Charina). If this relationship is accurate, it provides further support for a common Euro-American squamate fauna in the early Eocene. The majority of known Messel snake taxa are small-bodied with a small gape, suggesting that such forms may have played a greater role in the early evolutionary radiation of Booidea than their present diversity would suggest.
在德国Messel的Ypresian-Lutetian(早-中始新世)遗址发现了一具几乎完整的赤藓类动物骨架,我们建议将其命名为Rageryx schmidi n. gen., n. sp.该动物全长约52 cm,椎骨约258块。在头骨比例上,它类似于非加利菲亚类,尤其是非加利菲亚类,也类似于Tropidophis。所有活的丹毒生物共有的远尾椎骨的比例和独特的附属过程都存在于标本中,尽管这些过程不像许多现存物种那样复杂。与许多穴居蛇不同的是,它的前颌骨并没有突出来形成楔形的鼻子,鼻前关节似乎也没有任何特殊的支撑。此外,内耳缺乏适应积极的穴居生活方式的能力。我们得出的结论是,这种动物虽然可能是秘密的,但不是化石性的。利用最大简约性和贝叶斯推理的系统发育分析将Rageryx schmidi n. gen., n. sp.明确地定位在北美分支(Lichanura + Charina)的主干上。如果这种关系是准确的,那么它将进一步支持始新世早期欧美共同的鳞片动物群。大多数已知的梅塞尔蛇分类群都是小体小口的,这表明这些形式可能在布idea的早期进化辐射中发挥了比它们现在的多样性更大的作用。
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引用次数: 13
Cranial anatomy of Andinodelphys cochabambensis, a stem metatherian from the early Palaeocene of Bolivia 玻利维亚古新世早期的一种茎类美塔兽——科恰班班安斑龙(Andinodelphys cochabambensis)的颅骨解剖
IF 1.8 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2020-12-31 DOI: 10.5252/geodiversitas2020v42a30
C. de Muizon, Sandrine Ladevèze
ABSTRACT Andinodelphys cochabambensis Marshall & Muizon, 1988 is one of the best preserved metatherian species from the early Palaeocene fauna of Tiupampa (Bolivia). It is represented by five almost complete skulls, three of them being securely associated to sub-complete to partial skeleton. Four skulls could be extracted from a block including several intermingled skeletons. The present paper provides a thorough description of the dental, cranial, and dentary anatomy of A. cochabambensis. The cranial anatomy of A. cochabambensis is similar to that of Pucadelphys andinus. The skull of Andinodelphys however differs from that of Pucadelphys in its larger size and proportionally longer rostrum. Other differences include the presence, in Andinodelphys, of large anteriorly protruding I1s, small palatal vacuities, a transverse canal, and a small hypotympanic sinus. Andinodelphys has the same dental formula as Pucadelphys (I 5/4, C 1/1, P 3/3, M4/4), the plesiomorphic condition for metatherians. Furthermore, both genera share the lack a tympanic process of the alisphenoid, a deep groove for the internal carotid artery at the anterior apex of the promontorium, a small prootic canal perforating the lateral edge of the petrosal and opening laterally in the deep sulcus for the prootic sinus, and a vestigial anterior lamina of the petrosal. Dentally Andinodelphys closely resembles Pucadelphys, the two genera differing in the larger size of the former and in the inconstant presence in the former of a twinned stylar cusp C. Although 25% smaller, the cheek teeth of Andinodelphys closely resemble those of Itaboraidelphys camposi from the early Eocene of Itaboraí (Brazil). As far as dental morphology is concerned, both genera are likely to have diverged from a direct common ancestor, probably Andinodelphys-like, with Itaboraidelphys displaying more derived dental structures. Two isolated petrosal from Itaboraí (Type 2 petrosals) are morphologically close to those of Andinodelphys but distinctly larger. In this paper, a previous interpretation including the teeth of Itaboraidelphys and these petrosals in the same taxon is followed. A phylogenetic analysis retrieved Itaboraidelphys as a sister taxon of the clade Pucadelphys + Andinodelphys, thus lending support to inclusion of the former in the Pucadelphyidae. Three sets of parsimony analyses were performed. A first set of analyses (with all characters) retrieved a strict consensus tree with a clade as follows: (pucadelphyids, (deltatheroidans (stagodontids, Gurlin Tsav skull-GTS), sparassodonts)). An implied weighting analysis with the same data matrix placed the stagodontids in an early diverging position but retained a clade (pucadelphyids, (deltatheroidans, (GTS, sparassodonts))), the deltatheroidans, being therefore inserted in the pucadelphydans. This result implies an independent arrival of pucadelphyids and sparassodonts to South America, which consequently must have been present in North America in the Lat
摘要Andinodelphis cochabambensis Marshall&Muizon,1988是蒂乌帕(玻利维亚)古新世早期动物群中保存最完好的后生动物之一。它由五个几乎完整的头骨代表,其中三个与亚完整到部分骨骼紧密相连。四个头骨可以从一个包括几个混合骨骼的块中提取出来。本文对a.cochabambensis的牙齿、颅骨和牙齿解剖进行了全面的描述。A.cochabambensis的颅骨解剖结构与Pucadelphis andinus相似。然而,Andinodelphis的头骨与Pucadelphis的不同之处在于其更大的尺寸和相应的更长的喙。其他差异包括在Andinodelphis中存在大的向前突出的I1、小的腭部空洞、横管和小的下交感窦。Andinodelphis具有与Pucadelphis相同的牙科配方(I 5/4,C 1/1,P 3/3,M4/4),这是metatherians的准同形条件。此外,这两个属都有一个共同的缺点,即缺乏一个泽泻样的鼓突,一个位于海角前尖的颈内动脉深沟,一个穿过岩侧边缘并在深沟中侧向开口的小前耳管,以及一个残留的岩前板。Andinodelphis在牙齿上与Pucadelphis非常相似,这两个属的区别在于前者的体型较大,前者的柄尖C不稳定。尽管小25%,但Andinodelphes的颊齿与Itaboraí(巴西)始新世早期的Itaboraidelphis camposi的颊齿非常相似。就牙齿形态而言,这两个属都可能是从一个直接的共同祖先分化而来的,可能是类似Andinodelphis的,而Itaboraidelphis显示出更多衍生的牙齿结构。Itaboraí的两个孤立岩(2型岩)在形态上与Andinodelphis的岩相似,但明显更大。在这篇论文中,遵循了先前的解释,包括在同一分类单元中的梨形目和这些岩类的牙齿。一项系统发育分析检索到Itaboraidelphys是Pucadelphys+Andinodelphys分支的姐妹分类单元,从而支持将前者纳入Pucadelphidae。进行了三组简约分析。第一组分析(包括所有特征)检索到一个严格的一致树,其分支如下:(pucadelphiids,(三角兽纲(鹿齿目,Gurlin Tsav头骨GTS),sparassodonts))。具有相同数据矩阵的隐含加权分析将鹿齿目置于早期分叉位置,但保留了一个分支(pucadelphids,(三角兽齿目,(GTS,sparassodonts)),因此三角兽齿类被插入到pucadelfids中。这一结果意味着Pucadelphids和Sparassodons独立到达南美洲,因此它们一定在白垩纪晚期存在于北美洲。北美洲可能的裂头龙属可能是鲜为人知的Atokatheridium属和Olklatheridius属(目前被称为三角兽目),而pucadelphids可能与Aenigdadelphys属一起存在于北美洲白垩纪晚期。然而,这一假设(就古生物地理学而言)不如祖先Pucadelphida(Pucadelyidae+Sparassodonta)的一次南迁那么简单。由于第一组分析的结果可能是由与高碱性饮食相关的高度同源性牙齿特征引起的,因此进行了第二组分析,排除了所有牙齿特征。严格的一致性很难解决,但保留了单系有袋动物和雀形目。隐含加权分析检索到一个单系Pucadelphyda,但分裂了三角兽目,其多系性被认为是与缺乏牙齿特征有关的可能伪影。GTS是Pucadelphyda的姐妹分类单元。由于三角兽亚目的多聚性与之前的所有假设相矛盾,因此进行了第三组分析,仅排除了那些支持高食肉分支(三角兽亚纲、鹿齿亚目和盾齿亚目)密切关系的臼齿特征。严格一致树检索到了单系三角洲动物、有袋动物和裂头动物。隐含加权分析结果表明,三角兽目形成了Metatheria和单系Pucadelphida的系旁茎组合。GTS不再与雀形目有亲缘关系,而是一个分支的姐妹分类单元,包括数据矩阵的北美分类单元、亚洲目和有袋目。这种拓扑结构在这里很受欢迎,它支持(以及第二组分析的拓扑结构)可能在白垩纪晚期,伴随着南美洲肉食性后生动物的蒂潘派辐射,有一次pucadelphidan向南迁移。
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引用次数: 18
Contribution des motifs colorés résiduels dans la discrimination d'espèces nouvelles de Cryptochorda Mörch, 1858 (Mollusca: Gastropoda: Harpidae) de l'Éocène du bassin de Paris et du Cotentin 巴黎和科坦盆地新世Cryptochorda morch, 1858(软体动物:腹足纲:Harpidae)新种鉴定中残留彩色图案的贡献
IF 1.8 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2020-12-17 DOI: 10.5252/geodiversitas2020v42a29
Jean-Michel Pacaud, Frédéric Sautereau
RÉSUMÉ L'observation sous éclairage ultraviolet de motifs colorés résiduels sur les coquilles de très nombreux gastéropodes cénozoïques fournit des caractères permettant d'affiner la description des espèces fossiles. Nous appliquons ici cette méthode à la discrimination de trois espèces de Cryptochorda de l'Éocène du bassin de Paris et du Cotentin confondues jusqu'ici avec l'espèce classique Cryptochorda (s.str.) stromboides (Hermann, 1781). Ces espèces sont nommées: Cryptochorda (s.str.) altavesna n. sp., C. (s. str.) neptis n. sp. et C. (s.str.) cosediensis n. sp. La répartition stratigraphique et géographique de l'espèce type C. (s.str.) stromboides est ainsi redéfinie. Cryptochorda (s.str.) teae n. sp. est également décrite de l'Yprésien (Éocène inférieur) d'Italie. Une revue historique de la littérature portant sur la taxonomie du genre est donnée et une notice biographique est fournie pour Jean Hermann, descripteur de l'espèce type du genre Cryptochorda.
摘要:在紫外线照射下观察大量新生代腹足动物外壳上残留的彩色图案,为完善化石物种描述提供了特征。在这里,我们将该方法应用于巴黎盆地和科坦丁始新世的三种隐球虫物种的区分,迄今为止,它们与经典物种隐球虫(S.str.)stromboides(Hermann,1781)混淆。这些物种被命名为Cryptochorda(S.str.)、Altavesna n.sp.、C.(S.str.)、Neptis n.sp.和C.(S.str.)、Cosediensis n.sp。因此,重新定义了模式物种C.(S.str.)Stromboides的地层和地理分布。Cryptochorda(S.str.)teae n.sp.也描述于意大利的Ypresian(下始新世)。对该属分类学文献进行了历史回顾,并为Cryptochorda模式物种的描述符Jean Hermann提供了传记。
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引用次数: 2
Revision of the cranial anatomy of Ophisaurus acuminatus Jörg, 1965 (Anguimorpha, Anguidae) from the late Miocene of Germany 德国中新世晚期尖锐蛇鼻龙Jörg, 1965(鳗目,鳗科)颅骨解剖的修正
IF 1.8 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2020-12-03 DOI: 10.5252/geodiversitas2020v42a28
Jozef Klembara, A. Čerňanský
ABSTRACT The anguine species Ophisaurus acuminatus Jörg, 1965 is known on the basis of only one specimen consisting of skull bones and osteoderms from the late Miocene (MN 9) Höwenegg/Hegau locality in Germany. Since its first description, several other new species of Ophisaurus Daudin, 1803 have been described from various Miocene localities in Europe. The diagnoses of these new species are based mostly on characters on the parietal and partially frontal bones. Although most of the cranial elements of O. acuminatus are well preserved, its parietal is absent. The knowledge of the detailed anatomy of this anguine species is crucial for the understanding of its interrelationship within the genus Ophisaurus. For our re-study of this specimen we used a high-resolution X-ray microcomputed tomography. The application of this method enabled: 1) to reveal the anatomy of not visible portions of the previously described bones; 2) to discover the bones completely or almost completely embedded in the sediment; and 3) to identify previously not determined skull bones. Our study enabled to identify three distinguished features for this species and confirmed the validity of the species O. acuminatus.
摘要:在德国晚中新世(MN 9) Höwenegg/Hegau地区发现了一种由颅骨和骨皮组成的蛇类标本Jörg, 1965。自第一次描述以来,在欧洲中新世的不同地点已经描述了其他几个新的达乌丁蛇鼻龙。这些新种的诊断主要基于顶骨和部分额骨的特征。虽然大多数的尖顶鳄的头盖骨保存完好,但其顶骨是缺失的。了解这种蛇类物种的详细解剖结构对于了解它在蛇鼻龙属中的相互关系至关重要。为了重新研究这个标本,我们使用了高分辨率的x射线微计算机断层扫描。这种方法的应用使:1)揭示了先前描述的骨骼中不可见部分的解剖结构;2)发现完全或几乎完全埋在沉积物中的骨头;3)鉴定以前未确定的头骨。我们的研究鉴定了该物种的三个特征,并证实了该物种的有效性。
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引用次数: 5
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