Pub Date : 2021-04-08DOI: 10.5252/geodiversitas2021v43a7
M. Venczel, Zoltán Szentesi, J. Gardner
ABSTRACT The Late Cretaceous anuran Hungarobatrachus szukacsi Szentesi & Venczel, 2010 was erected for isolated ilia and tibio-fibulae from the Santonian-age Iharkút locality, in northwestern Hungary. On the strength of ilial features, H. szukacsi was interpreted as a neobatrachian and possible ranoid, making it the only pre-Cenozoic occurrence for both clades in Laurasia. New ilia and the first examples of skull bones (incomplete frontoparietals, squamosals, maxillae, and angulosplenials) from the type locality provide new insights into the taxonomic distinctiveness, osteology, and evolutionary history of H. szukacsi. In addition to its diagnostic ilia (e.g., dorsal crest tall and ornamented laterally with prominent ridges; extensive interiliac tubercle developed across entire medial surface of acetabular region), H. szukacsi is characterized further by having a moderately hyperossified skull exhibiting such traits as frontoparietals, squamosals, and maxillae externally covered with prominent pit-and-ridge and weakly developed tuberculate ornament (i.e., exostosis), frontoparietals solidly fused along midline, frontoparietals expanded posterolaterally to form a broad squamosal process, squamosals expanded anteroposteriorly to form a plate-like lamella alaris, and maxilla articulating posteriorly with the quadratojugal to form a solid bony ‘cheek'. The first cladistic analysis to include H. szukacsi corroborates its neobatrachian status, but consistently places it among hyloids, rather than ranoids as originally proposed. Indications of hyloids on the African continent and in Madagascar during the Late Cretaceous, suggest that the ancestor of H. szukacsi may have dispersed from Africa, across the proto-Mediterranean and into Europe, prior to the Santonian.
{"title":"New material of the frog Hungarobatrachus szukacsi Szentesi & Venczel, 2010, from the Santonian of Hungary, supports its neobatrachian affinities and reveals a Gondwanan influence on the European Late Cretaceous anuran fauna","authors":"M. Venczel, Zoltán Szentesi, J. Gardner","doi":"10.5252/geodiversitas2021v43a7","DOIUrl":"https://doi.org/10.5252/geodiversitas2021v43a7","url":null,"abstract":"ABSTRACT The Late Cretaceous anuran Hungarobatrachus szukacsi Szentesi & Venczel, 2010 was erected for isolated ilia and tibio-fibulae from the Santonian-age Iharkút locality, in northwestern Hungary. On the strength of ilial features, H. szukacsi was interpreted as a neobatrachian and possible ranoid, making it the only pre-Cenozoic occurrence for both clades in Laurasia. New ilia and the first examples of skull bones (incomplete frontoparietals, squamosals, maxillae, and angulosplenials) from the type locality provide new insights into the taxonomic distinctiveness, osteology, and evolutionary history of H. szukacsi. In addition to its diagnostic ilia (e.g., dorsal crest tall and ornamented laterally with prominent ridges; extensive interiliac tubercle developed across entire medial surface of acetabular region), H. szukacsi is characterized further by having a moderately hyperossified skull exhibiting such traits as frontoparietals, squamosals, and maxillae externally covered with prominent pit-and-ridge and weakly developed tuberculate ornament (i.e., exostosis), frontoparietals solidly fused along midline, frontoparietals expanded posterolaterally to form a broad squamosal process, squamosals expanded anteroposteriorly to form a plate-like lamella alaris, and maxilla articulating posteriorly with the quadratojugal to form a solid bony ‘cheek'. The first cladistic analysis to include H. szukacsi corroborates its neobatrachian status, but consistently places it among hyloids, rather than ranoids as originally proposed. Indications of hyloids on the African continent and in Madagascar during the Late Cretaceous, suggest that the ancestor of H. szukacsi may have dispersed from Africa, across the proto-Mediterranean and into Europe, prior to the Santonian.","PeriodicalId":55111,"journal":{"name":"Geodiversitas","volume":"43 1","pages":"187 - 207"},"PeriodicalIF":1.8,"publicationDate":"2021-04-08","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"49596164","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2021-03-25DOI: 10.5252/geodiversitas2021v43a6
A. Brignon
RÉSUMÉ Abel Vautier (1794-1863) avait constitué à Caen, un musée privé éclectique dans lequel était conservée une importante collection paléontologique. Il avait fait l'acquisition de plusieurs spécimens remarquables de reptiles marins jurassiques de Normandie (Ichthyosauria, Plesiosauria et Thalattosuchia). Après sa mort, ces spécimens furent achetés en novembre 1863 par Jacques-Amand Eudes-Deslongchamps et intégrèrent en 1935 les collections de la Faculté des Sciences de Caen. Une autre personnalité caennaise, Pierre-Gilles Morière (1817-1888), avait également collecté plusieurs spécimens importants de Thalattosuchia dans le Jurassique de Normandie. Contrairement à Vautier, Morière ne cherchait pas à constituer une collection personnelle et déposait directement ses trouvailles à la Faculté de Caen. L'ensemble des pièces collectées par Vautier et Morière subirent néanmoins le même sort et furent détruites dans les bombardements de Caen en 1944. Cet article retrace l'histoire des spécimens les plus emblématiques de ces collections, à savoir notamment, l'holotype de « Metriorhynchus » blainvillei (J.-A. Eudes-Deslongchamps, 1867) trouvé en 1844 dans le Callovien inférieur de Sannerville (Calvados), l'holotype de Deslongchampsina larteti (J.-A. Eudes-Deslongchamps, 1866) découvert avant 1853 dans le Calcaire de Caen (Bathonien moyen) à Fleury-sur-Orne, l'holotype de « Metriorhynchus » brachyrhynchus (J.-A. Eudes-Deslongchamps, 1867) trouvé en 1864 dans le Callovien au Mesnil de Bavent (Calvados) et enfin un crâne entier de Teleidosaurus calvadosii (J.-A. Eudes-Deslongchamps, 1866), désigné ici lectotype de l'espèce, découvert en 1864 dans le Calcaire de Caen (Bathonien moyen) à Fleury-sur-Orne. Eudes-Deslongchamps avait fait faire, avec le soutien d'Henri Milne Edwards et du ministère de l'Instruction publique, des moulages en plâtre de ces spécimens. Ces plâtres furent réalisés entre 1866 et 1868 par Jean-Benjamin Stahl, chef de l'atelier de moulage du Muséum national d'Histoire naturelle. Ils furent distribués à différents musées et facultés des sciences en France. Des plastotypes de « Metriorhynchus » blainvillei, Deslongchampsina larteti, « Metriorhynchus » brachyrhynchus et de Teleidosaurus calvadosii dont l'origine était oubliée depuis longtemps ont pu ainsi être retrouvés dans plusieurs collections publiques françaises.
{"title":"Les Thalattosuchia jurassiques de Normandie des collections Vautier et Morière : contexte historique et redécouverte des plastotypes","authors":"A. Brignon","doi":"10.5252/geodiversitas2021v43a6","DOIUrl":"https://doi.org/10.5252/geodiversitas2021v43a6","url":null,"abstract":"RÉSUMÉ Abel Vautier (1794-1863) avait constitué à Caen, un musée privé éclectique dans lequel était conservée une importante collection paléontologique. Il avait fait l'acquisition de plusieurs spécimens remarquables de reptiles marins jurassiques de Normandie (Ichthyosauria, Plesiosauria et Thalattosuchia). Après sa mort, ces spécimens furent achetés en novembre 1863 par Jacques-Amand Eudes-Deslongchamps et intégrèrent en 1935 les collections de la Faculté des Sciences de Caen. Une autre personnalité caennaise, Pierre-Gilles Morière (1817-1888), avait également collecté plusieurs spécimens importants de Thalattosuchia dans le Jurassique de Normandie. Contrairement à Vautier, Morière ne cherchait pas à constituer une collection personnelle et déposait directement ses trouvailles à la Faculté de Caen. L'ensemble des pièces collectées par Vautier et Morière subirent néanmoins le même sort et furent détruites dans les bombardements de Caen en 1944. Cet article retrace l'histoire des spécimens les plus emblématiques de ces collections, à savoir notamment, l'holotype de « Metriorhynchus » blainvillei (J.-A. Eudes-Deslongchamps, 1867) trouvé en 1844 dans le Callovien inférieur de Sannerville (Calvados), l'holotype de Deslongchampsina larteti (J.-A. Eudes-Deslongchamps, 1866) découvert avant 1853 dans le Calcaire de Caen (Bathonien moyen) à Fleury-sur-Orne, l'holotype de « Metriorhynchus » brachyrhynchus (J.-A. Eudes-Deslongchamps, 1867) trouvé en 1864 dans le Callovien au Mesnil de Bavent (Calvados) et enfin un crâne entier de Teleidosaurus calvadosii (J.-A. Eudes-Deslongchamps, 1866), désigné ici lectotype de l'espèce, découvert en 1864 dans le Calcaire de Caen (Bathonien moyen) à Fleury-sur-Orne. Eudes-Deslongchamps avait fait faire, avec le soutien d'Henri Milne Edwards et du ministère de l'Instruction publique, des moulages en plâtre de ces spécimens. Ces plâtres furent réalisés entre 1866 et 1868 par Jean-Benjamin Stahl, chef de l'atelier de moulage du Muséum national d'Histoire naturelle. Ils furent distribués à différents musées et facultés des sciences en France. Des plastotypes de « Metriorhynchus » blainvillei, Deslongchampsina larteti, « Metriorhynchus » brachyrhynchus et de Teleidosaurus calvadosii dont l'origine était oubliée depuis longtemps ont pu ainsi être retrouvés dans plusieurs collections publiques françaises.","PeriodicalId":55111,"journal":{"name":"Geodiversitas","volume":"43 1","pages":"151 - 186"},"PeriodicalIF":1.8,"publicationDate":"2021-03-25","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"42607451","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2021-03-11DOI: 10.5252/geodiversitas2021v43a5
Samir Zouhri, P. Gingerich, Bouziane Khalloufi, E. Bourdon, S. Adnet, S. Jouve, Najia Elboudali, Ayoub Amane, J. Rage, Rodolphe Tabuce, France de Lapparent de Broin
ABSTRACT In the Sahara Desert of southwestern Morocco, the Aridal Formation of Gueran is known for the world's richest Bartonian archaic whale assemblage, which includes both protocetids and basilosaurids. Gueran has also yielded another rich and diverse vertebrate fauna described in detail herein —The chondrichthyan assemblage of twelve species is quite similar to that of the Midawara Formation (Egypt). Actinopterygians include Siluriformes, Percomorpha and rostra of Cylindracanthus Leidy, 1856. Turtles are attributed to at least three indetermined species: two marine cryptodires – a cheloniid and a dermochelyid, and a possible littoral pleurodire, as found in Ad-Dakhla (Morocco) and Fayum (Egypt). The crocodylians comprise at least two longirostrine taxa, including a gavialoid that resembles the late Eocene-early Oligocene Eogavialis africanum Andrews, 1901 from Egypt. The second form is too fragmentary to be identified more precisely than Crocodyliformes indet. Two snake vertebrae belong to Pterosphenus cf. schweinfurthi Andrews, 1901. Two other incomplete snake vertebrae probably belong to Paleophiidae as well. Seabird remains belong to a gigantic soaring pseudo-toothed bird (Pelagornithidae) and constitute the earliest occurrence of the genus Pelagornis sp. Lartet, 1857. This material extends the fossil record of Pelagornis back in time by at least 10 million years. Based on their size and enamel microstructure, mammal dental fragments are attributed to the proboscidean ?Barytherium sp. The Bartonian age of the fauna, initially based on an archaeocete cetacean assemblage, is also supported by chondrichthyans. Affinities of the Gueran faunal assemblage are analyzed in comparison with those from other middle and upper Eocene deposits of North Africa and elsewhere.
{"title":"Middle Eocene vertebrate fauna from the Aridal Formation, Sabkha of Gueran, southwestern Morocco","authors":"Samir Zouhri, P. Gingerich, Bouziane Khalloufi, E. Bourdon, S. Adnet, S. Jouve, Najia Elboudali, Ayoub Amane, J. Rage, Rodolphe Tabuce, France de Lapparent de Broin","doi":"10.5252/geodiversitas2021v43a5","DOIUrl":"https://doi.org/10.5252/geodiversitas2021v43a5","url":null,"abstract":"ABSTRACT In the Sahara Desert of southwestern Morocco, the Aridal Formation of Gueran is known for the world's richest Bartonian archaic whale assemblage, which includes both protocetids and basilosaurids. Gueran has also yielded another rich and diverse vertebrate fauna described in detail herein —The chondrichthyan assemblage of twelve species is quite similar to that of the Midawara Formation (Egypt). Actinopterygians include Siluriformes, Percomorpha and rostra of Cylindracanthus Leidy, 1856. Turtles are attributed to at least three indetermined species: two marine cryptodires – a cheloniid and a dermochelyid, and a possible littoral pleurodire, as found in Ad-Dakhla (Morocco) and Fayum (Egypt). The crocodylians comprise at least two longirostrine taxa, including a gavialoid that resembles the late Eocene-early Oligocene Eogavialis africanum Andrews, 1901 from Egypt. The second form is too fragmentary to be identified more precisely than Crocodyliformes indet. Two snake vertebrae belong to Pterosphenus cf. schweinfurthi Andrews, 1901. Two other incomplete snake vertebrae probably belong to Paleophiidae as well. Seabird remains belong to a gigantic soaring pseudo-toothed bird (Pelagornithidae) and constitute the earliest occurrence of the genus Pelagornis sp. Lartet, 1857. This material extends the fossil record of Pelagornis back in time by at least 10 million years. Based on their size and enamel microstructure, mammal dental fragments are attributed to the proboscidean ?Barytherium sp. The Bartonian age of the fauna, initially based on an archaeocete cetacean assemblage, is also supported by chondrichthyans. Affinities of the Gueran faunal assemblage are analyzed in comparison with those from other middle and upper Eocene deposits of North Africa and elsewhere.","PeriodicalId":55111,"journal":{"name":"Geodiversitas","volume":"43 1","pages":"121 - 150"},"PeriodicalIF":1.8,"publicationDate":"2021-03-11","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"41422800","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2021-02-18DOI: 10.5252/geodiversitas2021v43a4
P. Fauré, Nejla Sekatni, Mohamed Sabri Arfaoui, R. Alouani
RÉSUMÉ La révision de la succession lithostratigraphique du Jurassique du Jebel Ressas (Dorsale tunisienne, Tunisie) permet d'identifier toutes les formations déjà définies par ailleurs dans la Dorsale tunisienne. La Formation Zaghouan y est nouvellement datée du Sinémurien supérieur par des ammonites appartenant aux chronozones à Obtusum (sous-chronozone à Stellare) et à Raricostatum (sous-chronozone à Raricostatum). Six taxons d'Ammonitina appartenant aux genres Asteroceras (A. gr. saltriense (Parona, 1896), A. cf. meridionalis Dommergues, Meister & Mettraux, 1990), Arnioceras (A. gr. ceratitoides (Quenstedt, 1849) – rejectum Fucini, 1902), Epophioceras (E. cf. landrioti (d'Orbigny, 1850)) et Paltechioceras (P. bavaricum (Böse, 1894), P. cf. charpentieri (Schafhäult, 1847)) sont décrits et figurés. Ces ammonites permettent de corréler la « Formation Zaghouan » avec les « Niveaux condensés du sommet de la Formation Oust », déjà bien connus pour leur riche ammonitofaune sinémurienne et dont les âges sont ici réévalués. Les affinités paléobiogéographiques des ammonites du Sinémurien de Tunisie sont discutées.
{"title":"Le Jurassique du Jebel Ressas (Tunisie). Identification du Sinémurien supérieur dans la Formation Zaghouan. Précisions stratigraphiques et paléobiogéographiques sur le Sinémurien de Tunisie","authors":"P. Fauré, Nejla Sekatni, Mohamed Sabri Arfaoui, R. Alouani","doi":"10.5252/geodiversitas2021v43a4","DOIUrl":"https://doi.org/10.5252/geodiversitas2021v43a4","url":null,"abstract":"RÉSUMÉ La révision de la succession lithostratigraphique du Jurassique du Jebel Ressas (Dorsale tunisienne, Tunisie) permet d'identifier toutes les formations déjà définies par ailleurs dans la Dorsale tunisienne. La Formation Zaghouan y est nouvellement datée du Sinémurien supérieur par des ammonites appartenant aux chronozones à Obtusum (sous-chronozone à Stellare) et à Raricostatum (sous-chronozone à Raricostatum). Six taxons d'Ammonitina appartenant aux genres Asteroceras (A. gr. saltriense (Parona, 1896), A. cf. meridionalis Dommergues, Meister & Mettraux, 1990), Arnioceras (A. gr. ceratitoides (Quenstedt, 1849) – rejectum Fucini, 1902), Epophioceras (E. cf. landrioti (d'Orbigny, 1850)) et Paltechioceras (P. bavaricum (Böse, 1894), P. cf. charpentieri (Schafhäult, 1847)) sont décrits et figurés. Ces ammonites permettent de corréler la « Formation Zaghouan » avec les « Niveaux condensés du sommet de la Formation Oust », déjà bien connus pour leur riche ammonitofaune sinémurienne et dont les âges sont ici réévalués. Les affinités paléobiogéographiques des ammonites du Sinémurien de Tunisie sont discutées.","PeriodicalId":55111,"journal":{"name":"Geodiversitas","volume":"43 1","pages":"95 - 119"},"PeriodicalIF":1.8,"publicationDate":"2021-02-18","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"44910355","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2021-02-11DOI: 10.5252/geodiversitas2021v43a3
A. Pérez‐García, Oier Suárez-Hernando, J. Hernández, S. García, X. Murelaga
ABSTRACT Abundant turtle fossils from two lower Miocene localities in which this lineage had not previously been studied are described herein. The localities are Monte Pila and Fuenmayor, which are 10 km apart, both located in the Autonomous Community of La Rioja (northern Spain), in the Ebro Basin. Information about the turtles from the lower Miocene levels in this basin was until now restricted to the finds made in the MN3 biozone of the Bardenas Reales of Navarre. Several turtle taxa had been recognized there. However, the material attributed to each of them was scarce. Monte Pila and Fuenmayor are located in biozone MN2. The remains of turtles are abundant in both sites. A terrestrial lineage, recognized at Monte Pila, is not represented in the Bardenas Reales fauna of Navarre. It represents the first record of Titanochelon Pérez-García & Vlachos, 2014 in La Rioja and is one of the oldest member of this genus, being assigned to Titanochelon cf. bolivari (Hernández-Pacheco, 1917). The only lineage of turtles identified in Fuenmayor, and the most abundant in Monte Pila, is Ptychogasterinae. All material of this clade of freshwater turtles identified there is compatible with a single form, for which a high range of intraspecific variability is recognized. It is attributed to Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola & Astibia, 1999, this species having been recognized, until now, as exclusive to the Bardenas Reales of Navarre.
{"title":"The freshwater and terrestrial turtles from Monte Pila and Fuenmayor (La Rioja, northern Spain): new data on the lower Miocene turtle diversity of the Ebro Basin","authors":"A. Pérez‐García, Oier Suárez-Hernando, J. Hernández, S. García, X. Murelaga","doi":"10.5252/geodiversitas2021v43a3","DOIUrl":"https://doi.org/10.5252/geodiversitas2021v43a3","url":null,"abstract":"ABSTRACT Abundant turtle fossils from two lower Miocene localities in which this lineage had not previously been studied are described herein. The localities are Monte Pila and Fuenmayor, which are 10 km apart, both located in the Autonomous Community of La Rioja (northern Spain), in the Ebro Basin. Information about the turtles from the lower Miocene levels in this basin was until now restricted to the finds made in the MN3 biozone of the Bardenas Reales of Navarre. Several turtle taxa had been recognized there. However, the material attributed to each of them was scarce. Monte Pila and Fuenmayor are located in biozone MN2. The remains of turtles are abundant in both sites. A terrestrial lineage, recognized at Monte Pila, is not represented in the Bardenas Reales fauna of Navarre. It represents the first record of Titanochelon Pérez-García & Vlachos, 2014 in La Rioja and is one of the oldest member of this genus, being assigned to Titanochelon cf. bolivari (Hernández-Pacheco, 1917). The only lineage of turtles identified in Fuenmayor, and the most abundant in Monte Pila, is Ptychogasterinae. All material of this clade of freshwater turtles identified there is compatible with a single form, for which a high range of intraspecific variability is recognized. It is attributed to Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola & Astibia, 1999, this species having been recognized, until now, as exclusive to the Bardenas Reales of Navarre.","PeriodicalId":55111,"journal":{"name":"Geodiversitas","volume":"43 1","pages":"75 - 94"},"PeriodicalIF":1.8,"publicationDate":"2021-02-11","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"41730582","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2021-01-28DOI: 10.5252/geodiversitas2021v43a2
J. Devillez, S. Charbonnier
ABSTRACT Erymoid lobsters (Crustacea, Decapoda, Erymoidea) are an important component of Mesozoic crustacean faunas in Europe, especially during the Jurassic. With 36 species reported, these lobsters reach their highest diversity during the Late Jurassic. After the review presented here, 23 species belonging to Eryma Meyer, 1840 (11 species), Palaeastacus Bell, 1850 (2 species), Pustulina Quenstedt, 1857 (2 species) and Stenodactylina Beurlen, 1928 (8 species) remain valid. One new species is described: Stenodactylina shotoverigiganti n. sp., and Eryma pseudoventrosa Beurlen, 1928 is integrated to Stenodactylina. We also notice the oldest representative of Enoploclytia M'Coy, 1849, known by a single specimen unidentified at specific level. Eryma ventrosum (Meyer, 1835) is the most common species in Western Europe, and may be seen as emblematic of the Middle-Late Jurassic. Moreover, the lithographic limestones of Germany yield an exceptionally diversified erymoid fauna, with four genera (Eryma, Palaeastacus, Pustulina, Stenodactylina) and 11 species listed. All the Late Jurassic representatives of Palaeastacus were found in this lithology. Finally, the examination of some specimens allows the observation of the strong effects of the decortication on the ornamentation of the erymoids and the resulting taxonomic issues.
{"title":"Review of the Late Jurassic erymoid lobsters (Crustacea: Decapoda)","authors":"J. Devillez, S. Charbonnier","doi":"10.5252/geodiversitas2021v43a2","DOIUrl":"https://doi.org/10.5252/geodiversitas2021v43a2","url":null,"abstract":"ABSTRACT Erymoid lobsters (Crustacea, Decapoda, Erymoidea) are an important component of Mesozoic crustacean faunas in Europe, especially during the Jurassic. With 36 species reported, these lobsters reach their highest diversity during the Late Jurassic. After the review presented here, 23 species belonging to Eryma Meyer, 1840 (11 species), Palaeastacus Bell, 1850 (2 species), Pustulina Quenstedt, 1857 (2 species) and Stenodactylina Beurlen, 1928 (8 species) remain valid. One new species is described: Stenodactylina shotoverigiganti n. sp., and Eryma pseudoventrosa Beurlen, 1928 is integrated to Stenodactylina. We also notice the oldest representative of Enoploclytia M'Coy, 1849, known by a single specimen unidentified at specific level. Eryma ventrosum (Meyer, 1835) is the most common species in Western Europe, and may be seen as emblematic of the Middle-Late Jurassic. Moreover, the lithographic limestones of Germany yield an exceptionally diversified erymoid fauna, with four genera (Eryma, Palaeastacus, Pustulina, Stenodactylina) and 11 species listed. All the Late Jurassic representatives of Palaeastacus were found in this lithology. Finally, the examination of some specimens allows the observation of the strong effects of the decortication on the ornamentation of the erymoids and the resulting taxonomic issues.","PeriodicalId":55111,"journal":{"name":"Geodiversitas","volume":"43 1","pages":"25 - 73"},"PeriodicalIF":1.8,"publicationDate":"2021-01-28","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"43041763","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2021-01-14DOI: 10.5252/geodiversitas2021v43a1
Krister T. Smith, A. Scanferla
ABSTRACT A nearly complete skeleton of an erycine boid is described from the Ypresian-Lutetian (early-middle Eocene) site of Messel, Germany, for which we propose the name Rageryx schmidi n. gen., n. sp. The animal had a total length of c. 52 cm, with c. 258 vertebrae. In skull proportions it is similar to ungaliophiine boids, especially Ungaliophis, and to Tropidophis. The proportions and distinctive accessory processes of the distal caudal vertebrae that are common to all living erycine boids are present in the specimen, although the processes are not as elaborate as in many extant species. The premaxilla was not protruded to form a wedge-shaped snout, and the nasofrontal joint does not appear to show any special buttressing, unlike in many burrowing snake species. Furthermore, the inner ear lacks adaptations to an actively burrowing mode of life. We conclude that the animal, while it was probably secretive, was not fossorial. Phylogenetic analyses using maximum parsimony and Bayesian inference place Rageryx schmidi n. gen., n. sp. unambiguously on the stem of the North American clade (Lichanura + Charina). If this relationship is accurate, it provides further support for a common Euro-American squamate fauna in the early Eocene. The majority of known Messel snake taxa are small-bodied with a small gape, suggesting that such forms may have played a greater role in the early evolutionary radiation of Booidea than their present diversity would suggest.
在德国Messel的Ypresian-Lutetian(早-中始新世)遗址发现了一具几乎完整的赤藓类动物骨架,我们建议将其命名为Rageryx schmidi n. gen., n. sp.该动物全长约52 cm,椎骨约258块。在头骨比例上,它类似于非加利菲亚类,尤其是非加利菲亚类,也类似于Tropidophis。所有活的丹毒生物共有的远尾椎骨的比例和独特的附属过程都存在于标本中,尽管这些过程不像许多现存物种那样复杂。与许多穴居蛇不同的是,它的前颌骨并没有突出来形成楔形的鼻子,鼻前关节似乎也没有任何特殊的支撑。此外,内耳缺乏适应积极的穴居生活方式的能力。我们得出的结论是,这种动物虽然可能是秘密的,但不是化石性的。利用最大简约性和贝叶斯推理的系统发育分析将Rageryx schmidi n. gen., n. sp.明确地定位在北美分支(Lichanura + Charina)的主干上。如果这种关系是准确的,那么它将进一步支持始新世早期欧美共同的鳞片动物群。大多数已知的梅塞尔蛇分类群都是小体小口的,这表明这些形式可能在布idea的早期进化辐射中发挥了比它们现在的多样性更大的作用。
{"title":"A nearly complete skeleton of the oldest definitive erycine boid (Messel, Germany)","authors":"Krister T. Smith, A. Scanferla","doi":"10.5252/geodiversitas2021v43a1","DOIUrl":"https://doi.org/10.5252/geodiversitas2021v43a1","url":null,"abstract":"ABSTRACT A nearly complete skeleton of an erycine boid is described from the Ypresian-Lutetian (early-middle Eocene) site of Messel, Germany, for which we propose the name Rageryx schmidi n. gen., n. sp. The animal had a total length of c. 52 cm, with c. 258 vertebrae. In skull proportions it is similar to ungaliophiine boids, especially Ungaliophis, and to Tropidophis. The proportions and distinctive accessory processes of the distal caudal vertebrae that are common to all living erycine boids are present in the specimen, although the processes are not as elaborate as in many extant species. The premaxilla was not protruded to form a wedge-shaped snout, and the nasofrontal joint does not appear to show any special buttressing, unlike in many burrowing snake species. Furthermore, the inner ear lacks adaptations to an actively burrowing mode of life. We conclude that the animal, while it was probably secretive, was not fossorial. Phylogenetic analyses using maximum parsimony and Bayesian inference place Rageryx schmidi n. gen., n. sp. unambiguously on the stem of the North American clade (Lichanura + Charina). If this relationship is accurate, it provides further support for a common Euro-American squamate fauna in the early Eocene. The majority of known Messel snake taxa are small-bodied with a small gape, suggesting that such forms may have played a greater role in the early evolutionary radiation of Booidea than their present diversity would suggest.","PeriodicalId":55111,"journal":{"name":"Geodiversitas","volume":"43 1","pages":"1 - 24"},"PeriodicalIF":1.8,"publicationDate":"2021-01-14","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"43810033","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2020-12-31DOI: 10.5252/geodiversitas2020v42a30
C. de Muizon, Sandrine Ladevèze
ABSTRACT Andinodelphys cochabambensis Marshall & Muizon, 1988 is one of the best preserved metatherian species from the early Palaeocene fauna of Tiupampa (Bolivia). It is represented by five almost complete skulls, three of them being securely associated to sub-complete to partial skeleton. Four skulls could be extracted from a block including several intermingled skeletons. The present paper provides a thorough description of the dental, cranial, and dentary anatomy of A. cochabambensis. The cranial anatomy of A. cochabambensis is similar to that of Pucadelphys andinus. The skull of Andinodelphys however differs from that of Pucadelphys in its larger size and proportionally longer rostrum. Other differences include the presence, in Andinodelphys, of large anteriorly protruding I1s, small palatal vacuities, a transverse canal, and a small hypotympanic sinus. Andinodelphys has the same dental formula as Pucadelphys (I 5/4, C 1/1, P 3/3, M4/4), the plesiomorphic condition for metatherians. Furthermore, both genera share the lack a tympanic process of the alisphenoid, a deep groove for the internal carotid artery at the anterior apex of the promontorium, a small prootic canal perforating the lateral edge of the petrosal and opening laterally in the deep sulcus for the prootic sinus, and a vestigial anterior lamina of the petrosal. Dentally Andinodelphys closely resembles Pucadelphys, the two genera differing in the larger size of the former and in the inconstant presence in the former of a twinned stylar cusp C. Although 25% smaller, the cheek teeth of Andinodelphys closely resemble those of Itaboraidelphys camposi from the early Eocene of Itaboraí (Brazil). As far as dental morphology is concerned, both genera are likely to have diverged from a direct common ancestor, probably Andinodelphys-like, with Itaboraidelphys displaying more derived dental structures. Two isolated petrosal from Itaboraí (Type 2 petrosals) are morphologically close to those of Andinodelphys but distinctly larger. In this paper, a previous interpretation including the teeth of Itaboraidelphys and these petrosals in the same taxon is followed. A phylogenetic analysis retrieved Itaboraidelphys as a sister taxon of the clade Pucadelphys + Andinodelphys, thus lending support to inclusion of the former in the Pucadelphyidae. Three sets of parsimony analyses were performed. A first set of analyses (with all characters) retrieved a strict consensus tree with a clade as follows: (pucadelphyids, (deltatheroidans (stagodontids, Gurlin Tsav skull-GTS), sparassodonts)). An implied weighting analysis with the same data matrix placed the stagodontids in an early diverging position but retained a clade (pucadelphyids, (deltatheroidans, (GTS, sparassodonts))), the deltatheroidans, being therefore inserted in the pucadelphydans. This result implies an independent arrival of pucadelphyids and sparassodonts to South America, which consequently must have been present in North America in the Lat
{"title":"Cranial anatomy of Andinodelphys cochabambensis, a stem metatherian from the early Palaeocene of Bolivia","authors":"C. de Muizon, Sandrine Ladevèze","doi":"10.5252/geodiversitas2020v42a30","DOIUrl":"https://doi.org/10.5252/geodiversitas2020v42a30","url":null,"abstract":"ABSTRACT Andinodelphys cochabambensis Marshall & Muizon, 1988 is one of the best preserved metatherian species from the early Palaeocene fauna of Tiupampa (Bolivia). It is represented by five almost complete skulls, three of them being securely associated to sub-complete to partial skeleton. Four skulls could be extracted from a block including several intermingled skeletons. The present paper provides a thorough description of the dental, cranial, and dentary anatomy of A. cochabambensis. The cranial anatomy of A. cochabambensis is similar to that of Pucadelphys andinus. The skull of Andinodelphys however differs from that of Pucadelphys in its larger size and proportionally longer rostrum. Other differences include the presence, in Andinodelphys, of large anteriorly protruding I1s, small palatal vacuities, a transverse canal, and a small hypotympanic sinus. Andinodelphys has the same dental formula as Pucadelphys (I 5/4, C 1/1, P 3/3, M4/4), the plesiomorphic condition for metatherians. Furthermore, both genera share the lack a tympanic process of the alisphenoid, a deep groove for the internal carotid artery at the anterior apex of the promontorium, a small prootic canal perforating the lateral edge of the petrosal and opening laterally in the deep sulcus for the prootic sinus, and a vestigial anterior lamina of the petrosal. Dentally Andinodelphys closely resembles Pucadelphys, the two genera differing in the larger size of the former and in the inconstant presence in the former of a twinned stylar cusp C. Although 25% smaller, the cheek teeth of Andinodelphys closely resemble those of Itaboraidelphys camposi from the early Eocene of Itaboraí (Brazil). As far as dental morphology is concerned, both genera are likely to have diverged from a direct common ancestor, probably Andinodelphys-like, with Itaboraidelphys displaying more derived dental structures. Two isolated petrosal from Itaboraí (Type 2 petrosals) are morphologically close to those of Andinodelphys but distinctly larger. In this paper, a previous interpretation including the teeth of Itaboraidelphys and these petrosals in the same taxon is followed. A phylogenetic analysis retrieved Itaboraidelphys as a sister taxon of the clade Pucadelphys + Andinodelphys, thus lending support to inclusion of the former in the Pucadelphyidae. Three sets of parsimony analyses were performed. A first set of analyses (with all characters) retrieved a strict consensus tree with a clade as follows: (pucadelphyids, (deltatheroidans (stagodontids, Gurlin Tsav skull-GTS), sparassodonts)). An implied weighting analysis with the same data matrix placed the stagodontids in an early diverging position but retained a clade (pucadelphyids, (deltatheroidans, (GTS, sparassodonts))), the deltatheroidans, being therefore inserted in the pucadelphydans. This result implies an independent arrival of pucadelphyids and sparassodonts to South America, which consequently must have been present in North America in the Lat","PeriodicalId":55111,"journal":{"name":"Geodiversitas","volume":"42 1","pages":"597 - 739"},"PeriodicalIF":1.8,"publicationDate":"2020-12-31","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"42676459","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2020-12-17DOI: 10.5252/geodiversitas2020v42a29
Jean-Michel Pacaud, Frédéric Sautereau
RÉSUMÉ L'observation sous éclairage ultraviolet de motifs colorés résiduels sur les coquilles de très nombreux gastéropodes cénozoïques fournit des caractères permettant d'affiner la description des espèces fossiles. Nous appliquons ici cette méthode à la discrimination de trois espèces de Cryptochorda de l'Éocène du bassin de Paris et du Cotentin confondues jusqu'ici avec l'espèce classique Cryptochorda (s.str.) stromboides (Hermann, 1781). Ces espèces sont nommées: Cryptochorda (s.str.) altavesna n. sp., C. (s. str.) neptis n. sp. et C. (s.str.) cosediensis n. sp. La répartition stratigraphique et géographique de l'espèce type C. (s.str.) stromboides est ainsi redéfinie. Cryptochorda (s.str.) teae n. sp. est également décrite de l'Yprésien (Éocène inférieur) d'Italie. Une revue historique de la littérature portant sur la taxonomie du genre est donnée et une notice biographique est fournie pour Jean Hermann, descripteur de l'espèce type du genre Cryptochorda.
{"title":"Contribution des motifs colorés résiduels dans la discrimination d'espèces nouvelles de Cryptochorda Mörch, 1858 (Mollusca: Gastropoda: Harpidae) de l'Éocène du bassin de Paris et du Cotentin","authors":"Jean-Michel Pacaud, Frédéric Sautereau","doi":"10.5252/geodiversitas2020v42a29","DOIUrl":"https://doi.org/10.5252/geodiversitas2020v42a29","url":null,"abstract":"RÉSUMÉ L'observation sous éclairage ultraviolet de motifs colorés résiduels sur les coquilles de très nombreux gastéropodes cénozoïques fournit des caractères permettant d'affiner la description des espèces fossiles. Nous appliquons ici cette méthode à la discrimination de trois espèces de Cryptochorda de l'Éocène du bassin de Paris et du Cotentin confondues jusqu'ici avec l'espèce classique Cryptochorda (s.str.) stromboides (Hermann, 1781). Ces espèces sont nommées: Cryptochorda (s.str.) altavesna n. sp., C. (s. str.) neptis n. sp. et C. (s.str.) cosediensis n. sp. La répartition stratigraphique et géographique de l'espèce type C. (s.str.) stromboides est ainsi redéfinie. Cryptochorda (s.str.) teae n. sp. est également décrite de l'Yprésien (Éocène inférieur) d'Italie. Une revue historique de la littérature portant sur la taxonomie du genre est donnée et une notice biographique est fournie pour Jean Hermann, descripteur de l'espèce type du genre Cryptochorda.","PeriodicalId":55111,"journal":{"name":"Geodiversitas","volume":"42 1","pages":"559 - 595"},"PeriodicalIF":1.8,"publicationDate":"2020-12-17","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"45783126","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2020-12-03DOI: 10.5252/geodiversitas2020v42a28
Jozef Klembara, A. Čerňanský
ABSTRACT The anguine species Ophisaurus acuminatus Jörg, 1965 is known on the basis of only one specimen consisting of skull bones and osteoderms from the late Miocene (MN 9) Höwenegg/Hegau locality in Germany. Since its first description, several other new species of Ophisaurus Daudin, 1803 have been described from various Miocene localities in Europe. The diagnoses of these new species are based mostly on characters on the parietal and partially frontal bones. Although most of the cranial elements of O. acuminatus are well preserved, its parietal is absent. The knowledge of the detailed anatomy of this anguine species is crucial for the understanding of its interrelationship within the genus Ophisaurus. For our re-study of this specimen we used a high-resolution X-ray microcomputed tomography. The application of this method enabled: 1) to reveal the anatomy of not visible portions of the previously described bones; 2) to discover the bones completely or almost completely embedded in the sediment; and 3) to identify previously not determined skull bones. Our study enabled to identify three distinguished features for this species and confirmed the validity of the species O. acuminatus.
{"title":"Revision of the cranial anatomy of Ophisaurus acuminatus Jörg, 1965 (Anguimorpha, Anguidae) from the late Miocene of Germany","authors":"Jozef Klembara, A. Čerňanský","doi":"10.5252/geodiversitas2020v42a28","DOIUrl":"https://doi.org/10.5252/geodiversitas2020v42a28","url":null,"abstract":"ABSTRACT The anguine species Ophisaurus acuminatus Jörg, 1965 is known on the basis of only one specimen consisting of skull bones and osteoderms from the late Miocene (MN 9) Höwenegg/Hegau locality in Germany. Since its first description, several other new species of Ophisaurus Daudin, 1803 have been described from various Miocene localities in Europe. The diagnoses of these new species are based mostly on characters on the parietal and partially frontal bones. Although most of the cranial elements of O. acuminatus are well preserved, its parietal is absent. The knowledge of the detailed anatomy of this anguine species is crucial for the understanding of its interrelationship within the genus Ophisaurus. For our re-study of this specimen we used a high-resolution X-ray microcomputed tomography. The application of this method enabled: 1) to reveal the anatomy of not visible portions of the previously described bones; 2) to discover the bones completely or almost completely embedded in the sediment; and 3) to identify previously not determined skull bones. Our study enabled to identify three distinguished features for this species and confirmed the validity of the species O. acuminatus.","PeriodicalId":55111,"journal":{"name":"Geodiversitas","volume":"42 1","pages":"539 - 557"},"PeriodicalIF":1.8,"publicationDate":"2020-12-03","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"44181454","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}