Pub Date : 2025-08-30DOI: 10.1016/j.cretres.2025.106210
Christa-Ch. Hofmann , Ulrich Heimhofer , Emily A. Roberts , Leyla J. Seyfullah
Twelve species of Eucommiidites pollen from deltaic sedimentary rocks of the Rio da Batateira and lowermost Crato formations are here described and depicted with light microscopy (LM) and scanning electron microscopy (SEM). A clump of Eucommiidites sp. 6 from sample CAS 27 of the lowermost Crato Formation is thought to be the dispersed equivalent of in situ pollen of Araripestrobus resiniferous found in the laminitic limestone of the Crato Formation. The other Eucommiidites taxa are known only as dispersed taxa. The separation of individual Eucommiidites species using LM is often very difficult and helped by the use of SEM to observe the position of lateral sulci/furrows, plus the sculpture and ornamentation of the sulcus membrane and ektexine. Only two Eucommiidites taxa, E. sp. 2 and E. sp. 3, occur frequently and generally abundantly in medium percentages (4–15 %) in the pollen sum throughout the section, and are interpreted to come from either wind pollinated plants or plants with no specific edaphic requirements. All other taxa are interpreted to have grown further away from the distributary channels, probably in more specialized stands. Two of them (Eucommiidites sp. 6 and E. sp. 8) are preserved in clumps and are suggested to be animal pollinated. The presence of unusually high percentages of 8–14 % of Eucommiidites in the pollen sums reveal that Eucommiidites taxa constituted a significant portion of the ancient vegetation habitats located upriver, but only comprise a few fragments in the macrofossil record of the Araripe Basin.
{"title":"More diverse and abundant than assumed: Eucommiidites pollen preserved in a deltaic setting (Lower Cretaceous) of the Araripe Basin (NE Brazil)","authors":"Christa-Ch. Hofmann , Ulrich Heimhofer , Emily A. Roberts , Leyla J. Seyfullah","doi":"10.1016/j.cretres.2025.106210","DOIUrl":"10.1016/j.cretres.2025.106210","url":null,"abstract":"<div><div>Twelve species of <em>Eucommiidites</em> pollen from deltaic sedimentary rocks of the Rio da Batateira and lowermost Crato formations are here described and depicted with light microscopy (LM) and scanning electron microscopy (SEM). A clump of <em>Eucommiidites</em> sp. 6 from sample CAS 27 of the lowermost Crato Formation is thought to be the dispersed equivalent of <em>in situ</em> pollen of <em>Araripestrobus resiniferous</em> found in the laminitic limestone of the Crato Formation. The other <em>Eucommiidites</em> taxa are known only as dispersed taxa. The separation of individual <em>Eucommiidites</em> species using LM is often very difficult and helped by the use of SEM to observe the position of lateral sulci/furrows, plus the sculpture and ornamentation of the sulcus membrane and ektexine. Only two <em>Eucommi</em><em>i</em><em>di</em><em>tes</em> taxa<em>, E</em>. sp. 2 and <em>E</em>. sp. 3, occur frequently and generally abundantly in medium percentages (4–15 %) in the pollen sum throughout the section, and are interpreted to come from either wind pollinated plants or plants with no specific edaphic requirements. All other taxa are interpreted to have grown further away from the distributary channels, probably in more specialized stands. Two of them (<em>Eucommiidites</em> sp. 6 and <em>E</em>. sp. 8) are preserved in clumps and are suggested to be animal pollinated. The presence of unusually high percentages of 8–14 % of <em>Eucommiidites</em> in the pollen sums reveal that <em>Eucommiidites</em> taxa constituted a significant portion of the ancient vegetation habitats located upriver, but only comprise a few fragments in the macrofossil record of the Araripe Basin.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"179 ","pages":"Article 106210"},"PeriodicalIF":1.7,"publicationDate":"2025-08-30","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145520350","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-08-28DOI: 10.1016/j.cretres.2025.106211
Cristian D. Benavides-Cabra , María E. Páramo-Fonseca , José A. Narváez-Rincón , Daniel E. Pomar
We describe a new specimen of a lamniform shark from the upper Aptian of the Arcillolitas abigarradas Member of the Paja Formation of Villa de Leiva (Colombia). It represents the first lamniform shark specimen from the Lower Cretaceous with both teeth and vertebrae preserved. The specimen consists of several disarticulated but well-preserved teeth and well-preserved partially articulated vertebral centra, denticles and soft tissues. We refer the specimen to Protolamna ricaurtei, a species recently erected from a specimen found in the same formation. It is a lamniform shark characterized by proportionally small tearing-type teeth with small triangular main cusp, two pairs of small triangular lateral cusplets, and massive bilobated roots. The specimen represents a mature individual to be 6.65 m in total length, making it the oldest record of a gigantic lamniform. This specimen is the first evidence that not all macrophagous lamniforms follow the linear function relating crown height to total body length. Our taphonomic analysis indicates that the specimen must have rapidly reached an anoxic bottom, with low benthic activity, gentle currents, and with microbial mats that facilitated the rapid phosphatization of the soft tissues. Based on the preserved anatomy, we propose that Protolamna ricaurtei was a relatively slow but active swimmer feeding in tropical near-shore areas over small preys such as bony fishes, small sharks, squids, and crustaceans.
我们描述了来自Villa de Leiva(哥伦比亚)Paja组的Arcillolitas abigarradas上Aptian的一种新的板形鲨鱼标本。它代表了下白垩纪第一个保存了牙齿和椎骨的板状鲨鱼标本。标本由几颗脱臼但保存完好的牙齿和保存完好的部分脱臼的椎体中心、牙髓和软组织组成。我们认为该标本属于richaurtei原olamna,这是最近从同一地层中发现的标本中生长出来的一个物种。它是一种板形鲨鱼,其特征是成比例的小撕裂型牙齿,具有小三角形的主尖,两对小三角形的侧尖,和巨大的双叶根。该标本代表了一个成熟的个体,总长度为6.65米,使其成为最古老的巨型板形动物记录。该标本首次证明,并非所有巨噬细胞板状体都遵循冠高与体长的线性关系。我们的埋藏学分析表明,标本必须迅速到达缺氧的底部,底栖生物活动低,水流温和,微生物席促进了软组织的快速磷化。根据保存下来的解剖结构,我们认为ricaurteprotoolamna是一种相对缓慢但活跃的游泳者,以热带近岸地区的小型猎物为食,如硬骨鱼、小鲨鱼、鱿鱼和甲壳类动物。
{"title":"A large lamniform shark from the Aptian of Villa de Leiva (Boyacá, Colombia), based on the first Lower Cretaceous shark specimen preserving both teeth and vertebrae","authors":"Cristian D. Benavides-Cabra , María E. Páramo-Fonseca , José A. Narváez-Rincón , Daniel E. Pomar","doi":"10.1016/j.cretres.2025.106211","DOIUrl":"10.1016/j.cretres.2025.106211","url":null,"abstract":"<div><div>We describe a new specimen of a lamniform shark from the upper Aptian of the Arcillolitas abigarradas Member of the Paja Formation of Villa de Leiva (Colombia). It represents the first lamniform shark specimen from the Lower Cretaceous with both teeth and vertebrae preserved. The specimen consists of several disarticulated but well-preserved teeth and well-preserved partially articulated vertebral centra, denticles and soft tissues. We refer the specimen to <em>Protolamna ricaurtei</em>, a species recently erected from a specimen found in the same formation. It is a lamniform shark characterized by proportionally small tearing-type teeth with small triangular main cusp, two pairs of small triangular lateral cusplets, and massive bilobated roots. The specimen represents a mature individual to be 6.65 m in total length, making it the oldest record of a gigantic lamniform. This specimen is the first evidence that not all macrophagous lamniforms follow the linear function relating crown height to total body length. Our taphonomic analysis indicates that the specimen must have rapidly reached an anoxic bottom, with low benthic activity, gentle currents, and with microbial mats that facilitated the rapid phosphatization of the soft tissues. Based on the preserved anatomy, we propose that <em>Protolamna ricaurtei</em> was a relatively slow but active swimmer feeding in tropical near-shore areas over small preys such as bony fishes, small sharks, squids, and crustaceans.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"178 ","pages":"Article 106211"},"PeriodicalIF":1.7,"publicationDate":"2025-08-28","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145057344","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-08-28DOI: 10.1016/j.cretres.2025.106212
Victor R. Silva , Vitor B. Guerrini , Filipe G. Varejão , Rafael C. Silva , Katie Collins , Ismar S. Carvalho , Simon Schneider , Marcello G. Simões
We revise and describe the Early Cretaceous (Berriasian to Albian) freshwater bivalves from northeastern Brazil, including representatives of Unionida, Trigoniida, and potentially Sphaeriida. Three genera previously thought to be endemic to the Aptian to Albian Crato and Romualdo formations, Araripe Basin, Ceará State, i.e. Cratonaia, Araripenaia, and Monginellopsis, are now recorded from other interior basins. The Berriasian to Barremian Mont Serrat Conglomerate (Salvador Formation) of the Recôncavo Basin, Bahia State, hosts a diverse Unionida-dominated fauna, including Mycetopoda, Anodontites?, Cratonaia, and potentially also hyriids. Specimens tentatively assigned to the neomiodontid Musculiopsis also occur. This fauna has mixed Gondwanan and Laurasian affinities and likely thrived in ephemeral streams and/or floodplain lakes. Araripenaia is recorded from the Aptian Marizal Formation of the Tucano Basin. Supposed representatives of Sphaerium from the same strata are clam shrimps (Diplostraca). Cratonaia and Monginellopsis occur in the Aptian-Albian Itapecuru Formation of the Parnaíba Basin, Maranhão State. Articulated shells of Cratonaia, Araripenaia, and Monginellopsis are typically associated with fluvial-lacustrine deposits. Their common occurrence in Aptian-Albian strata of northeastern Brazil suggests dispersal via fluvial systems connecting the different sedimentary basins. These northeastern Brazilian assemblages are distinct from those of the Upper Cretaceous Bauru Basin in southeastern Brazil. Despite South America and North Africa being connected for most of the Early Cretaceous, no shared freshwater bivalves are known. The newly described faunas provide insight into South American Early Cretaceous freshwater mussel diversity when the proto-South Atlantic was dominated by alluvial and lacustrine settings hosting abundant but poorly known freshwater bivalves.
{"title":"Early Cretaceous freshwater bivalves from northeastern Brazil: Insights into the evolutionary history of South American freshwater mussels","authors":"Victor R. Silva , Vitor B. Guerrini , Filipe G. Varejão , Rafael C. Silva , Katie Collins , Ismar S. Carvalho , Simon Schneider , Marcello G. Simões","doi":"10.1016/j.cretres.2025.106212","DOIUrl":"10.1016/j.cretres.2025.106212","url":null,"abstract":"<div><div>We revise and describe the Early Cretaceous (Berriasian to Albian) freshwater bivalves from northeastern Brazil, including representatives of Unionida, Trigoniida, and potentially Sphaeriida. Three genera previously thought to be endemic to the Aptian to Albian Crato and Romualdo formations, Araripe Basin, Ceará State, i.e. <em>Cratonaia</em>, <em>Araripenaia</em>, and <em>Monginellopsis</em>, are now recorded from other interior basins. The Berriasian to Barremian Mont Serrat Conglomerate (Salvador Formation) of the Recôncavo Basin, Bahia State, hosts a diverse Unionida-dominated fauna, including <em>Mycetopoda</em>, <em>Anodontites</em>?, <em>Cratonaia</em>, and potentially also hyriids. Specimens tentatively assigned to the neomiodontid <em>Musculiopsis</em> also occur. This fauna has mixed Gondwanan and Laurasian affinities and likely thrived in ephemeral streams and/or floodplain lakes. <em>Araripenaia</em> is recorded from the Aptian Marizal Formation of the Tucano Basin. Supposed representatives of <em>Sphaerium</em> from the same strata are clam shrimps (Diplostraca). <em>Cratonaia</em> and <em>Monginellopsis</em> occur in the Aptian-Albian Itapecuru Formation of the Parnaíba Basin, Maranhão State. Articulated shells of <em>Cratonaia</em>, <em>Araripenaia</em>, and <em>Monginellopsis</em> are typically associated with fluvial-lacustrine deposits. Their common occurrence in Aptian-Albian strata of northeastern Brazil suggests dispersal via fluvial systems connecting the different sedimentary basins. These northeastern Brazilian assemblages are distinct from those of the Upper Cretaceous Bauru Basin in southeastern Brazil. Despite South America and North Africa being connected for most of the Early Cretaceous, no shared freshwater bivalves are known. The newly described faunas provide insight into South American Early Cretaceous freshwater mussel diversity when the proto-South Atlantic was dominated by alluvial and lacustrine settings hosting abundant but poorly known freshwater bivalves.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"178 ","pages":"Article 106212"},"PeriodicalIF":1.7,"publicationDate":"2025-08-28","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145061544","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-08-22DOI: 10.1016/j.cretres.2025.106208
Matias Mitidieri , Marianella Talevi , Yanina Herrera , Bruce Rothschild , Marta S. Fernández
Histopathology plays a fundamental role in the detailed analysis of various pathologies, as it allows for diagnosing diseases through microscopic examination of the affected tissues. These studies on Mesozoic marine reptiles are scarce, and almost non-existent in plesiosaurs. Here, two elasmosaurid plesiosaur specimens were examined: specimen MLP-PV 93-I-5-1, recovered from the lower Maastrichtian Snow Hill Island Formation (Vega Island, Antarctica), and specimen MML-PV 5, retrieved from the upper Maastrichtian Jagüel Formation (Salinas de Trapalcó, Río Negro). To obtain the histological sections, a fragmented rib was selected from the first specimen; and a transverse process, dorsal rib, and phalanx were chosen from the second. Pathological bone callus was observed on the rib sections of MLP-PV 93-I-5-1. Periosteal growth perpendicular to the long axis of the bone was observed on the transverse process, dorsal rib, and phalanx sections of MML-PV 5. The characteristics observed in specimen MLP-PV 93-I-5-1 are consistent with the diagnosis of a healing fracture. The characteristics observed in specimen MML-PV 5 are indicative of periostitis. Based on the histopathological study of the two elasmosaurid specimens, it was possible to identify abnormal internal variations in bone tissue and determine that both pathologies developed during the life of the elasmosaurids.
组织病理学在各种病理的详细分析中起着基本的作用,因为它允许通过显微镜检查受影响的组织来诊断疾病。这些关于中生代海洋爬行动物的研究很少,关于蛇颈龙的研究几乎没有。本文研究了两个依拉丝模龙蛇颈龙标本,分别是来自马斯特拉赫特下雪山岛组(Vega Island, Antarctica)的MLP-PV 93- i -5和来自马斯特拉赫特上部jag el组(Salinas de Trapalcó, Río Negro)的MML-PV 5。为了获得组织学切片,从第一个标本中选择了一根破碎的肋骨;而横突,背肋骨和指骨则是从第二处选取的。MLP-PV 93-I-5-1肋骨切片可见病理性骨痂形成。在MML-PV 5的横突、背肋和指骨切片上观察到垂直于骨长轴的骨膜生长。在标本MLP-PV 93-I-5-1中观察到的特征与骨折愈合的诊断一致。在标本MML-PV 5中观察到的特征表明骨膜炎。根据对这两种骨模龙标本的组织病理学研究,可以识别出骨组织的异常内部变异,并确定这两种病理都是在骨模龙的一生中发展起来的。
{"title":"From the inside: Pathologies from a histological perspective of two elasmosaurids from the Upper Cretaceous of Antarctica and Argentina","authors":"Matias Mitidieri , Marianella Talevi , Yanina Herrera , Bruce Rothschild , Marta S. Fernández","doi":"10.1016/j.cretres.2025.106208","DOIUrl":"10.1016/j.cretres.2025.106208","url":null,"abstract":"<div><div>Histopathology plays a fundamental role in the detailed analysis of various pathologies, as it allows for diagnosing diseases through microscopic examination of the affected tissues. These studies on Mesozoic marine reptiles are scarce, and almost non-existent in plesiosaurs. Here, two elasmosaurid plesiosaur specimens were examined: specimen MLP-PV 93-I-5-1, recovered from the lower Maastrichtian Snow Hill Island Formation (Vega Island, Antarctica), and specimen MML-PV 5, retrieved from the upper Maastrichtian Jagüel Formation (Salinas de Trapalcó, Río Negro). To obtain the histological sections, a fragmented rib was selected from the first specimen; and a transverse process, dorsal rib, and phalanx were chosen from the second. Pathological bone callus was observed on the rib sections of MLP-PV 93-I-5-1. Periosteal growth perpendicular to the long axis of the bone was observed on the transverse process, dorsal rib, and phalanx sections of MML-PV 5. The characteristics observed in specimen MLP-PV 93-I-5-1 are consistent with the diagnosis of a healing fracture. The characteristics observed in specimen MML-PV 5 are indicative of periostitis. Based on the histopathological study of the two elasmosaurid specimens, it was possible to identify abnormal internal variations in bone tissue and determine that both pathologies developed during the life of the elasmosaurids.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"177 ","pages":"Article 106208"},"PeriodicalIF":1.7,"publicationDate":"2025-08-22","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144924946","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-08-05DOI: 10.1016/j.cretres.2025.106207
Tojo Chirakal , Jihede Haj Messaoud , Ali Alibrahim , Khalil Ibrahim , Carine Grélaud , Amir Kalifi , John H. Powell , Frans van Buchem
A new integrated age model of the uppermost Albian to Coniacian Ajlun Group in West-Central Jordan is presented based on four complete outcrop sections along a ∼124 km N–S transect. Carbon isotope curves from this work are integrated with published carbon isotope data and constrained by new and existing nannofossil and ammonite biostratigraphy. Key identified carbon isotope events include the Mid-Cenomanian Event 1 (MCE 1), the Oceanic Anoxic Event 2 (OAE 2) at the Cenomanian/Turonian boundary and the Pewsey Event in the middle Turonian. The findings of this study corroborate and revise previous chemostratigraphic definitions in the study area, while also demonstrating a coeval origin of different lithostratigraphic units within the Ajlun Group. On the Arabian Plate scale, a detailed (3rd order) sequence stratigraphic correlation is made between Jordan and time-equivalent strata from the well-studied Natih Formation in Oman. These correlations help to evaluate the relative contributions of eustasy and tectonics on different plate tectonic settings, since the northeastern and eastern margins were heavily influenced by tectonic processes associated with the closure of the Neo-Tethys Ocean. In Oman, the creation of increased accommodation space (ca. 40 m) is observed relative to Jordan commencing in the interval between the MCE 1 and OAE 2. This suggests the onset of tectonic precursor events in Oman during the middle to late Cenomanian interval before the main tectonic phase in the Turonian, resulting in the termination of shallow water carbonate deposition. In contrast, Jordan remained in a largely passive margin setting.
{"title":"Integrated carbon isotope stratigraphy and biostratigraphy of Cenomanian to Turonian carbonates from Jordan – An updated age model and sequence stratigraphic correlations with Oman","authors":"Tojo Chirakal , Jihede Haj Messaoud , Ali Alibrahim , Khalil Ibrahim , Carine Grélaud , Amir Kalifi , John H. Powell , Frans van Buchem","doi":"10.1016/j.cretres.2025.106207","DOIUrl":"10.1016/j.cretres.2025.106207","url":null,"abstract":"<div><div>A new integrated age model of the uppermost Albian to Coniacian Ajlun Group in West-Central Jordan is presented based on four complete outcrop sections along a ∼124 km N–S transect. Carbon isotope curves from this work are integrated with published carbon isotope data and constrained by new and existing nannofossil and ammonite biostratigraphy. Key identified carbon isotope events include the Mid-Cenomanian Event 1 (MCE 1), the Oceanic Anoxic Event 2 (OAE 2) at the Cenomanian/Turonian boundary and the Pewsey Event in the middle Turonian. The findings of this study corroborate and revise previous chemostratigraphic definitions in the study area, while also demonstrating a coeval origin of different lithostratigraphic units within the Ajlun Group. On the Arabian Plate scale, a detailed (3rd order) sequence stratigraphic correlation is made between Jordan and time-equivalent strata from the well-studied Natih Formation in Oman. These correlations help to evaluate the relative contributions of eustasy and tectonics on different plate tectonic settings, since the northeastern and eastern margins were heavily influenced by tectonic processes associated with the closure of the Neo-Tethys Ocean. In Oman, the creation of increased accommodation space (ca. 40 m) is observed relative to Jordan commencing in the interval between the MCE 1 and OAE 2. This suggests the onset of tectonic precursor events in Oman during the middle to late Cenomanian interval before the main tectonic phase in the Turonian, resulting in the termination of shallow water carbonate deposition. In contrast, Jordan remained in a largely passive margin setting.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"177 ","pages":"Article 106207"},"PeriodicalIF":1.7,"publicationDate":"2025-08-05","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144890383","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-08-05DOI: 10.1016/j.cretres.2025.106206
Zhi-Teng Chen , Pavel Sroka , Thomas van de Kamp , Arnold H. Staniczek
This study delves into the taxonomy of the stonefly family Peltoperlidae known from mid-Cretaceous Burmese amber. Based on a broad survey, the fossil family Petroperlidae Sroka, Staniczek & Kondratieff, 2018 is herein considered as a subfamily (Petroperlinae stat. nov.) within Peltoperlidae; and the subfamily Borisoperlinae Chen and Xu, 2020 syn. nov. is synonymized with Petroperlinae stat. nov. The diagnostic characters of all genera of Petroperlinae stat. nov. are revised and summarized. In addition, two new petroperline stoneflies, Graciloperla bulbosa gen. et sp. nov. and Graciloperla stylata gen. et sp. nov., are described based on male adults, which exhibit well-preserved aedeagal structures, the first known for fossil Peltoperlidae.
本研究探讨了中白垩纪缅甸琥珀中已知的石蝇科Peltoperlidae的分类。根据广泛的调查,化石科Petroperlidae Sroka, Staniczek & & & Kondratieff, 2018被认为是Peltoperlidae中的一个亚科(Petroperlinae stat. 11 .);Chen and Xu, 2020 syn11 .与Petroperlinae stat. 11同义。对Petroperlinae stat. 11各属的诊断特征进行了修订和总结。此外,本文还描述了两种新的石蝇,Graciloperla bulbosa gen. et sp. nov.和Graciloperla stylata gen. et sp. nov.,这两种石蝇基于雄性成虫,它们具有保存完好的aedeagal结构,这是在Peltoperlidae化石中首次发现的。
{"title":"Revision of mid-Cretaceous Peltoperlidae (Insecta: Plecoptera) from Burmese amber, with description of two new species in a new genus","authors":"Zhi-Teng Chen , Pavel Sroka , Thomas van de Kamp , Arnold H. Staniczek","doi":"10.1016/j.cretres.2025.106206","DOIUrl":"10.1016/j.cretres.2025.106206","url":null,"abstract":"<div><div>This study delves into the taxonomy of the stonefly family Peltoperlidae known from mid-Cretaceous Burmese amber. Based on a broad survey, the fossil family Petroperlidae Sroka, Staniczek & Kondratieff, 2018 is herein considered as a subfamily (Petroperlinae stat. nov.) within Peltoperlidae; and the subfamily Borisoperlinae Chen and Xu, 2020 syn. nov. is synonymized with Petroperlinae stat. nov. The diagnostic characters of all genera of Petroperlinae stat. nov. are revised and summarized. In addition, two new petroperline stoneflies, <em>Graciloperla bulbosa</em> gen. et sp. nov. and <em>Graciloperla stylata</em> gen. et sp. nov., are described based on male adults, which exhibit well-preserved aedeagal structures, the first known for fossil Peltoperlidae.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"177 ","pages":"Article 106206"},"PeriodicalIF":1.7,"publicationDate":"2025-08-05","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144864882","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
The hybodont shark (Elasmobranchii, Hybodontiformes) material from the Campanian of two non-marine localities in southern France are described. Some teeth and a dorsal fin spine from the upper Campanian of Velaux are assigned to the genus Meristodonoides (Hybodontidae), whereas a single tooth from the lower Campanian of Villeveyrac is referred to Parvodus (Lonchidiidae), a genus hitherto unknown in the Upper Cretaceous. Among the latest Cretaceous continental ichthyofaunas of the European archipelago, hybodont sharks may have been minor relictual components with spatially restricted distributions, as suggested by their scarcity in the fossil assemblages from France and Romania combined with their apparent absence in other areas (Iberian Peninsula, Hungary).
{"title":"Hybodont shark remains from Campanian (Upper Cretaceous) continental deposits of southern France","authors":"Xavier Valentin , Romain Vullo , Gilles Cuny , Olivier Jansen , Géraldine Garcia","doi":"10.1016/j.cretres.2025.106205","DOIUrl":"10.1016/j.cretres.2025.106205","url":null,"abstract":"<div><div>The hybodont shark (Elasmobranchii, Hybodontiformes) material from the Campanian of two non-marine localities in southern France are described. Some teeth and a dorsal fin spine from the upper Campanian of Velaux are assigned to the genus <em>Meristodonoides</em> (Hybodontidae), whereas a single tooth from the lower Campanian of Villeveyrac is referred to <em>Parvodus</em> (Lonchidiidae), a genus hitherto unknown in the Upper Cretaceous. Among the latest Cretaceous continental ichthyofaunas of the European archipelago, hybodont sharks may have been minor relictual components with spatially restricted distributions, as suggested by their scarcity in the fossil assemblages from France and Romania combined with their apparent absence in other areas (Iberian Peninsula, Hungary).</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"177 ","pages":"Article 106205"},"PeriodicalIF":1.7,"publicationDate":"2025-07-21","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144779563","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-07-19DOI: 10.1016/j.cretres.2025.106204
Giulia Amaglio , Maria Rose Petrizzo , Francesca Falzoni , Ann Holbourn , Wolfgang Kuhnt
The Cenomanian–Turonian Oceanic Anoxic Event 2 (OAE 2) severely disrupted the global carbon cycle with widespread deposition of organic-rich marine sediments, resulting in a positive carbon isotope excursion. The Eastbourne section in southeastern England offers extensive benthic and planktonic foraminiferal data, revealing four distinct paleoenvironmental intervals across OAE 2.
The Grey Chalk interval below OAE 2, which is characterized by the highest species diversity of benthic foraminifera, represents outer neritic-upper bathyal paleodepths, oxygenated environments and low organic fluxes at the seafloor. Deep- and thermocline-dwelling planktonic foraminifera suggest meso-oligotrophic regimes with a well-stratified water column.
The onset of OAE 2 in Bed 1a of the Plenus Marl is marked by a sea-level fall supported by the maximum peak in abundance of shallow water agglutinated foraminifera (Ataxophragmium depressum, Arenobulimina, Plectina cenomana) and by the disappearance of bathyal taxa (e.g., Tristix excavata, Kalamopsis grzybowsky).
In Bed 1b of the Plenus Marl, corresponding to the onset of the Plenus Cold Event, Eggerellina, Gaudryina, and Textularia replace shallow agglutinated taxa indicating a transgressive phase. This assemblage also coincides with the occurrence of Boreal planktonic foraminifera that suggests the incursion of Boreal waters into the Anglo-Paris Basin.
The White Chalk, in the upper part and the interval above OAE 2, is interpreted as a Transgressive and Highstand Systems Tract with a change in the benthic foraminiferal assemblage towards the dominance of Marssonella, Gavelinella, Lingulogavelinella and Tritaxia with warmer and more mesotrophic waters recorded by the dominance of Tethyan planktonic foraminifera.
{"title":"Sea-level and paleoenvironmental changes revealed by benthic foraminifera across Oceanic Anoxic Event 2 (OAE 2) at Eastbourne (SE England)","authors":"Giulia Amaglio , Maria Rose Petrizzo , Francesca Falzoni , Ann Holbourn , Wolfgang Kuhnt","doi":"10.1016/j.cretres.2025.106204","DOIUrl":"10.1016/j.cretres.2025.106204","url":null,"abstract":"<div><div>The Cenomanian–Turonian Oceanic Anoxic Event 2 (OAE 2) severely disrupted the global carbon cycle with widespread deposition of organic-rich marine sediments, resulting in a positive carbon isotope excursion. The Eastbourne section in southeastern England offers extensive benthic and planktonic foraminiferal data, revealing four distinct paleoenvironmental intervals across OAE 2.</div><div>The Grey Chalk interval below OAE 2, which is characterized by the highest species diversity of benthic foraminifera, represents outer neritic-upper bathyal paleodepths, oxygenated environments and low organic fluxes at the seafloor. Deep- and thermocline-dwelling planktonic foraminifera suggest meso-oligotrophic regimes with a well-stratified water column.</div><div>The onset of OAE 2 in Bed 1a of the Plenus Marl is marked by a sea-level fall supported by the maximum peak in abundance of shallow water agglutinated foraminifera (<em>Ataxophragmium depressum, Arenobulimina, Plectina cenomana</em>) and by the disappearance of bathyal taxa (e.g., <em>Tristix excavata</em>, <em>Kalamopsis grzybowsky</em>).</div><div>In Bed 1b of the Plenus Marl, corresponding to the onset of the Plenus Cold Event, <em>Eggerellina</em>, <em>Gaudryina</em>, and <em>Textularia</em> replace shallow agglutinated taxa indicating a transgressive phase. This assemblage also coincides with the occurrence of Boreal planktonic foraminifera that suggests the incursion of Boreal waters into the Anglo-Paris Basin.</div><div>The White Chalk, in the upper part and the interval above OAE 2, is interpreted as a Transgressive and Highstand Systems Tract with a change in the benthic foraminiferal assemblage towards the dominance of <em>Marssonella</em>, <em>Gavelinella</em>, <em>Lingulogavelinella</em> and <em>Tritaxia</em> with warmer and more mesotrophic waters recorded by the dominance of Tethyan planktonic foraminifera.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"177 ","pages":"Article 106204"},"PeriodicalIF":1.7,"publicationDate":"2025-07-19","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144810471","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-07-19DOI: 10.1016/j.cretres.2025.106201
Shan Li , Huaichun Wu , Qiang Fang , Runjian Chu , Meinan Shi , Guang Yang , Yuan Gao , Xiaojuan Ma , Youfeng Gao , Chengshan Wang
The continental scientific drilling borehole Songke 3 (SK-3) in the Songliao Basin, Northeastern China, recovers a continuous mid-Cretaceous stratigraphic record which offers a unique opportunity to study terrestrial climate and environmental change. In this study, high-resolution gamma-ray (GR) data were utilized as a paleoclimate proxy to conduct the cyclostratigraphic analysis of the Quantou Formation (K2q) and Denglouku Formation (K2d) within the SK-3 borehole. The cyclic variations in GR values, which reflect fluctuating clay influxes, document Cretaceous Milankovitch cycles including the 405-kyr long eccentricity, ∼100-kyr short eccentricity, ∼38-kyr obliquity, and ∼20-kyr precession cycles. Astronomical tuning of GR data series to the 405-kyr and 100-kyr eccentricity cycles yields an 8.89 ± 0.32 Myr duration for the K2d2 to K2q interval at SK-3. Based on the age of 91.81 ± 0.22 Ma at the boundary of Qingshankou (K2qn) and K2q formations derived from Songke 1 south borehole (SK-1s), our newly constructed astronomical time scale (ATS) suggests that the Turonian/Cenomanian and Cenomanian/Albian boundaries are at the depth ∼1055.5 m within the mid-K2q3 and ∼2350 m within the lower K2d2, respectively. Our study also indicates the complex processes of the sedimentary system in response to the Milankovitch cycles. The eccentricity cycles are more effectively preserved in the lacustrine-deltaic facies. In contrast, both obliquity and precession cycles exhibit better preservation in the meandering river and braided river facies.
{"title":"Astrochronology of the terrestrial mid-Cretaceous Quantou and Denglouku formations in the SK-3 borehole from the Songliao Basin, Northeastern China","authors":"Shan Li , Huaichun Wu , Qiang Fang , Runjian Chu , Meinan Shi , Guang Yang , Yuan Gao , Xiaojuan Ma , Youfeng Gao , Chengshan Wang","doi":"10.1016/j.cretres.2025.106201","DOIUrl":"10.1016/j.cretres.2025.106201","url":null,"abstract":"<div><div>The continental scientific drilling borehole Songke 3 (SK-3) in the Songliao Basin, Northeastern China, recovers a continuous mid-Cretaceous stratigraphic record which offers a unique opportunity to study terrestrial climate and environmental change. In this study, high-resolution gamma-ray (GR) data were utilized as a paleoclimate proxy to conduct the cyclostratigraphic analysis of the Quantou Formation (K<sub>2</sub>q) and Denglouku Formation (K<sub>2</sub>d) within the SK-3 borehole. The cyclic variations in GR values, which reflect fluctuating clay influxes, document Cretaceous Milankovitch cycles including the 405-kyr long eccentricity, ∼100-kyr short eccentricity, ∼38-kyr obliquity, and ∼20-kyr precession cycles. Astronomical tuning of GR data series to the 405-kyr and 100-kyr eccentricity cycles yields an 8.89 ± 0.32 Myr duration for the K<sub>2</sub>d<sup>2</sup> to K<sub>2</sub>q interval at SK-3. Based on the age of 91.81 ± 0.22 Ma at the boundary of Qingshankou (K<sub>2</sub>qn) and K<sub>2</sub>q formations derived from Songke 1 south borehole (SK-1s), our newly constructed astronomical time scale (ATS) suggests that the Turonian/Cenomanian and Cenomanian/Albian boundaries are at the depth ∼1055.5 m within the mid-K<sub>2</sub>q<sup>3</sup> and ∼2350 m within the lower K<sub>2</sub>d<sup>2</sup>, respectively. Our study also indicates the complex processes of the sedimentary system in response to the Milankovitch cycles. The eccentricity cycles are more effectively preserved in the lacustrine-deltaic facies. In contrast, both obliquity and precession cycles exhibit better preservation in the meandering river and braided river facies.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"177 ","pages":"Article 106201"},"PeriodicalIF":1.7,"publicationDate":"2025-07-19","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144757660","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-07-15DOI: 10.1016/j.cretres.2025.106203
Andreas Gärtner , Birgit Niebuhr , H. Tim Breitfeld , Delia Rösel , Marlene C. Schulze , Markus Wilmsen
The Kreibitz-Zittau area (northern margin of the Bohemian Massif, Czech-German border region) exposes a ∼1000-m-thick Cenomanian–middle Coniacian succession of quartz arenites with high compositional but low textural maturity. Translucent heavy minerals are almost exclusively composed of zircon, tourmaline and rutile (Ø-ZTR maturity index of 91) with tourmaline predominating (average of 60 %). Zr-in-rutile temperatures document upper amphibolite-eclogite to granulite facies between 700 and 930 °C. Cr/Nb discrimination of rutile and geochemistry of tourmaline indicate that >95 % of these grains originate from Al-rich and Fe–Mg-poor metapelites. U–Pb-ages of detrital rutile show a distinct peak at 320–330 Ma. If present, garnets of the almandine–pyrope series are dominant. The 95 % predominance of Variscan U–Pb rutile ages in all samples and the high/ultrahigh temperatures of metamorphism exclude the Neoproterozoic greywackes and granitoids of the Lusatian Massif and the granitoids of the Jizera–Krkonoše Massif in the northeast as sources. Potential proto source rocks were high-grade metamorphic rocks similar to those today exposed within the gneissic-migmatitic Góry Sowie Massif, ca. 120 km east of the depositional area. Since the late Devonian, this massif was uplifted, reaching the surface in late Carboniferous times. Consequently, the widespread Permo–Carboniferous Variscan molasse is the major sediment source for the Cretaceous sandstones, which thus were deposited during at least their 2nd sedimentary cycle. The study shows that integrated approaches combining careful petrography, heavy mineral analyses, mineral-specific geochemistry and thermometry as well as U–Pb age dating with high-resolution stratigraphy are suited to solve complex provenance puzzles.
{"title":"Multi-proxy provenance study of sandstones from the northern Bohemian Cretaceous Basin (Germany, Czechia) focused on tourmaline, rutile and garnet","authors":"Andreas Gärtner , Birgit Niebuhr , H. Tim Breitfeld , Delia Rösel , Marlene C. Schulze , Markus Wilmsen","doi":"10.1016/j.cretres.2025.106203","DOIUrl":"10.1016/j.cretres.2025.106203","url":null,"abstract":"<div><div>The Kreibitz-Zittau area (northern margin of the Bohemian Massif, Czech-German border region) exposes a ∼1000-m-thick Cenomanian–middle Coniacian succession of quartz arenites with high compositional but low textural maturity. Translucent heavy minerals are almost exclusively composed of zircon, tourmaline and rutile (Ø-ZTR maturity index of 91) with tourmaline predominating (average of 60 %). Zr-in-rutile temperatures document upper amphibolite-eclogite to granulite facies between 700 and 930 °C. Cr/Nb discrimination of rutile and geochemistry of tourmaline indicate that >95 % of these grains originate from Al-rich and Fe–Mg-poor metapelites. U–Pb-ages of detrital rutile show a distinct peak at 320–330 Ma. If present, garnets of the almandine–pyrope series are dominant. The 95 % predominance of Variscan U–Pb rutile ages in all samples and the high/ultrahigh temperatures of metamorphism exclude the Neoproterozoic greywackes and granitoids of the Lusatian Massif and the granitoids of the Jizera–Krkonoše Massif in the northeast as sources. Potential proto source rocks were high-grade metamorphic rocks similar to those today exposed within the gneissic-migmatitic Góry Sowie Massif, ca. 120 km east of the depositional area. Since the late Devonian, this massif was uplifted, reaching the surface in late Carboniferous times. Consequently, the widespread Permo–Carboniferous Variscan molasse is the major sediment source for the Cretaceous sandstones, which thus were deposited during at least their 2nd sedimentary cycle. The study shows that integrated approaches combining careful petrography, heavy mineral analyses, mineral-specific geochemistry and thermometry as well as U–Pb age dating with high-resolution stratigraphy are suited to solve complex provenance puzzles.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"177 ","pages":"Article 106203"},"PeriodicalIF":1.7,"publicationDate":"2025-07-15","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144722251","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}