Pub Date : 2025-09-03DOI: 10.1016/j.cretres.2025.106213
Noushin Moradi Lirkashasi , Sakineh Arefifard , Thomas J. Algeo , Jonathan D. Schueth
Calcareous nannofossil biostratigraphy has been investigated for Upper Cretaceous and lowermost Paleocene deposits in the Garab section located on the northeastern limb of the Kabir Kuh Anticline, Zagros Basin (Iran). The results reveal a relatively complete record that spans the Cenomanian to Danian stages. The base of the section contains Eiffellithus turriseiffelii, implying placement in or above the UC0 Biozone, and the occurrence of Corollithion kennedyi at 68.8 m confirms placement in the Cenomanian. An unconformity is suggested by the missing UC15a-b Biozone in the upper Campanian Gurpi Formation. The Cenomanian/Turonian Boundary (CTB) and Oceanic Anoxic Event 2 (OAE2) is characterized by abundant small taxa including Biscutum constans, Discorhabdus ignotus, and Zeugrhabdotus noeliae, among which the last two taxa are indicators of surface-water fertility during OAE2. The Cretaceous/Paleogene Boundary (KPB) is marked by an increase of abundance of calcareous nannofossils such as Braarudosphaera bigelowii and Neobiscutum spp. and a rapid increase of calcareous dinoflagellates. Our results document high nannofossil productivity during the OAE2, as observed in other Tethyan locations, and rapid turnover and proliferation of disaster taxa such as Braarudosphaera bigelowii, and Neobiscutum spp. at the KPB. These results significantly enhance our understanding of these geological events and suggest that the Garab section is a strong candidate for further study of both the Cenomanian/Turonian boundary and K/Pg boundary.
{"title":"Calcareous nannofossil biostratigraphy of the upper Cretaceous to lower Paleocene in Zagros Basin, western Iran","authors":"Noushin Moradi Lirkashasi , Sakineh Arefifard , Thomas J. Algeo , Jonathan D. Schueth","doi":"10.1016/j.cretres.2025.106213","DOIUrl":"10.1016/j.cretres.2025.106213","url":null,"abstract":"<div><div>Calcareous nannofossil biostratigraphy has been investigated for Upper Cretaceous and lowermost Paleocene deposits in the Garab section located on the northeastern limb of the Kabir Kuh Anticline, Zagros Basin (Iran). The results reveal a relatively complete record that spans the Cenomanian to Danian stages. The base of the section contains <em>Eiffellithus turriseiffelii</em>, implying placement in or above the UC0 Biozone, and the occurrence of <em>Corollithion kennedyi</em> at 68.8 m confirms placement in the Cenomanian. An unconformity is suggested by the missing UC15a-b Biozone in the upper Campanian Gurpi Formation. The Cenomanian/Turonian Boundary (CTB) and Oceanic Anoxic Event 2 (OAE2) is characterized by abundant small taxa including <em>Biscutum constans</em>, <em>Discorhabdus ignotus</em>, and <em>Zeugrhabdotus noeliae</em>, among which the last two taxa are indicators of surface-water fertility during OAE2. The Cretaceous/Paleogene Boundary (KPB) is marked by an increase of abundance of calcareous nannofossils such as <em>Braarudosphaera bigelowii</em> and <em>Neobiscutum</em> spp. and a rapid increase of calcareous dinoflagellates. Our results document high nannofossil productivity during the OAE2, as observed in other Tethyan locations, and rapid turnover and proliferation of disaster taxa such as <em>Braarudosphaera bigelowii</em>, and <em>Neobiscutum</em> spp. at the KPB. These results significantly enhance our understanding of these geological events and suggest that the Garab section is a strong candidate for further study of both the Cenomanian/Turonian boundary and K/Pg boundary.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"178 ","pages":"Article 106213"},"PeriodicalIF":1.7,"publicationDate":"2025-09-03","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145057343","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-09-03DOI: 10.1016/j.cretres.2025.106217
Miaoqin Lin , Jianguo Li , Yixiao Wu , Tan Tan , Yigang Xu
The International Continental Scientific Drilling Project (ICDP) in the Songliao Basin that has produced a continuous core of the Lower Cretaceous in 4134.81 m thickness, i.e. the SK-II borehole, provides an exceptional archive for studying tectonic and palaeoenvironmental history of the basin under the Cretaceous greenhouse climate. The Shahezi Formation that consists of the main part of this core, represents the primary basin fills at the syn-rift stage of the Songliao Basin and thus marks its commencement and is crucial for studying the timing and mechanism of this basin's early evolution. The precise age of the Shahezi Formation, however, remains controversial and little is known about the palaeovegetation and palaeoenvironment during its deposition. In this study, plenty of pollen and spores are extracted from core samples near the base of the Shahezi Formation. They are the oldest fossils so-far identified from the SK-Ⅱ borehole. The high abundance and diversity of Cicatricosisporites, in association with some other fossils as Pilosisporites, indicate a clear early–middle Aptian age for the base of the Shahezi Formation, i.e. the primary formation of the Songliao Basin. The vegetation was overall featured by abundant ferns, common cycads and some bryophytes in the ground storey, alongside conifer trees or shrubs of Pinaceae, Taxodiaceae, Araucariaceae and Cheirolepidiaceae in the upper canopy, with increases of Cheirolepidiaceae and cycads over time. A generally warm and humid tropical to subtropical climate with a trend to be slightly drier was developed, which might be a response of terrestrial ecosystem to the Oceanic Anoxic Event 1a.
{"title":"Vegetation and climate during the primary formation of the Songliao Basin, NE China","authors":"Miaoqin Lin , Jianguo Li , Yixiao Wu , Tan Tan , Yigang Xu","doi":"10.1016/j.cretres.2025.106217","DOIUrl":"10.1016/j.cretres.2025.106217","url":null,"abstract":"<div><div>The International Continental Scientific Drilling Project (ICDP) in the Songliao Basin that has produced a continuous core of the Lower Cretaceous in 4134.81 m thickness, i.e. the SK-II borehole, provides an exceptional archive for studying tectonic and palaeoenvironmental history of the basin under the Cretaceous greenhouse climate. The Shahezi Formation that consists of the main part of this core, represents the primary basin fills at the syn-rift stage of the Songliao Basin and thus marks its commencement and is crucial for studying the timing and mechanism of this basin's early evolution. The precise age of the Shahezi Formation, however, remains controversial and little is known about the palaeovegetation and palaeoenvironment during its deposition. In this study, plenty of pollen and spores are extracted from core samples near the base of the Shahezi Formation. They are the oldest fossils so-far identified from the SK-Ⅱ borehole. The high abundance and diversity of <em>Cicatricosisporites</em>, in association with some other fossils as <em>Pilosisporites</em>, indicate a clear early–middle Aptian age for the base of the Shahezi Formation, i.e. the primary formation of the Songliao Basin. The vegetation was overall featured by abundant ferns, common cycads and some bryophytes in the ground storey, alongside conifer trees or shrubs of Pinaceae, Taxodiaceae, Araucariaceae and Cheirolepidiaceae in the upper canopy, with increases of Cheirolepidiaceae and cycads over time. A generally warm and humid tropical to subtropical climate with a trend to be slightly drier was developed, which might be a response of terrestrial ecosystem to the Oceanic Anoxic Event 1a.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"178 ","pages":"Article 106217"},"PeriodicalIF":1.7,"publicationDate":"2025-09-03","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145107828","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-09-02DOI: 10.1016/j.cretres.2025.106216
Wei-Wei Wang , Dong Ren , Zhi-Peng Zhao
Nototracans have lived on Earth for over 350 million years, yet limited species records and poor specimen preservation have significantly impeded our understanding of their early evolutionary history. Herein, we supplement the information of a known species and report a new species, Weichangiops trangularis Yang and Hong, 1980 and Weichangiops squamosus sp. nov., from the Lower Cretaceous Yixian Formation in Duolun, Inner Mongolia, China. The diagnosis characters of Weichangiops Yang and Hong, 1980 and W. trangularis were revised based on the new well-preserved specimens. These newly discovered fossil specimens allow for a detailed summary of morphological characteristics in Notostraca, with a systematic review of features observed in Cretaceous lineages. Given the relatively rarity of Cretaceous notostracan fossils, these findings not only expand the known species diversity of this period but also provide valuable insights into the early evolutionary processes of Notostraca.
{"title":"New fossil notostracans (Branchiopoda, Notostraca) from the Lower Cretaceous of Inner Mongolia, China","authors":"Wei-Wei Wang , Dong Ren , Zhi-Peng Zhao","doi":"10.1016/j.cretres.2025.106216","DOIUrl":"10.1016/j.cretres.2025.106216","url":null,"abstract":"<div><div>Nototracans have lived on Earth for over 350 million years, yet limited species records and poor specimen preservation have significantly impeded our understanding of their early evolutionary history. Herein, we supplement the information of a known species and report a new species, <em>Weichangiops trangularis</em> Yang and Hong, 1980 and <em>Weichangiops squamosus</em> sp. nov., from the Lower Cretaceous Yixian Formation in Duolun, Inner Mongolia, China. The diagnosis characters of <em>Weichangiops</em> Yang and Hong, 1980 and <em>W. trangularis</em> were revised based on the new well-preserved specimens. These newly discovered fossil specimens allow for a detailed summary of morphological characteristics in Notostraca, with a systematic review of features observed in Cretaceous lineages. Given the relatively rarity of Cretaceous notostracan fossils, these findings not only expand the known species diversity of this period but also provide valuable insights into the early evolutionary processes of Notostraca.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"178 ","pages":"Article 106216"},"PeriodicalIF":1.7,"publicationDate":"2025-09-02","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145107829","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-08-30DOI: 10.1016/j.cretres.2025.106215
Ian Jarvis , Zofia Dubicka , Markéta Chroustová
The uppermost Turonian – lower Campanian Chalk succession at Seaford Head, southern England, is described and the placement of stage and substage boundaries are defined. Previous studies of foraminifera and ostracods in the section are reviewed. New data are provided for 99 samples spanning the full exposed section. Samples yielded abundant well-preserved diverse microfossil assemblages. The stratigraphic distributions of 38 selected stratigraphically important foraminifera species of Gavelinella, Stensioeina, Protostensioeina, Bolivina, Bolivinoides, Pyramidina, Pseudouvigerina and planktonic foraminifera (Contusotruncana, Globotruncana, Marginotruncana), and 40 ostracod taxa are presented. A list of taxa with taxonomic notes and descriptions of five new foraminifera species, Gavelinella praestelligera, G. praethalmanni, G. praetumida, Protostensioeina ukrainica and Stensioeina praeexsculpta, and three informal species, Pyramidina sp. A, B and C is included. Evolutionary lineages of foraminifera genera provide the basis for a refined biostratigraphy that is successfully correlated to Salzgitter-Salder, Germany and Dubivtsi, Ukraine. Records are compared to literature data and benthic foraminifera zonations across northern Europe. However, differing taxonomic concepts of authors hamper comparison with literature data, exemplified by critical review of previous work at Seaford Head. This precludes rigorous assessment of diachronism of marker species, although regional stratigraphic differences in the distribution of the first stensioeinids and representatives of the Gavelinella clementiana group in Europe are apparent. The first detailed ostracod records from the English Coniacian – Campanian are presented. Four new ostracod biozones defined by the lowest occurrences of the index taxa are proposed: Neocythere (Physocythere) virginea; Limburgina senonensis; Phacorhabdotus lonsdaleianus; and Pterygocythere laticristata zones.
{"title":"Foraminifera and ostracod biostratigraphy of the English Coniacian – Campanian (Upper Cretaceous) Chalk: New results from Seaford Head and correlation to Eastern Europe","authors":"Ian Jarvis , Zofia Dubicka , Markéta Chroustová","doi":"10.1016/j.cretres.2025.106215","DOIUrl":"10.1016/j.cretres.2025.106215","url":null,"abstract":"<div><div>The uppermost Turonian – lower Campanian Chalk succession at Seaford Head, southern England, is described and the placement of stage and substage boundaries are defined. Previous studies of foraminifera and ostracods in the section are reviewed. New data are provided for 99 samples spanning the full exposed section. Samples yielded abundant well-preserved diverse microfossil assemblages. The stratigraphic distributions of 38 selected stratigraphically important foraminifera species of <em>Gavelinella</em>, <em>Stensioeina</em>, <em>Protostensioeina</em>, <em>Bolivina</em>, <em>Bolivinoides</em>, <em>Pyramidina</em>, <em>Pseudouvigerina</em> and planktonic foraminifera (<em>Contusotruncana, Globotruncana, Marginotruncana</em>), and 40 ostracod taxa are presented. A list of taxa with taxonomic notes and descriptions of five new foraminifera species, <em>Gavelinella praestelligera, G. praethalmanni, G. praetumida, Protostensioeina ukrainica</em> and <em>Stensioeina praeexsculpta</em>, and three informal species, <em>Pyramidina</em> sp. A, B and C is included. Evolutionary lineages of foraminifera genera provide the basis for a refined biostratigraphy that is successfully correlated to Salzgitter-Salder, Germany and Dubivtsi, Ukraine. Records are compared to literature data and benthic foraminifera zonations across northern Europe. However, differing taxonomic concepts of authors hamper comparison with literature data, exemplified by critical review of previous work at Seaford Head. This precludes rigorous assessment of diachronism of marker species, although regional stratigraphic differences in the distribution of the first stensioeinids and representatives of the <em>Gavelinella clementiana</em> group in Europe are apparent. The first detailed ostracod records from the English Coniacian – Campanian are presented. Four new ostracod biozones defined by the lowest occurrences of the index taxa are proposed: <em>Neocythere (Physocythere) virginea</em>; <em>Limburgina senonensis</em>; <em>Phacorhabdotus lonsdaleianus</em>; and <em>Pterygocythere laticristata</em> zones.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"178 ","pages":"Article 106215"},"PeriodicalIF":1.7,"publicationDate":"2025-08-30","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145121162","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-08-30DOI: 10.1016/j.cretres.2025.106210
Christa-Ch. Hofmann , Ulrich Heimhofer , Emily A. Roberts , Leyla J. Seyfullah
Twelve species of Eucommiidites pollen from deltaic sedimentary rocks of the Rio da Batateira and lowermost Crato formations are here described and depicted with light microscopy (LM) and scanning electron microscopy (SEM). A clump of Eucommiidites sp. 6 from sample CAS 27 of the lowermost Crato Formation is thought to be the dispersed equivalent of in situ pollen of Araripestrobus resiniferous found in the laminitic limestone of the Crato Formation. The other Eucommiidites taxa are known only as dispersed taxa. The separation of individual Eucommiidites species using LM is often very difficult and helped by the use of SEM to observe the position of lateral sulci/furrows, plus the sculpture and ornamentation of the sulcus membrane and ektexine. Only two Eucommiidites taxa, E. sp. 2 and E. sp. 3, occur frequently and generally abundantly in medium percentages (4–15 %) in the pollen sum throughout the section, and are interpreted to come from either wind pollinated plants or plants with no specific edaphic requirements. All other taxa are interpreted to have grown further away from the distributary channels, probably in more specialized stands. Two of them (Eucommiidites sp. 6 and E. sp. 8) are preserved in clumps and are suggested to be animal pollinated. The presence of unusually high percentages of 8–14 % of Eucommiidites in the pollen sums reveal that Eucommiidites taxa constituted a significant portion of the ancient vegetation habitats located upriver, but only comprise a few fragments in the macrofossil record of the Araripe Basin.
{"title":"More diverse and abundant than assumed: Eucommiidites pollen preserved in a deltaic setting (Lower Cretaceous) of the Araripe Basin (NE Brazil)","authors":"Christa-Ch. Hofmann , Ulrich Heimhofer , Emily A. Roberts , Leyla J. Seyfullah","doi":"10.1016/j.cretres.2025.106210","DOIUrl":"10.1016/j.cretres.2025.106210","url":null,"abstract":"<div><div>Twelve species of <em>Eucommiidites</em> pollen from deltaic sedimentary rocks of the Rio da Batateira and lowermost Crato formations are here described and depicted with light microscopy (LM) and scanning electron microscopy (SEM). A clump of <em>Eucommiidites</em> sp. 6 from sample CAS 27 of the lowermost Crato Formation is thought to be the dispersed equivalent of <em>in situ</em> pollen of <em>Araripestrobus resiniferous</em> found in the laminitic limestone of the Crato Formation. The other <em>Eucommiidites</em> taxa are known only as dispersed taxa. The separation of individual <em>Eucommiidites</em> species using LM is often very difficult and helped by the use of SEM to observe the position of lateral sulci/furrows, plus the sculpture and ornamentation of the sulcus membrane and ektexine. Only two <em>Eucommi</em><em>i</em><em>di</em><em>tes</em> taxa<em>, E</em>. sp. 2 and <em>E</em>. sp. 3, occur frequently and generally abundantly in medium percentages (4–15 %) in the pollen sum throughout the section, and are interpreted to come from either wind pollinated plants or plants with no specific edaphic requirements. All other taxa are interpreted to have grown further away from the distributary channels, probably in more specialized stands. Two of them (<em>Eucommiidites</em> sp. 6 and <em>E</em>. sp. 8) are preserved in clumps and are suggested to be animal pollinated. The presence of unusually high percentages of 8–14 % of <em>Eucommiidites</em> in the pollen sums reveal that <em>Eucommiidites</em> taxa constituted a significant portion of the ancient vegetation habitats located upriver, but only comprise a few fragments in the macrofossil record of the Araripe Basin.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"179 ","pages":"Article 106210"},"PeriodicalIF":1.7,"publicationDate":"2025-08-30","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145520350","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-08-28DOI: 10.1016/j.cretres.2025.106211
Cristian D. Benavides-Cabra , María E. Páramo-Fonseca , José A. Narváez-Rincón , Daniel E. Pomar
We describe a new specimen of a lamniform shark from the upper Aptian of the Arcillolitas abigarradas Member of the Paja Formation of Villa de Leiva (Colombia). It represents the first lamniform shark specimen from the Lower Cretaceous with both teeth and vertebrae preserved. The specimen consists of several disarticulated but well-preserved teeth and well-preserved partially articulated vertebral centra, denticles and soft tissues. We refer the specimen to Protolamna ricaurtei, a species recently erected from a specimen found in the same formation. It is a lamniform shark characterized by proportionally small tearing-type teeth with small triangular main cusp, two pairs of small triangular lateral cusplets, and massive bilobated roots. The specimen represents a mature individual to be 6.65 m in total length, making it the oldest record of a gigantic lamniform. This specimen is the first evidence that not all macrophagous lamniforms follow the linear function relating crown height to total body length. Our taphonomic analysis indicates that the specimen must have rapidly reached an anoxic bottom, with low benthic activity, gentle currents, and with microbial mats that facilitated the rapid phosphatization of the soft tissues. Based on the preserved anatomy, we propose that Protolamna ricaurtei was a relatively slow but active swimmer feeding in tropical near-shore areas over small preys such as bony fishes, small sharks, squids, and crustaceans.
我们描述了来自Villa de Leiva(哥伦比亚)Paja组的Arcillolitas abigarradas上Aptian的一种新的板形鲨鱼标本。它代表了下白垩纪第一个保存了牙齿和椎骨的板状鲨鱼标本。标本由几颗脱臼但保存完好的牙齿和保存完好的部分脱臼的椎体中心、牙髓和软组织组成。我们认为该标本属于richaurtei原olamna,这是最近从同一地层中发现的标本中生长出来的一个物种。它是一种板形鲨鱼,其特征是成比例的小撕裂型牙齿,具有小三角形的主尖,两对小三角形的侧尖,和巨大的双叶根。该标本代表了一个成熟的个体,总长度为6.65米,使其成为最古老的巨型板形动物记录。该标本首次证明,并非所有巨噬细胞板状体都遵循冠高与体长的线性关系。我们的埋藏学分析表明,标本必须迅速到达缺氧的底部,底栖生物活动低,水流温和,微生物席促进了软组织的快速磷化。根据保存下来的解剖结构,我们认为ricaurteprotoolamna是一种相对缓慢但活跃的游泳者,以热带近岸地区的小型猎物为食,如硬骨鱼、小鲨鱼、鱿鱼和甲壳类动物。
{"title":"A large lamniform shark from the Aptian of Villa de Leiva (Boyacá, Colombia), based on the first Lower Cretaceous shark specimen preserving both teeth and vertebrae","authors":"Cristian D. Benavides-Cabra , María E. Páramo-Fonseca , José A. Narváez-Rincón , Daniel E. Pomar","doi":"10.1016/j.cretres.2025.106211","DOIUrl":"10.1016/j.cretres.2025.106211","url":null,"abstract":"<div><div>We describe a new specimen of a lamniform shark from the upper Aptian of the Arcillolitas abigarradas Member of the Paja Formation of Villa de Leiva (Colombia). It represents the first lamniform shark specimen from the Lower Cretaceous with both teeth and vertebrae preserved. The specimen consists of several disarticulated but well-preserved teeth and well-preserved partially articulated vertebral centra, denticles and soft tissues. We refer the specimen to <em>Protolamna ricaurtei</em>, a species recently erected from a specimen found in the same formation. It is a lamniform shark characterized by proportionally small tearing-type teeth with small triangular main cusp, two pairs of small triangular lateral cusplets, and massive bilobated roots. The specimen represents a mature individual to be 6.65 m in total length, making it the oldest record of a gigantic lamniform. This specimen is the first evidence that not all macrophagous lamniforms follow the linear function relating crown height to total body length. Our taphonomic analysis indicates that the specimen must have rapidly reached an anoxic bottom, with low benthic activity, gentle currents, and with microbial mats that facilitated the rapid phosphatization of the soft tissues. Based on the preserved anatomy, we propose that <em>Protolamna ricaurtei</em> was a relatively slow but active swimmer feeding in tropical near-shore areas over small preys such as bony fishes, small sharks, squids, and crustaceans.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"178 ","pages":"Article 106211"},"PeriodicalIF":1.7,"publicationDate":"2025-08-28","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145057344","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-08-28DOI: 10.1016/j.cretres.2025.106212
Victor R. Silva , Vitor B. Guerrini , Filipe G. Varejão , Rafael C. Silva , Katie Collins , Ismar S. Carvalho , Simon Schneider , Marcello G. Simões
We revise and describe the Early Cretaceous (Berriasian to Albian) freshwater bivalves from northeastern Brazil, including representatives of Unionida, Trigoniida, and potentially Sphaeriida. Three genera previously thought to be endemic to the Aptian to Albian Crato and Romualdo formations, Araripe Basin, Ceará State, i.e. Cratonaia, Araripenaia, and Monginellopsis, are now recorded from other interior basins. The Berriasian to Barremian Mont Serrat Conglomerate (Salvador Formation) of the Recôncavo Basin, Bahia State, hosts a diverse Unionida-dominated fauna, including Mycetopoda, Anodontites?, Cratonaia, and potentially also hyriids. Specimens tentatively assigned to the neomiodontid Musculiopsis also occur. This fauna has mixed Gondwanan and Laurasian affinities and likely thrived in ephemeral streams and/or floodplain lakes. Araripenaia is recorded from the Aptian Marizal Formation of the Tucano Basin. Supposed representatives of Sphaerium from the same strata are clam shrimps (Diplostraca). Cratonaia and Monginellopsis occur in the Aptian-Albian Itapecuru Formation of the Parnaíba Basin, Maranhão State. Articulated shells of Cratonaia, Araripenaia, and Monginellopsis are typically associated with fluvial-lacustrine deposits. Their common occurrence in Aptian-Albian strata of northeastern Brazil suggests dispersal via fluvial systems connecting the different sedimentary basins. These northeastern Brazilian assemblages are distinct from those of the Upper Cretaceous Bauru Basin in southeastern Brazil. Despite South America and North Africa being connected for most of the Early Cretaceous, no shared freshwater bivalves are known. The newly described faunas provide insight into South American Early Cretaceous freshwater mussel diversity when the proto-South Atlantic was dominated by alluvial and lacustrine settings hosting abundant but poorly known freshwater bivalves.
{"title":"Early Cretaceous freshwater bivalves from northeastern Brazil: Insights into the evolutionary history of South American freshwater mussels","authors":"Victor R. Silva , Vitor B. Guerrini , Filipe G. Varejão , Rafael C. Silva , Katie Collins , Ismar S. Carvalho , Simon Schneider , Marcello G. Simões","doi":"10.1016/j.cretres.2025.106212","DOIUrl":"10.1016/j.cretres.2025.106212","url":null,"abstract":"<div><div>We revise and describe the Early Cretaceous (Berriasian to Albian) freshwater bivalves from northeastern Brazil, including representatives of Unionida, Trigoniida, and potentially Sphaeriida. Three genera previously thought to be endemic to the Aptian to Albian Crato and Romualdo formations, Araripe Basin, Ceará State, i.e. <em>Cratonaia</em>, <em>Araripenaia</em>, and <em>Monginellopsis</em>, are now recorded from other interior basins. The Berriasian to Barremian Mont Serrat Conglomerate (Salvador Formation) of the Recôncavo Basin, Bahia State, hosts a diverse Unionida-dominated fauna, including <em>Mycetopoda</em>, <em>Anodontites</em>?, <em>Cratonaia</em>, and potentially also hyriids. Specimens tentatively assigned to the neomiodontid <em>Musculiopsis</em> also occur. This fauna has mixed Gondwanan and Laurasian affinities and likely thrived in ephemeral streams and/or floodplain lakes. <em>Araripenaia</em> is recorded from the Aptian Marizal Formation of the Tucano Basin. Supposed representatives of <em>Sphaerium</em> from the same strata are clam shrimps (Diplostraca). <em>Cratonaia</em> and <em>Monginellopsis</em> occur in the Aptian-Albian Itapecuru Formation of the Parnaíba Basin, Maranhão State. Articulated shells of <em>Cratonaia</em>, <em>Araripenaia</em>, and <em>Monginellopsis</em> are typically associated with fluvial-lacustrine deposits. Their common occurrence in Aptian-Albian strata of northeastern Brazil suggests dispersal via fluvial systems connecting the different sedimentary basins. These northeastern Brazilian assemblages are distinct from those of the Upper Cretaceous Bauru Basin in southeastern Brazil. Despite South America and North Africa being connected for most of the Early Cretaceous, no shared freshwater bivalves are known. The newly described faunas provide insight into South American Early Cretaceous freshwater mussel diversity when the proto-South Atlantic was dominated by alluvial and lacustrine settings hosting abundant but poorly known freshwater bivalves.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"178 ","pages":"Article 106212"},"PeriodicalIF":1.7,"publicationDate":"2025-08-28","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145061544","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-08-22DOI: 10.1016/j.cretres.2025.106208
Matias Mitidieri , Marianella Talevi , Yanina Herrera , Bruce Rothschild , Marta S. Fernández
Histopathology plays a fundamental role in the detailed analysis of various pathologies, as it allows for diagnosing diseases through microscopic examination of the affected tissues. These studies on Mesozoic marine reptiles are scarce, and almost non-existent in plesiosaurs. Here, two elasmosaurid plesiosaur specimens were examined: specimen MLP-PV 93-I-5-1, recovered from the lower Maastrichtian Snow Hill Island Formation (Vega Island, Antarctica), and specimen MML-PV 5, retrieved from the upper Maastrichtian Jagüel Formation (Salinas de Trapalcó, Río Negro). To obtain the histological sections, a fragmented rib was selected from the first specimen; and a transverse process, dorsal rib, and phalanx were chosen from the second. Pathological bone callus was observed on the rib sections of MLP-PV 93-I-5-1. Periosteal growth perpendicular to the long axis of the bone was observed on the transverse process, dorsal rib, and phalanx sections of MML-PV 5. The characteristics observed in specimen MLP-PV 93-I-5-1 are consistent with the diagnosis of a healing fracture. The characteristics observed in specimen MML-PV 5 are indicative of periostitis. Based on the histopathological study of the two elasmosaurid specimens, it was possible to identify abnormal internal variations in bone tissue and determine that both pathologies developed during the life of the elasmosaurids.
组织病理学在各种病理的详细分析中起着基本的作用,因为它允许通过显微镜检查受影响的组织来诊断疾病。这些关于中生代海洋爬行动物的研究很少,关于蛇颈龙的研究几乎没有。本文研究了两个依拉丝模龙蛇颈龙标本,分别是来自马斯特拉赫特下雪山岛组(Vega Island, Antarctica)的MLP-PV 93- i -5和来自马斯特拉赫特上部jag el组(Salinas de Trapalcó, Río Negro)的MML-PV 5。为了获得组织学切片,从第一个标本中选择了一根破碎的肋骨;而横突,背肋骨和指骨则是从第二处选取的。MLP-PV 93-I-5-1肋骨切片可见病理性骨痂形成。在MML-PV 5的横突、背肋和指骨切片上观察到垂直于骨长轴的骨膜生长。在标本MLP-PV 93-I-5-1中观察到的特征与骨折愈合的诊断一致。在标本MML-PV 5中观察到的特征表明骨膜炎。根据对这两种骨模龙标本的组织病理学研究,可以识别出骨组织的异常内部变异,并确定这两种病理都是在骨模龙的一生中发展起来的。
{"title":"From the inside: Pathologies from a histological perspective of two elasmosaurids from the Upper Cretaceous of Antarctica and Argentina","authors":"Matias Mitidieri , Marianella Talevi , Yanina Herrera , Bruce Rothschild , Marta S. Fernández","doi":"10.1016/j.cretres.2025.106208","DOIUrl":"10.1016/j.cretres.2025.106208","url":null,"abstract":"<div><div>Histopathology plays a fundamental role in the detailed analysis of various pathologies, as it allows for diagnosing diseases through microscopic examination of the affected tissues. These studies on Mesozoic marine reptiles are scarce, and almost non-existent in plesiosaurs. Here, two elasmosaurid plesiosaur specimens were examined: specimen MLP-PV 93-I-5-1, recovered from the lower Maastrichtian Snow Hill Island Formation (Vega Island, Antarctica), and specimen MML-PV 5, retrieved from the upper Maastrichtian Jagüel Formation (Salinas de Trapalcó, Río Negro). To obtain the histological sections, a fragmented rib was selected from the first specimen; and a transverse process, dorsal rib, and phalanx were chosen from the second. Pathological bone callus was observed on the rib sections of MLP-PV 93-I-5-1. Periosteal growth perpendicular to the long axis of the bone was observed on the transverse process, dorsal rib, and phalanx sections of MML-PV 5. The characteristics observed in specimen MLP-PV 93-I-5-1 are consistent with the diagnosis of a healing fracture. The characteristics observed in specimen MML-PV 5 are indicative of periostitis. Based on the histopathological study of the two elasmosaurid specimens, it was possible to identify abnormal internal variations in bone tissue and determine that both pathologies developed during the life of the elasmosaurids.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"177 ","pages":"Article 106208"},"PeriodicalIF":1.7,"publicationDate":"2025-08-22","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144924946","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-08-05DOI: 10.1016/j.cretres.2025.106207
Tojo Chirakal , Jihede Haj Messaoud , Ali Alibrahim , Khalil Ibrahim , Carine Grélaud , Amir Kalifi , John H. Powell , Frans van Buchem
A new integrated age model of the uppermost Albian to Coniacian Ajlun Group in West-Central Jordan is presented based on four complete outcrop sections along a ∼124 km N–S transect. Carbon isotope curves from this work are integrated with published carbon isotope data and constrained by new and existing nannofossil and ammonite biostratigraphy. Key identified carbon isotope events include the Mid-Cenomanian Event 1 (MCE 1), the Oceanic Anoxic Event 2 (OAE 2) at the Cenomanian/Turonian boundary and the Pewsey Event in the middle Turonian. The findings of this study corroborate and revise previous chemostratigraphic definitions in the study area, while also demonstrating a coeval origin of different lithostratigraphic units within the Ajlun Group. On the Arabian Plate scale, a detailed (3rd order) sequence stratigraphic correlation is made between Jordan and time-equivalent strata from the well-studied Natih Formation in Oman. These correlations help to evaluate the relative contributions of eustasy and tectonics on different plate tectonic settings, since the northeastern and eastern margins were heavily influenced by tectonic processes associated with the closure of the Neo-Tethys Ocean. In Oman, the creation of increased accommodation space (ca. 40 m) is observed relative to Jordan commencing in the interval between the MCE 1 and OAE 2. This suggests the onset of tectonic precursor events in Oman during the middle to late Cenomanian interval before the main tectonic phase in the Turonian, resulting in the termination of shallow water carbonate deposition. In contrast, Jordan remained in a largely passive margin setting.
{"title":"Integrated carbon isotope stratigraphy and biostratigraphy of Cenomanian to Turonian carbonates from Jordan – An updated age model and sequence stratigraphic correlations with Oman","authors":"Tojo Chirakal , Jihede Haj Messaoud , Ali Alibrahim , Khalil Ibrahim , Carine Grélaud , Amir Kalifi , John H. Powell , Frans van Buchem","doi":"10.1016/j.cretres.2025.106207","DOIUrl":"10.1016/j.cretres.2025.106207","url":null,"abstract":"<div><div>A new integrated age model of the uppermost Albian to Coniacian Ajlun Group in West-Central Jordan is presented based on four complete outcrop sections along a ∼124 km N–S transect. Carbon isotope curves from this work are integrated with published carbon isotope data and constrained by new and existing nannofossil and ammonite biostratigraphy. Key identified carbon isotope events include the Mid-Cenomanian Event 1 (MCE 1), the Oceanic Anoxic Event 2 (OAE 2) at the Cenomanian/Turonian boundary and the Pewsey Event in the middle Turonian. The findings of this study corroborate and revise previous chemostratigraphic definitions in the study area, while also demonstrating a coeval origin of different lithostratigraphic units within the Ajlun Group. On the Arabian Plate scale, a detailed (3rd order) sequence stratigraphic correlation is made between Jordan and time-equivalent strata from the well-studied Natih Formation in Oman. These correlations help to evaluate the relative contributions of eustasy and tectonics on different plate tectonic settings, since the northeastern and eastern margins were heavily influenced by tectonic processes associated with the closure of the Neo-Tethys Ocean. In Oman, the creation of increased accommodation space (ca. 40 m) is observed relative to Jordan commencing in the interval between the MCE 1 and OAE 2. This suggests the onset of tectonic precursor events in Oman during the middle to late Cenomanian interval before the main tectonic phase in the Turonian, resulting in the termination of shallow water carbonate deposition. In contrast, Jordan remained in a largely passive margin setting.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"177 ","pages":"Article 106207"},"PeriodicalIF":1.7,"publicationDate":"2025-08-05","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144890383","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-08-05DOI: 10.1016/j.cretres.2025.106206
Zhi-Teng Chen , Pavel Sroka , Thomas van de Kamp , Arnold H. Staniczek
This study delves into the taxonomy of the stonefly family Peltoperlidae known from mid-Cretaceous Burmese amber. Based on a broad survey, the fossil family Petroperlidae Sroka, Staniczek & Kondratieff, 2018 is herein considered as a subfamily (Petroperlinae stat. nov.) within Peltoperlidae; and the subfamily Borisoperlinae Chen and Xu, 2020 syn. nov. is synonymized with Petroperlinae stat. nov. The diagnostic characters of all genera of Petroperlinae stat. nov. are revised and summarized. In addition, two new petroperline stoneflies, Graciloperla bulbosa gen. et sp. nov. and Graciloperla stylata gen. et sp. nov., are described based on male adults, which exhibit well-preserved aedeagal structures, the first known for fossil Peltoperlidae.
本研究探讨了中白垩纪缅甸琥珀中已知的石蝇科Peltoperlidae的分类。根据广泛的调查,化石科Petroperlidae Sroka, Staniczek & & & Kondratieff, 2018被认为是Peltoperlidae中的一个亚科(Petroperlinae stat. 11 .);Chen and Xu, 2020 syn11 .与Petroperlinae stat. 11同义。对Petroperlinae stat. 11各属的诊断特征进行了修订和总结。此外,本文还描述了两种新的石蝇,Graciloperla bulbosa gen. et sp. nov.和Graciloperla stylata gen. et sp. nov.,这两种石蝇基于雄性成虫,它们具有保存完好的aedeagal结构,这是在Peltoperlidae化石中首次发现的。
{"title":"Revision of mid-Cretaceous Peltoperlidae (Insecta: Plecoptera) from Burmese amber, with description of two new species in a new genus","authors":"Zhi-Teng Chen , Pavel Sroka , Thomas van de Kamp , Arnold H. Staniczek","doi":"10.1016/j.cretres.2025.106206","DOIUrl":"10.1016/j.cretres.2025.106206","url":null,"abstract":"<div><div>This study delves into the taxonomy of the stonefly family Peltoperlidae known from mid-Cretaceous Burmese amber. Based on a broad survey, the fossil family Petroperlidae Sroka, Staniczek & Kondratieff, 2018 is herein considered as a subfamily (Petroperlinae stat. nov.) within Peltoperlidae; and the subfamily Borisoperlinae Chen and Xu, 2020 syn. nov. is synonymized with Petroperlinae stat. nov. The diagnostic characters of all genera of Petroperlinae stat. nov. are revised and summarized. In addition, two new petroperline stoneflies, <em>Graciloperla bulbosa</em> gen. et sp. nov. and <em>Graciloperla stylata</em> gen. et sp. nov., are described based on male adults, which exhibit well-preserved aedeagal structures, the first known for fossil Peltoperlidae.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"177 ","pages":"Article 106206"},"PeriodicalIF":1.7,"publicationDate":"2025-08-05","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144864882","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}