Pub Date : 2026-02-01Epub Date: 2025-11-03DOI: 10.1016/j.cretres.2025.106261
Nils Schlüter , Rafika Slami , Madani Benyoucef , Abdelmoumen Garah , W. James Kennedy , Ireneusz Walaszczyk
Thirteen echinoid taxa are described from Coniacian marl-limestone alternations in three sections south and east of Djebel Metlili, on the southwestern margin of the Aures Mountains in northeastern Algeria. The biostratigraphic framework, based on ammonites and inoceramids, indicates an early to middle Coniacian age. The identified taxa include: Rachiosoma rectilineatum, Phymosoma tamarinense, P. cf. P. mansour, Gomphechinus meslei, Coenholectypus subcrassus, Parapygus coquandi, Petalobrissus trigonopygus, P. rimula, Hemiaster sp., and Mecaster fourneli. Three new species are described, Orthopsis metliliae sp. nov., Coenholectypus sulcatus sp. nov. and a new genus and species referred to the family Phymosomatoidae, Triboporus luluatus gen. et sp. nov. A large amount of well-documented material, comprising 579 specimens, was collected, with the fauna being dominated by the spatangoid Mecaster fourneli. At the genus level, the fauna exhibits characteristics typical of the Tethyan realm, and includes Rachiosoma, Gomphechinus, Mecaster, Parapygus and Petalobrissus. However, at the species level, most taxa are restricted to Algeria. Some are also found in adjacent countries such as Tunisia and Egypt.
在阿尔及利亚东北部Aures山脉西南边缘Djebel Metlili以南和以东三个剖面的Coniacian泥灰岩交替中,描述了13个针孔类动物。以菊石和菊石为基础的生物地层格架表明其时代为早至中Coniacian时代。已鉴定的分类群包括:直纹Rachiosoma直纹Rachiosoma tamarinense, P. cf. P. mansour, Gomphechinus meslei, Coenholectypus subcrassus, coquandi, Petalobrissus trigonopygus, P. rimula, Hemiaster sp.和Mecaster fourneli。报告了3个新种(骨科)、3个新属(新种)和1个新属(新种)(藻科)、1个新属(新种)(三角孔虫)(三角孔虫)。收集了579个标本,其中以蛛形目Mecaster fourneli为主。在属水平上,动物群表现出典型的特提斯王国特征,包括Rachiosoma、Gomphechinus、Mecaster、Parapygus和Petalobrissus。然而,在物种水平上,大多数分类群仅限于阿尔及利亚。在突尼斯和埃及等邻近国家也发现了一些。
{"title":"Coniacian (Late Cretaceous) echinoids from the southern Tethyan shelf (NE Algeria) with remarks on their palaeobiology and palaeobiogeography","authors":"Nils Schlüter , Rafika Slami , Madani Benyoucef , Abdelmoumen Garah , W. James Kennedy , Ireneusz Walaszczyk","doi":"10.1016/j.cretres.2025.106261","DOIUrl":"10.1016/j.cretres.2025.106261","url":null,"abstract":"<div><div>Thirteen echinoid taxa are described from Coniacian marl-limestone alternations in three sections south and east of Djebel Metlili, on the southwestern margin of the Aures Mountains in northeastern Algeria. The biostratigraphic framework, based on ammonites and inoceramids, indicates an early to middle Coniacian age. The identified taxa include: <em>Rachiosoma rectilineatum</em>, <em>Phymosoma tamarinense</em>, <em>P</em>. cf. <em>P. mansour</em>, <em>Gomphechinus meslei</em>, <em>Coenholectypus subcrassus, Parapygus coquandi</em>, <em>Petalobrissus trigonopygus</em>, <em>P</em>. <em>rimula</em>, <em>Hemiaster</em> sp., and <em>Mecaster fourneli</em>. Three new species are described, <em>Orthopsis metliliae</em> sp. nov., <em>Coenholectypus sulcatus</em> sp. nov. and a new genus and species referred to the family Phymosomatoidae, <em>Triboporus luluatus</em> gen. et sp. nov. A large amount of well-documented material, comprising 579 specimens, was collected, with the fauna being dominated by the spatangoid <em>Mecaster fourneli</em>. At the genus level, the fauna exhibits characteristics typical of the Tethyan realm, and includes <em>Rachiosoma</em>, <em>Gomphechinus</em>, <em>Mecaster</em>, <em>Parapygus</em> and <em>Petalobrissus</em>. However, at the species level, most taxa are restricted to Algeria. Some are also found in adjacent countries such as Tunisia and Egypt.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"179 ","pages":"Article 106261"},"PeriodicalIF":1.7,"publicationDate":"2026-02-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145579114","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2026-02-01Epub Date: 2025-09-17DOI: 10.1016/j.cretres.2025.106241
Husamaldeen Zubi , Brian F. Platt , Muneer Abdalla , Jennifer Gifford , Belkasim Khameiss
The Tuscaloosa Group (Grp), the basal unit of Upper Cretaceous strata in the northern Gulf of Mexico (GoM), formed during the late Cenomanian transgression across the southeastern United States. Despite its significance, comprehensive sequence stratigraphic analyses of the Tuscaloosa Grp within the south-central Mississippi Embayment (MSE) are limited. This study integrates core data, wireline logs, and field observations to characterize its depositional evolution and sequence stratigraphic architecture, delineating stratigraphic surfaces, systems tracts, and depositional sequences, and examining controlling factors. Four third-order sequences were identified from four sequence boundaries, two transgressive surfaces, and three maximum flooding surfaces. Sequence 1 (S1) includes a lowstand systems tract (LST), transgressive systems tract (TST), and highstand systems tract (HST); Sequence 2 (S2) has a TST and HST; Sequence 3 (S3) contains a LST, TST, and HST; and Sequence 4 (S4) includes a LST and TST. S1 comprises fluvial and incised valley deposits (LST), transitioned to deltaic TST during rising sea levels, and culminated in barrier island and shelf HST. S2 features distal shelf TST deposits followed by a prograding deltaic HST. S3 records a full cycle: fluvial LST, retrograding deltaic TST, and progradational deltaic HST. S4 is less developed, with fluvial LST and localized retrograding deltaic TST deposits. The Tuscaloosa's evolution was driven by accommodation space changes and sediment supply. Subsidence of the MSE in the Late Cretaceous may have increased accommodation and facilitated TST formation in the Upper Tuscaloosa. The Appalachian Highlands supplied most sediments, with the Ouachita Highlands as a secondary source.
{"title":"Depositional system transitions and sequence stratigraphic evolution of the Tuscaloosa Group in the Mississippi Embayment, USA: Accommodation space and sediment supply controls","authors":"Husamaldeen Zubi , Brian F. Platt , Muneer Abdalla , Jennifer Gifford , Belkasim Khameiss","doi":"10.1016/j.cretres.2025.106241","DOIUrl":"10.1016/j.cretres.2025.106241","url":null,"abstract":"<div><div>The Tuscaloosa Group (Grp), the basal unit of Upper Cretaceous strata in the northern Gulf of Mexico (GoM), formed during the late Cenomanian transgression across the southeastern United States. Despite its significance, comprehensive sequence stratigraphic analyses of the Tuscaloosa Grp within the south-central Mississippi Embayment (MSE) are limited. This study integrates core data, wireline logs, and field observations to characterize its depositional evolution and sequence stratigraphic architecture, delineating stratigraphic surfaces, systems tracts, and depositional sequences, and examining controlling factors. Four third-order sequences were identified from four sequence boundaries, two transgressive surfaces, and three maximum flooding surfaces. Sequence 1 (S1) includes a lowstand systems tract (LST), transgressive systems tract (TST), and highstand systems tract (HST); Sequence 2 (S2) has a TST and HST; Sequence 3 (S3) contains a LST, TST, and HST; and Sequence 4 (S4) includes a LST and TST. S1 comprises fluvial and incised valley deposits (LST), transitioned to deltaic TST during rising sea levels, and culminated in barrier island and shelf HST. S2 features distal shelf TST deposits followed by a prograding deltaic HST. S3 records a full cycle: fluvial LST, retrograding deltaic TST, and progradational deltaic HST. S4 is less developed, with fluvial LST and localized retrograding deltaic TST deposits. The Tuscaloosa's evolution was driven by accommodation space changes and sediment supply. Subsidence of the MSE in the Late Cretaceous may have increased accommodation and facilitated TST formation in the Upper Tuscaloosa. The Appalachian Highlands supplied most sediments, with the Ouachita Highlands as a secondary source.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"178 ","pages":"Article 106241"},"PeriodicalIF":1.7,"publicationDate":"2026-02-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145221744","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2026-02-01Epub Date: 2025-10-25DOI: 10.1016/j.cretres.2025.106253
Michael J. Novacek , William M. Wolfe , Maureen A. O'Leary , Eric M. Roberts
We describe Zoslestes gongori, a new species of non-placental eutherian mammal from Upper Cretaceous rocks of Zos Canyon in the Nemegt Basin of the western Gobi Desert, Mongolia. Zoslestes gongori is the largest known zalambdalestid. The Zos Canyon locality, Red Rum, that yielded the specimen is adjacent to beds that have produced several extinct crocodyliforms and the ornithischian dinosaur Haya griva. We provide the first formal stratigraphic description of these fossiliferous Zos Canyon rocks by naming and describing the Zos Formation for the unit at Red Rum. The Zos Formation has a distinct set of sedimentologic associations that differentiate it from the overlying Djadokhta Formation and also has the only fauna in the Nemegt Basin older than that of the Djadokhta Formation. We consider the Zos Formation roughly comparable in age to the Javkhlant Formation of the Eastern Gobi Desert, and likely older than the Alagteeg Formation of the Ulan Nur Basin.
Zoslestes gongori is known from the holotype, a well-preserved posterior cranium, worn dentition, and fragmentary postcranial elements, and one referred specimen. The referred specimen consists of a damaged anterior skull and dentition, and a fragmentary left mandible with roots and alveoli, including an enlarged tooth resembling the elongate first lower incisor of zalambdalestids. Zoslestes shares several derived features with zalambdalestids, including the presence of a prominent entoglenoid process that, in ventral view, partly conceals the anteriorly-positioned postglenoid foramen, a large maxillary tuberosity, multiple hypoglossal foramina, a large, elongate i1 with a horizontally directed root extending deep within the mandible, lower molars with mesiodistally compressed trigonids and relatively wide talonids, a relatively large P4, and upper molars that are distinctly wider labiolingually than long mesiodistally and that are not interlocked mesiodistally. The significantly large size of Zoslestes and the large contribution of its jugal to the zygomatic process are autapomorphies distinguishing this taxon from other zalambdalestids.
{"title":"A new large zalambdalestid mammal from the Gobi Desert and formal stratigraphic description of the Upper Cretaceous Zos Formation","authors":"Michael J. Novacek , William M. Wolfe , Maureen A. O'Leary , Eric M. Roberts","doi":"10.1016/j.cretres.2025.106253","DOIUrl":"10.1016/j.cretres.2025.106253","url":null,"abstract":"<div><div>We describe <em>Zoslestes gongori</em>, a new species of non-placental eutherian mammal from Upper Cretaceous rocks of Zos Canyon in the Nemegt Basin of the western Gobi Desert, Mongolia. <em>Zoslestes gongori</em> is the largest known zalambdalestid. The Zos Canyon locality, Red Rum, that yielded the specimen is adjacent to beds that have produced several extinct crocodyliforms and the ornithischian dinosaur <em>Haya griva</em>. We provide the first formal stratigraphic description of these fossiliferous Zos Canyon rocks by naming and describing the Zos Formation for the unit at Red Rum. The Zos Formation has a distinct set of sedimentologic associations that differentiate it from the overlying Djadokhta Formation and also has the only fauna in the Nemegt Basin older than that of the Djadokhta Formation. We consider the Zos Formation roughly comparable in age to the Javkhlant Formation of the Eastern Gobi Desert, and likely older than the Alagteeg Formation of the Ulan Nur Basin.</div><div><em>Zoslestes gongori</em> is known from the holotype, a well-preserved posterior cranium, worn dentition, and fragmentary postcranial elements, and one referred specimen. The referred specimen consists of a damaged anterior skull and dentition, and a fragmentary left mandible with roots and alveoli, including an enlarged tooth resembling the elongate first lower incisor of zalambdalestids. <em>Zoslestes</em> shares several derived features with zalambdalestids, including the presence of a prominent entoglenoid process that, in ventral view, partly conceals the anteriorly-positioned postglenoid foramen, a large maxillary tuberosity, multiple hypoglossal foramina, a large, elongate i1 with a horizontally directed root extending deep within the mandible, lower molars with mesiodistally compressed trigonids and relatively wide talonids, a relatively large P4, and upper molars that are distinctly wider labiolingually than long mesiodistally and that are not interlocked mesiodistally. The significantly large size of <em>Zoslestes</em> and the large contribution of its jugal to the zygomatic process are autapomorphies distinguishing this taxon from other zalambdalestids.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"179 ","pages":"Article 106253"},"PeriodicalIF":1.7,"publicationDate":"2026-02-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145520351","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2026-02-01Epub Date: 2025-09-15DOI: 10.1016/j.cretres.2025.106235
Javier García-Guerrero , Fredy Parra Ruge , German A. Prieto , Edwin-Alberto Cadena
The fossil record of Lower Cretaceous pliosaurids, particularly from the Valanginian is still poorly known globally, hindering efforts to reconstruct their evolution and palaeogeographical distribution within a broader time frame. Here, we describe an isolated cervical centrum (14 cm maximum diameter) of a pliosaurid belonging to a Brachaucheniinae gen. et sp. indet., from the Lower Cretaceous (upper Valanginian) Rosa Blanca Formation in Zapatoca, Colombia. The presence of large brachaucheniines in the Rosa Blanca Formation suggests that these large pliosaurids were part of the abundant and diverse fauna that inhabited the northwestern margin of Gondwana during the Early Cretaceous and extends their record in northern South America from the Barremian to the late Valanginian. These pliosaurids potentially acted as top predators in the food chain of lower latitude shallow marine environments. This discovery contributes new data to the taxonomic and chronostratigraphic framework of Early Cretaceous marine reptiles from the Proto-Caribbean domain.
下白垩纪上龙的化石记录,特别是来自Valanginian的化石记录,在全球范围内仍然知之甚少,这阻碍了在更广泛的时间框架内重建它们的进化和古地理分布的努力。在这里,我们描述了一个分离的颈椎体(最大直径14厘米),属于brachacheniinae gen. et sp. indet的上龙。来自哥伦比亚萨帕托卡的下白垩纪(上瓦兰吉尼亚)Rosa Blanca组。Rosa Blanca组中大型brachacheniines的存在表明,这些大型上龙是早白垩纪时期居住在冈瓦纳西北边缘的丰富多样的动物群的一部分,并将其在南美洲北部的记录从巴雷米亚延伸到晚瓦兰吉尼亚。这些上龙有可能成为低纬度浅海环境食物链中的顶级捕食者。这一发现为原加勒比海地区早白垩世海生爬行动物的分类和年代地层格架提供了新的数据。
{"title":"Evidence of large pliosaurids in the late Valanginian of Colombia","authors":"Javier García-Guerrero , Fredy Parra Ruge , German A. Prieto , Edwin-Alberto Cadena","doi":"10.1016/j.cretres.2025.106235","DOIUrl":"10.1016/j.cretres.2025.106235","url":null,"abstract":"<div><div>The fossil record of Lower Cretaceous pliosaurids, particularly from the Valanginian is still poorly known globally, hindering efforts to reconstruct their evolution and palaeogeographical distribution within a broader time frame. Here, we describe an isolated cervical centrum (14 cm maximum diameter) of a pliosaurid belonging to a Brachaucheniinae gen. et sp. indet., from the Lower Cretaceous (upper Valanginian) Rosa Blanca Formation in Zapatoca, Colombia. The presence of large brachaucheniines in the Rosa Blanca Formation suggests that these large pliosaurids were part of the abundant and diverse fauna that inhabited the northwestern margin of Gondwana during the Early Cretaceous and extends their record in northern South America from the Barremian to the late Valanginian. These pliosaurids potentially acted as top predators in the food chain of lower latitude shallow marine environments. This discovery contributes new data to the taxonomic and chronostratigraphic framework of Early Cretaceous marine reptiles from the Proto-Caribbean domain.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"178 ","pages":"Article 106235"},"PeriodicalIF":1.7,"publicationDate":"2026-02-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145159477","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2026-02-01Epub Date: 2025-09-03DOI: 10.1016/j.cretres.2025.106213
Noushin Moradi Lirkashasi , Sakineh Arefifard , Thomas J. Algeo , Jonathan D. Schueth
Calcareous nannofossil biostratigraphy has been investigated for Upper Cretaceous and lowermost Paleocene deposits in the Garab section located on the northeastern limb of the Kabir Kuh Anticline, Zagros Basin (Iran). The results reveal a relatively complete record that spans the Cenomanian to Danian stages. The base of the section contains Eiffellithus turriseiffelii, implying placement in or above the UC0 Biozone, and the occurrence of Corollithion kennedyi at 68.8 m confirms placement in the Cenomanian. An unconformity is suggested by the missing UC15a-b Biozone in the upper Campanian Gurpi Formation. The Cenomanian/Turonian Boundary (CTB) and Oceanic Anoxic Event 2 (OAE2) is characterized by abundant small taxa including Biscutum constans, Discorhabdus ignotus, and Zeugrhabdotus noeliae, among which the last two taxa are indicators of surface-water fertility during OAE2. The Cretaceous/Paleogene Boundary (KPB) is marked by an increase of abundance of calcareous nannofossils such as Braarudosphaera bigelowii and Neobiscutum spp. and a rapid increase of calcareous dinoflagellates. Our results document high nannofossil productivity during the OAE2, as observed in other Tethyan locations, and rapid turnover and proliferation of disaster taxa such as Braarudosphaera bigelowii, and Neobiscutum spp. at the KPB. These results significantly enhance our understanding of these geological events and suggest that the Garab section is a strong candidate for further study of both the Cenomanian/Turonian boundary and K/Pg boundary.
{"title":"Calcareous nannofossil biostratigraphy of the upper Cretaceous to lower Paleocene in Zagros Basin, western Iran","authors":"Noushin Moradi Lirkashasi , Sakineh Arefifard , Thomas J. Algeo , Jonathan D. Schueth","doi":"10.1016/j.cretres.2025.106213","DOIUrl":"10.1016/j.cretres.2025.106213","url":null,"abstract":"<div><div>Calcareous nannofossil biostratigraphy has been investigated for Upper Cretaceous and lowermost Paleocene deposits in the Garab section located on the northeastern limb of the Kabir Kuh Anticline, Zagros Basin (Iran). The results reveal a relatively complete record that spans the Cenomanian to Danian stages. The base of the section contains <em>Eiffellithus turriseiffelii</em>, implying placement in or above the UC0 Biozone, and the occurrence of <em>Corollithion kennedyi</em> at 68.8 m confirms placement in the Cenomanian. An unconformity is suggested by the missing UC15a-b Biozone in the upper Campanian Gurpi Formation. The Cenomanian/Turonian Boundary (CTB) and Oceanic Anoxic Event 2 (OAE2) is characterized by abundant small taxa including <em>Biscutum constans</em>, <em>Discorhabdus ignotus</em>, and <em>Zeugrhabdotus noeliae</em>, among which the last two taxa are indicators of surface-water fertility during OAE2. The Cretaceous/Paleogene Boundary (KPB) is marked by an increase of abundance of calcareous nannofossils such as <em>Braarudosphaera bigelowii</em> and <em>Neobiscutum</em> spp. and a rapid increase of calcareous dinoflagellates. Our results document high nannofossil productivity during the OAE2, as observed in other Tethyan locations, and rapid turnover and proliferation of disaster taxa such as <em>Braarudosphaera bigelowii</em>, and <em>Neobiscutum</em> spp. at the KPB. These results significantly enhance our understanding of these geological events and suggest that the Garab section is a strong candidate for further study of both the Cenomanian/Turonian boundary and K/Pg boundary.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"178 ","pages":"Article 106213"},"PeriodicalIF":1.7,"publicationDate":"2026-02-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145057343","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2026-02-01Epub Date: 2025-08-30DOI: 10.1016/j.cretres.2025.106215
Ian Jarvis , Zofia Dubicka , Markéta Chroustová
The uppermost Turonian – lower Campanian Chalk succession at Seaford Head, southern England, is described and the placement of stage and substage boundaries are defined. Previous studies of foraminifera and ostracods in the section are reviewed. New data are provided for 99 samples spanning the full exposed section. Samples yielded abundant well-preserved diverse microfossil assemblages. The stratigraphic distributions of 38 selected stratigraphically important foraminifera species of Gavelinella, Stensioeina, Protostensioeina, Bolivina, Bolivinoides, Pyramidina, Pseudouvigerina and planktonic foraminifera (Contusotruncana, Globotruncana, Marginotruncana), and 40 ostracod taxa are presented. A list of taxa with taxonomic notes and descriptions of five new foraminifera species, Gavelinella praestelligera, G. praethalmanni, G. praetumida, Protostensioeina ukrainica and Stensioeina praeexsculpta, and three informal species, Pyramidina sp. A, B and C is included. Evolutionary lineages of foraminifera genera provide the basis for a refined biostratigraphy that is successfully correlated to Salzgitter-Salder, Germany and Dubivtsi, Ukraine. Records are compared to literature data and benthic foraminifera zonations across northern Europe. However, differing taxonomic concepts of authors hamper comparison with literature data, exemplified by critical review of previous work at Seaford Head. This precludes rigorous assessment of diachronism of marker species, although regional stratigraphic differences in the distribution of the first stensioeinids and representatives of the Gavelinella clementiana group in Europe are apparent. The first detailed ostracod records from the English Coniacian – Campanian are presented. Four new ostracod biozones defined by the lowest occurrences of the index taxa are proposed: Neocythere (Physocythere) virginea; Limburgina senonensis; Phacorhabdotus lonsdaleianus; and Pterygocythere laticristata zones.
{"title":"Foraminifera and ostracod biostratigraphy of the English Coniacian – Campanian (Upper Cretaceous) Chalk: New results from Seaford Head and correlation to Eastern Europe","authors":"Ian Jarvis , Zofia Dubicka , Markéta Chroustová","doi":"10.1016/j.cretres.2025.106215","DOIUrl":"10.1016/j.cretres.2025.106215","url":null,"abstract":"<div><div>The uppermost Turonian – lower Campanian Chalk succession at Seaford Head, southern England, is described and the placement of stage and substage boundaries are defined. Previous studies of foraminifera and ostracods in the section are reviewed. New data are provided for 99 samples spanning the full exposed section. Samples yielded abundant well-preserved diverse microfossil assemblages. The stratigraphic distributions of 38 selected stratigraphically important foraminifera species of <em>Gavelinella</em>, <em>Stensioeina</em>, <em>Protostensioeina</em>, <em>Bolivina</em>, <em>Bolivinoides</em>, <em>Pyramidina</em>, <em>Pseudouvigerina</em> and planktonic foraminifera (<em>Contusotruncana, Globotruncana, Marginotruncana</em>), and 40 ostracod taxa are presented. A list of taxa with taxonomic notes and descriptions of five new foraminifera species, <em>Gavelinella praestelligera, G. praethalmanni, G. praetumida, Protostensioeina ukrainica</em> and <em>Stensioeina praeexsculpta</em>, and three informal species, <em>Pyramidina</em> sp. A, B and C is included. Evolutionary lineages of foraminifera genera provide the basis for a refined biostratigraphy that is successfully correlated to Salzgitter-Salder, Germany and Dubivtsi, Ukraine. Records are compared to literature data and benthic foraminifera zonations across northern Europe. However, differing taxonomic concepts of authors hamper comparison with literature data, exemplified by critical review of previous work at Seaford Head. This precludes rigorous assessment of diachronism of marker species, although regional stratigraphic differences in the distribution of the first stensioeinids and representatives of the <em>Gavelinella clementiana</em> group in Europe are apparent. The first detailed ostracod records from the English Coniacian – Campanian are presented. Four new ostracod biozones defined by the lowest occurrences of the index taxa are proposed: <em>Neocythere (Physocythere) virginea</em>; <em>Limburgina senonensis</em>; <em>Phacorhabdotus lonsdaleianus</em>; and <em>Pterygocythere laticristata</em> zones.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"178 ","pages":"Article 106215"},"PeriodicalIF":1.7,"publicationDate":"2026-02-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145121162","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2026-02-01Epub Date: 2025-09-24DOI: 10.1016/j.cretres.2025.106243
Vladimir N. Makarkin , Arnold H. Staniczek
Rhachisymphrasis raehlei gen. et sp. nov. (Neuroptera: Mantispidae: Symphrasinae) is described from mid-Cretaceous Kachin amber. The genus is remarkable for possessing many character states which do not occur in other Symphrasinae: the profemur is slender; the protibia is very short; the lanceolate process of the probasitarsus arises from its tip and is relatively thin; the mesotibia is strongly expanded; and the forewing CuP is shallowly forked. The protarsus of Rhachisymphrasis gen. nov. is rather similar to that of Rhachiberothidae. However, it is unclear whether Rhachiberothidae is phylogenetically closer to Mantispidae or Berothidae. The most plausible hypothesis is that Rhachiberothidae constitute an ancient branch of Mantispoidea leading to Mantispidae (including Symphrasinae), but retaining many character states of Berothidae. The symphrasine affinity of SinomesomantispaJepson et al., 2013 from the Lower Cretaceous of China is confirmed.
rachisymphrasis raehlei gen. et sp. 11 .(神经翅目:螳螂科:螳螂科)来自中白垩世克钦琥珀。该属的显著特点是具有许多在其他合孢科中所没有的特征状态:柱头纤细;时间很短;近跖骨的披针形突起起于其尖端,相对较薄;中胫肌强烈扩张;前杯呈浅分叉。石竹属的前跗关节与石竹科的前跗关节非常相似。然而,目前尚不清楚在系统发育上Rhachiberothidae是否更接近Mantispidae或Berothidae。最合理的假设是,尾蛛科构成了螳螂科的一个古老分支,导致了螳螂科(包括合蛛科),但保留了许多蛛科的特征状态。中国下白垩统Sinomesomantispa Jepson et al., 2013证实了共phrasine亲和力。
{"title":"A remarkable new genus of Symphrasinae (Neuroptera: Mantispidae) from mid-Cretaceous amber of Myanmar, and the problem of the phylogenetic affinities of Rhachiberothidae","authors":"Vladimir N. Makarkin , Arnold H. Staniczek","doi":"10.1016/j.cretres.2025.106243","DOIUrl":"10.1016/j.cretres.2025.106243","url":null,"abstract":"<div><div><em>Rhachisymphrasis raehlei</em> gen. et sp. nov. (Neuroptera: Mantispidae: Symphrasinae) is described from mid-Cretaceous Kachin amber. The genus is remarkable for possessing many character states which do not occur in other Symphrasinae: the profemur is slender; the protibia is very short; the lanceolate process of the probasitarsus arises from its tip and is relatively thin; the mesotibia is strongly expanded; and the forewing CuP is shallowly forked. The protarsus of <em>Rhachisymphrasis</em> gen. nov. is rather similar to that of Rhachiberothidae. However, it is unclear whether Rhachiberothidae is phylogenetically closer to Mantispidae or Berothidae. The most plausible hypothesis is that Rhachiberothidae constitute an ancient branch of Mantispoidea leading to Mantispidae (including Symphrasinae), but retaining many character states of Berothidae. The symphrasine affinity of <em>Sinomesomantispa</em> <span><span>Jepson et al., 2013</span></span> from the Lower Cretaceous of China is confirmed.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"179 ","pages":"Article 106243"},"PeriodicalIF":1.7,"publicationDate":"2026-02-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145242356","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2026-02-01Epub Date: 2025-11-19DOI: 10.1016/j.cretres.2025.106268
Paolo Sandroni , Nathan S. Church , Rodolfo Coccioni , Fabrizio Frontalini , Maurizio Mainiero , Alessandro Montanari
A group of free climbers by chance discovered a large number of footprint traces deeply impressed on a vast pelagic limestone slab on the steep northeastern limb of the Monte Cònero anticline, near the city of Ancona (Marche region of central-eastern Italy). The footprints probably represent a stampede of panicking sea turtles that were mobilized en masse by an earthquake. These tracks were subsequently covered by a fluxoturbidite triggered by the same earthquake. The same layer is exposed in a 40-m-thick section along the littoral zone below. This new section provides the ability, through combined bio- and magneto-stratigraphic analysis, to place the footprint layer in the lower Campanian foraminiferal biozone Globotruncanita elevata and the lowermost part of magnetochron C33n. Most of this section comprises calcarenitic and calcilutitic turbidites interbedded with pelagic biomicrites, which witnessed a period of enhanced seismic activity exacerbated by a climate change-driven eustatic sea level fluctuation. Following a review of the sedimentological and tectono-seismic history of the Cretaceous Umbria-Marche paleobasin with particular attention to the Monte Cònero area, we describe and document our integrated stratigraphic analysis of the new section exposed along the northeastern littoral known as La Vela Beach.
{"title":"Reptile footprints on a pelagic seafloor as a vestige of a synsedimentary seismic event in the lower Campanian Scaglia Rossa basin of the Umbria-Marche Apennines (Italy)","authors":"Paolo Sandroni , Nathan S. Church , Rodolfo Coccioni , Fabrizio Frontalini , Maurizio Mainiero , Alessandro Montanari","doi":"10.1016/j.cretres.2025.106268","DOIUrl":"10.1016/j.cretres.2025.106268","url":null,"abstract":"<div><div>A group of free climbers by chance discovered a large number of footprint traces deeply impressed on a vast pelagic limestone slab on the steep northeastern limb of the Monte Cònero anticline, near the city of Ancona (Marche region of central-eastern Italy). The footprints probably represent a stampede of panicking sea turtles that were mobilized <em>en masse</em> by an earthquake. These tracks were subsequently covered by a fluxoturbidite triggered by the same earthquake. The same layer is exposed in a 40-m-thick section along the littoral zone below. This new section provides the ability, through combined bio- and magneto-stratigraphic analysis, to place the footprint layer in the lower Campanian foraminiferal biozone <em>Globotruncanita elevata</em> and the lowermost part of magnetochron C33n. Most of this section comprises calcarenitic and calcilutitic turbidites interbedded with pelagic biomicrites, which witnessed a period of enhanced seismic activity exacerbated by a climate change-driven eustatic sea level fluctuation. Following a review of the sedimentological and tectono-seismic history of the Cretaceous Umbria-Marche paleobasin with particular attention to the Monte Cònero area, we describe and document our integrated stratigraphic analysis of the new section exposed along the northeastern littoral known as La Vela Beach.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"179 ","pages":"Article 106268"},"PeriodicalIF":1.7,"publicationDate":"2026-02-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145623594","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2026-02-01Epub Date: 2025-08-28DOI: 10.1016/j.cretres.2025.106211
Cristian D. Benavides-Cabra , María E. Páramo-Fonseca , José A. Narváez-Rincón , Daniel E. Pomar
We describe a new specimen of a lamniform shark from the upper Aptian of the Arcillolitas abigarradas Member of the Paja Formation of Villa de Leiva (Colombia). It represents the first lamniform shark specimen from the Lower Cretaceous with both teeth and vertebrae preserved. The specimen consists of several disarticulated but well-preserved teeth and well-preserved partially articulated vertebral centra, denticles and soft tissues. We refer the specimen to Protolamna ricaurtei, a species recently erected from a specimen found in the same formation. It is a lamniform shark characterized by proportionally small tearing-type teeth with small triangular main cusp, two pairs of small triangular lateral cusplets, and massive bilobated roots. The specimen represents a mature individual to be 6.65 m in total length, making it the oldest record of a gigantic lamniform. This specimen is the first evidence that not all macrophagous lamniforms follow the linear function relating crown height to total body length. Our taphonomic analysis indicates that the specimen must have rapidly reached an anoxic bottom, with low benthic activity, gentle currents, and with microbial mats that facilitated the rapid phosphatization of the soft tissues. Based on the preserved anatomy, we propose that Protolamna ricaurtei was a relatively slow but active swimmer feeding in tropical near-shore areas over small preys such as bony fishes, small sharks, squids, and crustaceans.
我们描述了来自Villa de Leiva(哥伦比亚)Paja组的Arcillolitas abigarradas上Aptian的一种新的板形鲨鱼标本。它代表了下白垩纪第一个保存了牙齿和椎骨的板状鲨鱼标本。标本由几颗脱臼但保存完好的牙齿和保存完好的部分脱臼的椎体中心、牙髓和软组织组成。我们认为该标本属于richaurtei原olamna,这是最近从同一地层中发现的标本中生长出来的一个物种。它是一种板形鲨鱼,其特征是成比例的小撕裂型牙齿,具有小三角形的主尖,两对小三角形的侧尖,和巨大的双叶根。该标本代表了一个成熟的个体,总长度为6.65米,使其成为最古老的巨型板形动物记录。该标本首次证明,并非所有巨噬细胞板状体都遵循冠高与体长的线性关系。我们的埋藏学分析表明,标本必须迅速到达缺氧的底部,底栖生物活动低,水流温和,微生物席促进了软组织的快速磷化。根据保存下来的解剖结构,我们认为ricaurteprotoolamna是一种相对缓慢但活跃的游泳者,以热带近岸地区的小型猎物为食,如硬骨鱼、小鲨鱼、鱿鱼和甲壳类动物。
{"title":"A large lamniform shark from the Aptian of Villa de Leiva (Boyacá, Colombia), based on the first Lower Cretaceous shark specimen preserving both teeth and vertebrae","authors":"Cristian D. Benavides-Cabra , María E. Páramo-Fonseca , José A. Narváez-Rincón , Daniel E. Pomar","doi":"10.1016/j.cretres.2025.106211","DOIUrl":"10.1016/j.cretres.2025.106211","url":null,"abstract":"<div><div>We describe a new specimen of a lamniform shark from the upper Aptian of the Arcillolitas abigarradas Member of the Paja Formation of Villa de Leiva (Colombia). It represents the first lamniform shark specimen from the Lower Cretaceous with both teeth and vertebrae preserved. The specimen consists of several disarticulated but well-preserved teeth and well-preserved partially articulated vertebral centra, denticles and soft tissues. We refer the specimen to <em>Protolamna ricaurtei</em>, a species recently erected from a specimen found in the same formation. It is a lamniform shark characterized by proportionally small tearing-type teeth with small triangular main cusp, two pairs of small triangular lateral cusplets, and massive bilobated roots. The specimen represents a mature individual to be 6.65 m in total length, making it the oldest record of a gigantic lamniform. This specimen is the first evidence that not all macrophagous lamniforms follow the linear function relating crown height to total body length. Our taphonomic analysis indicates that the specimen must have rapidly reached an anoxic bottom, with low benthic activity, gentle currents, and with microbial mats that facilitated the rapid phosphatization of the soft tissues. Based on the preserved anatomy, we propose that <em>Protolamna ricaurtei</em> was a relatively slow but active swimmer feeding in tropical near-shore areas over small preys such as bony fishes, small sharks, squids, and crustaceans.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"178 ","pages":"Article 106211"},"PeriodicalIF":1.7,"publicationDate":"2026-02-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145057344","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2026-02-01Epub Date: 2025-10-15DOI: 10.1016/j.cretres.2025.106250
Lucie Vaňková , Barbara V. Radulović , Martin Košťák , Dragoman Rabrenović , Martin Mazuch , Dejan Barjaktarović , Slađana Dušanić
The Tithonian‒Berriasian boundary interval at the Dedina section (Serbia) has been evaluated from the perspective of rare invertebrate fossils (excluding ammonites and their aptychi) and stable isotope data obtained from the bulk rock samples (δ13Ccarb and δ18Ocarb). The majority of the described benthic fauna consists of brachiopods (Lacunosella hoheneggeri, Tropeothyris tychaviensis, Svaljavithyris bilimeki and Helvetella arzierensis); additionally, a single bivalve species Camptonectes (Camptonectes) cottaldinus is present. This fauna occurs predominantly in the uppermost part of the section, corresponding to the middle Berriasian. Rare belemnites (genera Duvalia, Berriasibelus, Tithonobelus, and Hibolites) exhibit a faunal turnover around the boundary between the calpionellid Remaniella ferasini and Calpionella elliptica subzones, where the family Duvaliidae is largely replaced by Belemnopseidae. The stable carbon isotope data are consistent with previously published records from the north-western Tethys, showing typical slightly negative excursions and trends: at the base of the Calpionella alpina Subzone, within the Ferasini Subzone (a well documented negative δ13Ccarb excursion), and a negative peak within the Elliptica Subzone. The latter may hold potential correlative value but this requires confirmation. The stable oxygen isotope data are more affected by diagenesis, but partially reflect trends observed in other sections and may therefore preserve a palaeoclimatic signal.
{"title":"Macrofauna and stable isotope record of the Jurassic‒Cretaceous transition at the Dedina section (Golubac, eastern Serbia)","authors":"Lucie Vaňková , Barbara V. Radulović , Martin Košťák , Dragoman Rabrenović , Martin Mazuch , Dejan Barjaktarović , Slađana Dušanić","doi":"10.1016/j.cretres.2025.106250","DOIUrl":"10.1016/j.cretres.2025.106250","url":null,"abstract":"<div><div>The Tithonian‒Berriasian boundary interval at the Dedina section (Serbia) has been evaluated from the perspective of rare invertebrate fossils (excluding ammonites and their aptychi) and stable isotope data obtained from the bulk rock samples (δ<sup>13</sup>C<sub>carb</sub> and δ<sup>18</sup>O<sub>carb</sub>). The majority of the described benthic fauna consists of brachiopods (<em>Lacunosella hoheneggeri</em>, <em>Tropeothyris tychaviensis</em>, <em>Svaljavithyris bilimeki</em> and <em>Helvetella arzierensis</em>); additionally, a single bivalve species <em>Camptonectes</em> (<em>Camptonectes</em>) <em>cottaldinus</em> is present. This fauna occurs predominantly in the uppermost part of the section, corresponding to the middle Berriasian. Rare belemnites (genera <em>Duvalia</em>, <em>Berriasibelus</em>, <em>Tithonobelus</em>, and <em>Hibolites</em>) exhibit a faunal turnover around the boundary between the calpionellid <em>Remaniella ferasini</em> and <em>Calpionella elliptica</em> subzones, where the family Duvaliidae is largely replaced by Belemnopseidae. The stable carbon isotope data are consistent with previously published records from the north-western Tethys, showing typical slightly negative excursions and trends: at the base of the <em>Calpionella alpina</em> Subzone, within the <em>Ferasini</em> Subzone (a well documented negative δ<sup>13</sup>C<sub>carb</sub> excursion), and a negative peak within the <em>Elliptica</em> Subzone. The latter may hold potential correlative value but this requires confirmation. The stable oxygen isotope data are more affected by diagenesis, but partially reflect trends observed in other sections and may therefore preserve a palaeoclimatic signal.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"179 ","pages":"Article 106250"},"PeriodicalIF":1.7,"publicationDate":"2026-02-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145466985","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}