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Valanginian ammonite biostratigraphy of the La Peña del Águila section, Zacatecas State, northern Mexico 墨西哥北部萨卡特卡斯州 La Peña del Águila 断面的瓦朗基尼安虫生物地层学
IF 2.1 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2024-02-20 DOI: 10.1016/j.cretres.2024.105865
J.R. Ovando-Figueroa , J.A. Jacobo-Delgado , M. Company , R. Barragán , C.F. Ramírez-Peña , G. Chávez-Cabello , J.A. Moreno-Bedmar

An ammonite biostratigraphic analysis was carried out for the La Peña del Águila section, Zacatecas State, northern Mexico, within Valanginian strata of the Taraises Formation. Bed-by-bed sampling yielded 208 ammonites identified in 13 taxa. The biostratigraphic analysis allowed us to identify two ammonite zones: Neocomites neocomiensiformis and Karakaschiceras inostranzewi; and one subzone: Neocomites platycostatus. These biostratigraphic units range within the lower Valanginian. The identified fauna in the La Peña del Águila section has been previously reported in the Mediterranean region. The faunal similarity allows us to employ the Standard Mediterranean Ammonite Scale for the analysis of the current section. This affinity between the Mexican and Mediterranean ammonite faunas could be explained by the proximity and communication between these two areas through the Hispanic corridor during the Valanginian.

对墨西哥北部萨卡特卡斯州 La Peña del Águila 地段的塔拉伊塞斯地层瓦朗基年地层中的氨水母进行了生物地层分析。通过逐层取样,共鉴定出 13 个类群的 208 只氨甲昆虫。通过生物地层分析,我们确定了两个鹦鹉螺区:Neocomites neocomiensiformis 和 Karakaschiceras inostranzewi;以及一个亚区:Neocomites platycostatus。这些生物地层单元属于下瓦朗基尼期。La Peña del Águila 剖面中已确定的动物群以前在地中海地区也有报道。动物群的相似性使我们能够采用标准地中海鲑鱼尺度来分析当前的剖面。墨西哥和地中海氨石群之间的这种亲缘关系可以解释为在瓦朗基尼期,这两个地区之间通过西班牙走廊进行的接近和交流。
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引用次数: 0
Paleoecology of the foraminifer Acruliammina longa (Tappan, 1940) from the upper Hauterivian of the Neuquén Basin (northern Patagonia, Argentina) 内乌肯盆地(阿根廷巴塔哥尼亚北部)上豪特里维世有孔虫 Acruliammina longa(Tappan,1940 年)的古生态学研究
IF 2.1 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2024-02-15 DOI: 10.1016/j.cretres.2024.105857
M. Caratelli , P. Citton , F. Archuby , J. Pignatti

Herein the epibiont agglutinated placopsilinid foraminifer Acruliammina longa from the upper Hauterivian of the Neuquén Basin is discussed. This is the first record from South America of A. longa, already known from the upper Valanginian to lower Turonian? of North America and Europe. The studied material consists of foraminiferal tests forming macroids and encrusting the valves of Ptychomya koeneni, coming from poorly lithified claystones/siltstones to marlstones forming high frequency (6th-order) depositional sequences within the upper Hauterivian Agua de la Mula Member of the Agrio Formation. This new record extends the paleobiogeographic distribution of A. longa, and allows us to investigate the paleoenvironmental and paleoecological significance of this species in a mixed carbonate and siliciclastic ramp. Energy dispersive spectroscopy and X-ray computed microtomography were used to evaluate taphonomic features of foraminiferal tests, highlighting differences between tests that encrust bivalves and those forming macroids, which also differ in their position within the 3rd-order sedimentary sequences. A. longa tests encrusting bivalves are less damaged, show coarser grain size, and occur throughout the 3rd-order regressive systems tracts, whereas tests forming macroids show higher breakage and finer grain size, occurring throughout the 3rd-order transgressive systems tracts. This distribution allows us to interpret changes in rates of terrigenous input at the time of encrustation, and also in energy conditions, highlighting the opportunistic behavior of A. longa. Low sedimentation rates represent the main paleoenvironmental condition that favored foraminiferal propagule settlement and the growth of A. longa, both on bivalve fragments exposed on the seafloor in the case of macroids, and on living Ptychomya koeneni shells.

本文讨论了来自内乌肯盆地上豪特里维世的附生凝集胎生有孔虫Acruliammina longa。这是南美洲首次记录到长角有孔虫(Acruliammina longa)。所研究的材料包括有孔虫的测试结果,这些测试结果形成了大型有孔虫,并包覆在 Ptychomya koeneni 的瓣膜上,这些测试结果来自岩性较差的粘土岩/粉砂岩到泥灰岩,这些泥灰岩形成了阿格里奥地层上豪特里维世 Agua de la Mula 成员中的高频率(第 6 阶)沉积序列。这一新的记录扩展了长尾鳕的古生物地理分布,使我们能够研究该物种在碳酸盐岩和硅质岩混合斜坡中的古环境和古生态意义。我们利用能量色散光谱法和 X 射线计算机显微层析成像法评估了有孔虫试验的岩石学特征,突出了包覆双壳类动物的试验与形成大型类动物的试验之间的差异,这些试验在三阶沉积序列中的位置也有所不同。包裹双壳类动物的有孔虫化石破坏较少,粒度较粗,出现在整个三阶退行系统道中,而形成大型类群的有孔虫化石破损程度较高,粒度较细,出现在整个三阶退行系统道中。这种分布使我们能够解释在结壳时土著输入率的变化以及能量条件的变化,突出了 A. longa 的机会主义行为。低沉积速率是有孔虫繁殖体沉降和长尾藻生长的主要古环境条件,无论是在暴露于海底的双壳贝类(大型类)碎片上,还是在活着的 Ptychomya koeneni 贝壳上都是如此。
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引用次数: 0
A caenagnathid tibia (Theropoda: Oviraptorosauria) from the upper Campanian Kirtland Formation of New Mexico 新墨西哥州坎帕尼亚上统克特兰德地层中的胫骨(兽脚类:卵翼龙科
IF 2.1 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2024-02-08 DOI: 10.1016/j.cretres.2024.105856
Gregory F. Funston , Thomas E. Williamson , Stephen L. Brusatte

Caenagnathid oviraptorosaurian dinosaurs, long considered enigmas, have now become relatively easy to recognize in the fossil record of Asia and North America. This has revealed their presence in several formations spanning the Late Cretaceous, providing new insights including showing that they were more widespread in the southern parts of North America than previously recognized. Here we add to this record by describing a nearly complete right tibia from the upper Campanian De-na-zin Member of the Kirtland Formation of New Mexico, which represents the first caenagnathid material from these beds. The bone is nearly identical in morphology and size to its counterpart in the Mongolian Elmisaurus rarus, reinforcing a suite of previously-identified features that allow for straightforward recognition of caenagnathid tibiae. The growing body of caenagnathid material continues to blur boundaries between features once considered distinct, raising doubts that some taxa established from sparse material, including Ojoraptorsaurus boerei, are actually valid.

长期以来一直被认为是谜的腔棘龙类恐龙,如今在亚洲和北美洲的化石记录中变得相对容易辨认。这揭示了它们在晚白垩世的几个地层中的存在,提供了新的见解,包括表明它们在北美洲南部的分布比以前认识到的更为广泛。在这里,我们描述了来自新墨西哥州克特兰地层上坎帕尼亚期 De-na-zin 成员的一块近乎完整的右胫骨,这是在这些地层中发现的第一块 caenagnathid 骨。该骨骼在形态和大小上与蒙古榆马龙的对应骨骼几乎完全相同,从而强化了之前确定的一系列特征,使人们可以直接识别出 caenagnathid 的胫骨。越来越多的尾兽脚类动物材料不断模糊曾经被认为是不同特征之间的界限,使人怀疑一些根据稀少材料建立的类群(包括 Ojoraptorsaurus boerei)是否真正有效。
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引用次数: 0
Mortoniceratinae (Ammonoidea) from the lower upper Albian of the Basque-Cantabrian basin (Western Pyrenees): New records, new taxa and their taxonomic and biostratigraphical value 来自巴斯克-坎塔布里亚盆地(西比利牛斯山)上阿尔卑斯山下层的Mortoniceratinae(Amonoidea):新记录、新类群及其分类学和生物地层学价值
IF 2.1 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2024-02-07 DOI: 10.1016/j.cretres.2024.105855
Mikel A. López-Horgue , Hugh G. Owen

The subfamily Mortoniceratinae comprises Albian to early Cenomanian ammonoids whose phylogeny forms the basis of a biozonation for the upper Albian due to their highly evolving nature and worldwide distribution. However, both morphological plasticity of the group and the known common occurrences in many condensed successions call for accurate taxonomy and stratigraphy in order to ascertain the phylogeny and temporal/spatial relationships. Here, early late Albian Mortoniceratinae from mostly uncondensed successions, with a continuous record, are described from a hyperextended pericratonic rift basin in the Western Pyrenees. Although quite rare in the study area, we can here provide a detailed taxonomic study of 55 specimens, representing 19 species, with a single new genus and five new subspecies, all recovered over a period of 30 years from upper Albian strata in eight sedimentary areas covering nearly 1000 square kilometres and more than 4000 m of measured logs. Most of these mortoniceratines show morphologies that are in agreement with their known and accepted phyletic lineage in which tuberculation stages are key; however, we also record tri- and quadrituberculate forms that depart from the main and represent unsuccessful offshoots. A reassessment of Mortoniceras (M.) fallax is presented, inclusive of a more complete, emended diagnosis, and referral of the eponymous biozone to the lower upper Albian. The palaeobiogeographical distribution of species studied corresponds to the western Mediterranean area, part of the central Tethyan Realm, in the Basque-Cantabrian Basin (Western Pyrenees) which faces the northern European basins of the Boreal Realm. This strategic location is interpreted to be a nexus between worldwide basins, as the striking cosmopolitan character of most species studied suggests. The ammonoid fauna studied is proposed as support of the western Mediterranean biozonation; it is tentatively correlated interregionally as well.

Mortoniceratinae亚科由白垩纪至新元古代早期的氨类动物组成,由于其高度演化的性质和全球分布,其系统发育构成了上白垩纪生物区系的基础。然而,由于该类群形态上的可塑性和已知的在许多浓缩演替中的常见性,需要进行准确的分类和地层学研究,以确定其系统发育和时空关系。本文描述了来自西比利牛斯山的一个高延伸围岩裂谷盆地的早期晚阿尔卑斯Mortoniceratinae,它们大多来自非凝聚演替,具有连续的记录。虽然这些标本在该研究地区非常罕见,但我们在此对 55 个标本进行了详细的分类研究,这些标本代表 19 个物种,其中包括一个新属和五个新亚种,这些标本都是历时 30 年从上阿尔比统地层中采集的,分布在 8 个沉积区域,面积近 1000 平方公里,测量的原木长度超过 4000 米。其中大多数 Mortoniceratines 的形态与其已知的、公认的、以瘤化阶段为关键的植物谱系一致;但是,我们也记录到了三瘤化和四瘤化的形态,它们偏离了主系,代表了不成功的分支。本文对 Mortoniceras (M.) fallax 进行了重新评估,包括更完整的修正诊断,并将同名生物区归入上阿尔比统下层。所研究物种的古生物地理学分布与地中海西部地区相对应,该地区是泰提安盆地中部的一部分,位于巴斯克-坎塔布里亚盆地(西比利牛斯山),该盆地面向北欧的北方盆地。这一战略位置被认为是世界盆地之间的连接点,所研究的大多数物种都具有显著的世界性特征。所研究的类氨动物群被认为是西地中海生物区系的支持者;它在区域间也具有初步的相关性。
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引用次数: 0
The first report of Lower Cretaceous corals from the Baghin section, west of Kerman, Iran 首次报道伊朗克尔曼以西巴格欣地段的下白垩世珊瑚
IF 2.1 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2024-02-05 DOI: 10.1016/j.cretres.2024.105851
Toran Khamoshi, Ahmad Lotfabad Arab, Mohammad Reza Vaziri

In the Baghin section, west of Kerman, Iran, Aptian scleractinian corals are described for the first time in relation to taxonomically related genera and species. The superfamilies are Actinastraeoidea, Cladocoroidea, Cyclolitoidea, Eugyroidea, Stylinoidea, Montlivaltioidea, Caryophyllioidea and Misistelloidea. Other macrofossil groups are echinoids, gastropods, brachiopods and bivalves and confirm the Aptian age. Regarding identified corals, it was concluded that the study area can be correlated with Aptian-Albian of other Lower Cretaceous coral faunas of Basque-Cantabrian Basin in the Central Tethys, and also most closely related to faunas of the Aptian Pelagonium South margin and the Aptian -Albian Bisbee Basin in the western Atlantic.

在伊朗克尔曼以西的 Baghin 地段,首次描述了始新世硬手珊瑚与分类学上相关的属和种的关系。其超科为 Actinastraeoidea、Cladocoroidea、Cyclolitoidea、Eugyroidea、Stylinoidea、Montlivaltioidea、Caryophyllioidea 和 Misistelloidea。其他大型化石类群包括回声类、腹足类、腕足类和双壳类,证实了该地区的始新世年龄。关于已鉴定的珊瑚,结论是该研究区域可与特提斯中部巴斯克-坎塔布里亚盆地的其他下白垩世珊瑚动物群的始新世-白垩世时期相关联,也与大西洋西部始新世-白垩世时期的伯拉贡尼姆南缘和始新世-白垩世时期的比斯比盆地的动物群关系最为密切。
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引用次数: 0
Integrated stratigraphy across the Jurassic–Cretaceous boundary in the Rettenbacher section (Northern Calcareous Alps, Salzburg, Austria) 雷滕巴赫(Rettenbacher)段侏罗纪-白垩纪界线的综合地层学(奥地利萨尔茨堡北钙质阿尔卑斯山
IF 2.1 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2024-02-01 DOI: 10.1016/j.cretres.2024.105854
T. Elbra , P. Skupien , M. Bubík , M. Košťák , M. Molčan Matejová , P. Pruner , D. Reháková , L. Švábenická , L. Vaňková , V. Cígler , J. Geist , Š. Kdýr , A. Lukeneder , P. Rybová , M. Mazuch , P. Schnabl , A. Svobodová , J. Trubač , H. Ucar

Rettenbacher Quarry carbonate sequence in the Northern Calcareous Alps represents highly dynamic pelagic carbonate sedimentation in the lower slope environment. The section spans from upper Tithonian Crassicollaria colomi Subzone to middle Berriasian Calpionella elliptica Subzone. Radiolarian samples belong to Pseudodictyomitra carpatica Zone. Considerable redeposition of calpionellids, organic walled dinoflagellate cysts, calcareous nannofossils and foraminifera in the allodapic layers, as well as extensive normal polarity remagnetisation event with substantial clockwise rotation is documented. Quantitative predominance of genera Watznaueria and Cyclagelosphaera indicate strong nannofossil modification during diagenesis. The almost straight trend in the δ13C with some slight carbon-isotope perturbations is recorded at Rettenbacher. Three significant and stratigraphically important negative excursions of the δ13Ccarb values were identified — including tentative indication of a new “Negative peak 4” in the upper part of the Elliptica Subzone — and correlated with other sections confirming stability, respectively isochronity within carbon chemostratigraphy.

北钙质阿尔卑斯山的雷滕巴赫采石场碳酸盐序列代表了下斜坡环境中高度动态的浮游碳酸盐沉积。该剖面从上提托尼世的 Crassicollaria colomi 亚区到中贝里亚世的 Calpionella elliptica 亚区。放射虫样本属于假鲤科鲤属(Pseudodictyomitra carpatica)区。在异钙化层中有大量的钙藻类、有机壁甲藻胞囊、钙质化石和有孔虫重新沉积,并记录了大量顺时针旋转的正常极性再磁化事件。Watznaueria属和Cyclagelosphaera属在数量上占优势,表明在成岩过程中,有孔虫化石发生了强烈的改变。在雷滕巴赫(Rettenbacher),δ13C 几乎呈直线趋势,并伴有一些轻微的碳同位素扰动。发现了δ13Ccarb值的三个重要的、地层学上重要的负偏移--包括在埃利普卡亚区上部的一个新的 "负峰4 "的初步迹象--并与其他断面相关联,证实了碳化学地层学中的稳定性和等时性。
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引用次数: 0
Body fossil assemblages from the Lower Idzików Beds (Coniacian) in the Upper Nysa Kłodzka Graben, south-west Poland: Preliminary taphonomic, palaeoecological and palaeogeographical data 波兰西南部上Nysa Kłodzka地堑下Idzików床(科尼亚克期)的体化石组合:初步的岩石学、古生态学和古地理学数据
IF 2.1 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2024-02-01 DOI: 10.1016/j.cretres.2024.105853
Alina Chrząstek , Elena A. Jagt-Yazykova , John W.M. Jagt

From mudstones and sandstones referred to as the Lower Idzików Beds, as exposed at Stary Waliszów (Upper Nysa Kłodzka Graben, south-west Poland), a rich assemblage of body fossils has been recovered. This comprises rare ammonites (Baculites incurvatus, Forresteria cf. petrocoriensis, ?Scalarites sp.), nautiloids (Anglonautilus sinuatoplicatus), numerous bivalves, inclusive of inoceramids such as Inoceramus frechi and Volviceramus involutus, oysters (Ceratostreon, Vultogryphaea and Hyotissa) and other common genera such as Scabrotrigonia, Protocardia, Cucullaea, Neithea, Pinna, Liopistha, Panopea and Goniomya. Gastropods include mainly aporrhaids, turritellids, naticids and ampullinids. Echinoids are rare and comparatively poorly preserved, comprising only irregular forms (cardiasterids, micrasterids and hemiasterids), some of them being recorded from the Upper Cretaceous of Poland for the first time. Decapod crustaceans are fairly abundant in the Lower Idzików Beds, with axiideans predominating. A taphonomic analysis shows that this assemblage is well preserved, suggesting rapid burial as a result of storm events, while a palaeoecological assessment of taxonomic composition and trophic modes implies that deposition took place in a shallow-marine, well-oxygenated environment with a soft-bottom substrate, rich in nutrients and close to the fair-weather wave base. The presence of stenohaline taxa such as ammonites and echinoderms provides evidence of normal salinity.

在 Stary Waliszów(波兰西南部,上 Nysa Kłodzka 地堑)出露的被称为下 Idzików 床的泥岩和砂岩中,发现了丰富的尸体化石。其中包括罕见的氨虫(Baculites incurvatus、Forresteria cf. petrocoriensis、Scalarites sp.)、鹦鹉螺(Anglonautilus sinuatoplicatus)、大量双壳类,包括 Inoceramus frechi 和 Volviceramus involutus 等 Inoceramus 类、牡蛎(Ceratostreon、Vultogryphaea 和 Hyotissa)以及 Scabrotrigonia、Protocardia、Cucullaea、Neithea、Pinna、Liopistha、Panopea 和 Goniomya 等其他常见属。腹足类主要包括porrhaids、turritellids、naticids 和 ampullinids。棘皮动物很少见,而且保存相对较差,只有不规则的形态(贲门虫类、小贲门虫类和半贲门虫类),其中一些还是首次在波兰的上白垩世记录到。十足类甲壳动物在 Idzików 下白垩统床层中相当丰富,其中以xiideans 类为主。岩石学分析表明,该集合体保存完好,表明是风暴事件导致的快速埋藏,而对分类组成和营养模式的古生态学评估则表明,沉积发生在一个浅海、氧气充足、底质松软、富含营养物质且靠近风浪基底的环境中。氨虫和棘皮动物等盐碱类群的存在为正常盐度提供了证据。
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引用次数: 0
Two new kateretid beetle species (Coleoptera: Kateretidae) with specialized antennal scapes in males from mid-Cretaceous Kachin amber 白垩纪中期克钦琥珀中发现的两个雄性具有特化触角的甲虫新种(鞘翅目:甲虫科
IF 2.1 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2024-01-29 DOI: 10.1016/j.cretres.2024.105848
Qian Zhao , David Peris , Jia Liu , Diying Huang , Chenyang Cai

Kateretidae is a beetle family with 15 reported fossil species, four of them with specialized antennal scape. Specialized scape represents a rare form of sexual dimorphism in Kateretidae. Here we report two new species with a special scape in males of Protokateretes Zhao, Huang and Cai, 2023, and one female individual of an unnamed species of Protokateretes. The discovery of new species provides a supplementary diagnosis of Protokateretes, and increases the biodiversity of the fossil genus. The high diversity of species in Protokateretes provides new insights into sexual dimorphism of Kateretidae in mid-Cretaceous, enriching the knowledge of potential courtship strategy of kateretid beetles in the late Mesozoic.

甲虫科(Kateretidae)是一个甲虫家族,据报道有 15 个化石物种,其中 4 个物种具有特化的触角。特化的触角是甲虫科中一种罕见的性二态形式。在此,我们报告了两个具有特殊触角的新种,雄性个体为赵,黄和蔡,2023 年发现的 Protokateretes,雌性个体为一个未命名的 Protokateretes 种。新物种的发现为 Protokateretes 的诊断提供了补充,并增加了该化石属的生物多样性。Protokateretes物种的高度多样性为研究白垩纪中期甲虫的性二型提供了新的视角,丰富了对中生代晚期甲虫潜在求偶策略的认识。
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引用次数: 0
The first stegosaurian dinosaur from Gansu Province, China 中国甘肃省发现的第一只盗龙类恐龙
IF 2.1 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2024-01-26 DOI: 10.1016/j.cretres.2024.105852
Ning Li , Daqing Li , Guangzhao Peng , Hailu You

Stegosaurs are a minor but iconic clade of ornithischian dinosaurs, and along with their sister taxon ankylosaurs form the clade Eurypoda, the major radiation of Thyreophora (armoured dinosaurs). We here report some stegosaurian materials from the Lower Cretaceous Hekou Group of the Zhongpu area, Lanzhou-Minhe Basin, Gansu Province, China. Most of the morphology of the specimen is similar to Stegosaurus stenops and Stegosaurus homheni. However, its dorsal vertebrae have a higher neural arch and smaller neural canal than Stegosaurus stenops. The neural arches of the dorsal vertebrae of Stegosaurus homheni are deeply excavated dorsal to the neural canal in anterior view, which is not present in GSAU 201201. Because the material is fragmentary, we consider the new specimen as Stegosaurus sp. In phylogenetic analysis, it is also recovered as the sister taxon of Stegosaurus stenops. This is the first stegosaurian dinosaur from Gansu Province, which extends the geographical range of Stegosauria and enriches the Cretaceous stegosaurian record. The Ankylosaur Taohelong jinchengensis is also from the same area and same layer as this stegosaur, which is new evidence that they lived in the same ecosystems alongside each other.

剑龙是鸟臀目恐龙中一个次要但具有标志性的支系,与其姊妹类群踝龙一起组成了剑龙支系(Eurypoda),是甲龙(Thyreophora)的主要分支。我们在此报告了来自中国甘肃省兰州-民和盆地中铺地区下白垩统河口组的一些剑龙类材料。该标本的大部分形态与剑龙(Stegosaurus stenops)和家龙(Stegosaurus homheni)相似。然而,与剑龙相比,它的背椎具有更高的神经弓和更小的神经管。在前视图中,同氏剑龙背椎的神经弓在神经管的背面有很深的凹陷,而 GSAU 201201 没有这种凹陷。在系统发育分析中,它也被认为是 Stenops 剑龙的姊妹类群。这是甘肃省首次发现的剑龙类恐龙,扩大了剑龙类的地理分布范围,丰富了白垩纪剑龙类的记录。踝龙 "陶合龙 "和 "金城龙 "产于同一地区、同一地层,是它们生活在同一生态系统中的新证据。
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引用次数: 0
‘Changing of the guard’ amongst holasteroid echinoids in the upper Maastrichtian of the south-east Netherlands: Exit Echinocorys, enter Hemipneustes 荷兰东南部马斯特里赫特上统类回声体的 "换防":退出棘尾回声体,进入半棘尾回声体
IF 2.1 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2024-01-26 DOI: 10.1016/j.cretres.2024.105850
John W.M. Jagt , Mart J.M. Deckers , Elena A. Jagt-Yazykova

The search, over several decades, for overlap (if any) in the stratigraphical ranges of two common holasteroid sea urchin genera in the type area of the Maastrichtian Stage (south-east Netherlands, north-east Belgium), has come to an end. Here, the first examples of representatives of Echinocorys and Hemipneustes are recorded in situ from the middle portion (between flint levels 10 and 12/12a) of the Lanaye Member (Gulpen Formation) at the former ENCI-HeidelbergCement Group quarry (Maastricht, the Netherlands). At this level, the typically Tethyan genus Hemipneustes is accompanied by other echinoid ‘newcomers’ such as Nucleopygus scrobiculatus, N. coravium, Oolopygus pyriformis, Catopygus fenestratus, Bolbaster prunella, B. koninckanus and Leymeriaster eluvialis. Most of these genera can be linked to shallower settings, and two at least (Nucleopygus, Catopygus) connected with shallower, coarser-grained and more permeable facies. The ‘last’ Echinocorys and the ‘first’ Hemipneustes in the Lanaye Member are separated by a single flint/chalk couplet, indicative of the c. 21-kyr precession cycle. At flint level 10 of the Lanaye Member, a negative carbon isotope excursion, suggesting a regressive phase, has recently been determined and tentatively equated with M3-(b) in the Stevns-1 borehole (Denmark), situated within calcareous nannoplankton zone UC20a and close to the last appearance datum (LAD) of the dinoflagellate marker species Triblastula utinensis. The exchange in echinoid faunas at this level may thus have been controlled by bathymetry (and concomitant differences in temperature), as well as by increased nutrient levels on the seafloor.

几十年来,在马斯特里赫特期类型区(荷兰东南部、比利时东北部)寻找两个常见的海胆属地层范围重叠(如果有的话)的工作已经结束。在此,我们在前 ENCI-海德堡水泥集团采石场(荷兰马斯特里赫特)Lanaye 组(Gulpen Formation)的中间部分(燧石层 10 和 12/12a 之间)记录到了 Echinocorys 和 Hemipneustes 的第一个代表实例。在这一层中,典型的四叠纪属 Hemipneustes 与其他回声虫 "新成员 "如 Nucleopygus scrobiculatus、N. coravium、Oolopygus pyriformis、Catopygus fenestratus、Bolbaster prunella、B. koninckanus 和 Leymeriaster eluvialis 相伴而生。这些属中的大多数都与较浅的地层有关,至少有两个属(Nucleopygus 和 Catopygus)与较浅、颗粒较粗和渗透性较强的地层有关。Lanaye岩层中 "最后 "的Echinocorys和 "最初 "的Hemipneustes被一个燧石/白垩岩对联隔开,表明了约21世纪的前向周期。在拉纳耶成岩的燧石层 10,最近确定了一个负的碳同位素偏移,表明了一个回归阶段,并暂时与 Stevns-1 号钻孔(丹麦)中的 M3-(b)相等,该钻孔位于钙质浮游动物区 UC20a 内,靠近甲藻标记物种 Triblastula utinensis 的最后出现基准(LAD)。因此,这一层的回声动物群交换可能受水深(以及随之而来的温度差异)和海底营养水平增加的控制。
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Cretaceous Research
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