Journal of Anthropological Sciences最新文献

El Castillo cave is a well-known site because of its Paleolithic archaeology and parietal rock art. This paper is focused on the human remains found by V. Cabrera in the Mousterian Unit XX assigned to MIS 4 and early MIS 3. The fossils consist of one upper left second premolar (ULP4), one incomplete proximal hand phalanx, and one partial femoral head. The tooth and the phalanx were assigned to adults, whereas the femoral head belonged to an immature individual due to the absence of fusion traces to the metaphyseal surface. The external morphology and metrical characterization of the Castillo-1466 (ULP4) tooth crown was quantified and compared to the variability of other Neanderthal dental remains and a sample of modern human populations. We also quantified its 3D enamel thickness distribution, its roots morphology, as well as the presence of chipping, and their possible relation to masticatory or paramasticatory activities. Castillo-1466 shows crown dimensions compatible with middle-sized Neanderthal teeth, but with a remarkably thicker enamel than other Neanderthal premolars, such as Marillac 13. The femoral head and the hand phalanx fragment are compared to published values for Neanderthals, although both partial fossils lack diagnostic features precluding any clear taxonomic diagnostic. Therefore, their attribution to Neanderthals is assumed based on the dating of the layers in which they were discovered. El Castillo cave Mousterian fossils represent another contribution to the knowledge of the Middle Paleolithic populations of Northern Spain, where different sites along the Cantabrian mountains yielded several human remains assigned to MIS 4 and early MIS 3.

North Africa is a key area for understanding cultural processes that led to the Acheulean pan - African emergence and expressions and the related hominin population dynamics. Unfortunately, little is known about the early Acheulean in this vast area of the African continent due to the scarceness of archaeological sites in stratigraphic context with reliable chronometric data, human remains, and technological analyses of the lithic industries. Here, we present the first comprehensive techno-economic analysis of the early Acheulean assemblage from Thomas Quarry I - Unit L1 (ThI-L1, Casablanca, Morocco), which is the earliest Acheulean site of North Africa, unambiguously dated to 1.3 Ma. Fieldwork has unearthed faunal remains and a lithic collection containing over 3800 artefacts, which represents one of the largest series for the early African Acheulean. The assemblage is mainly composed of quartzites and to a lesser extent of silicites, both abundantly available near the site. Previously published results of the silicite study revealed two different productions for the extraction of small flakes and of bladelet-like flakes. In this work, we analyse the techno-economic systems of the quartzite assemblage. Two distinct quartzite productions co-occur, one devoted to the manufacture of Large Cutting Tools (LCTs), the other focused on the extraction of small-medium sized flakes. LCTs were usually produced from large cobbles, less often from large flakes detached mainly using the entame core method. The main technical objective was to obtain large pointed tools and, more rarely, large tools with a transversal cutting edge. Results support the existence of a strong synergy between conceptual and operational schemes regulated by the ability to anticipate the final tool morphology and to apply standardized shaping procedures to manufacture recurrent morphotypes. Quartzite small-medium sized flakes were produced by a diversity of flaking methods adapted to the cobble blank morphologies and were not retouched. The results allow to assess that the earliest technical expression of the Acheulean in North Africa is characterised by a high diversification of the stone knapping outcomes, the complexity of the mental templates, and the flexible structure of the operational schemes.

Cortical bone and dentine share similarities in their embryological origin, development, and genetic background. Few analyses have combined the study of cortical bone and dentine to quantify their covariation relative to endogenous and exogenous factors. However, knowing how these tissues relate in individuals is of great importance to decipher the factors acting on their evolution, and ultimately to understand the mechanisms responsible for the different patterns of tissue proportions shown in hominins. The aims of this study are to examine age-, sex-, and ancestry-related variation in cortical bone and dentine volumes, and to preliminary assess the possible covariation between these tissues in modern humans and in five composite Neandertals. The modern analytical sample includes 12 immature individuals from France and 49 adults from France and South Africa. Three-dimensional tissue proportions were assessed from microtomographic records of radii and permanent maxillary canines. Results suggest ontogenic differences and a strong sexual dimorphism in cortical bone and dentine developments. The developmental pattern of dentine also seems to vary according to individual's ancestry. We measure a stronger covariation signal between cortical bone and dentine volumes than with any other dental tissue. A more complex covariation pattern is shown when splitting the modern sample by age, sex, and ancestry, as no signal is found in some subsamples while others show a covariation between cortical bone and either crown or radicular dentine. Finally, no difference in cortical bone volume is noticed between the modern young adults and the five young adult composite Neandertals from Marine Isotopic Stages (MIS) 5 and 3. Greater dentine Cortical bone and dentine (co)variation volumes are measured in the MIS 5 chimeric Neandertals whereas a strong interpopulation variation in dentine thickness is noticed in the MIS 3 chimeric Neandertals. Further research on the cortical bonedentine covariation will increase understanding of the impact of endogenous and exogenous factors on the development of the mineralized tissues.

Here we present the results of a microcontextual analysis of purported combustion features recovered from Middle and Upper Paleolithic occupations at the cave site of Fumane, Italy. Our analyses, which integrate micromorphology with organic petrology, show that only a few of the features represent primary, intact hearths; some of them show evidence for various phases of anthropogenic reworking, either through trampling or sweeping and dumping. Several of the features are multi-layered and reflect a complex formation history of various activities related to combustion and site maintenance. Many appear to be the remnants of occupation horizons only partially preserved and peripherally related to combustion. Within several of the intact hearths from the Mousterian, we were able to identify variable fuel sources in different features, implying a degree of flexibility in the fuel-selection strategies of the Neanderthal occupants of Fumane. In this study we design a classification system of the anthropogenic features and also conduct a spatial analysis, through which we can infer diachronic patterns in the frequency and intensity of site occupation and the spatial distribution of activities. We note a decrease in frequency of combustion features throughout the Mousterian which continues into the Uluzzian. The features associated with the Protoaurignacian occupation, in contrast with those from the Mousterian, are multi-layered and well-defined. We argue that these trends, which correspond with other trends in artefact frequency, imply changes in the settlement dynamics of the site during the transition from the last Neanderthal occupation of the cave to the arrival of modern humans.

Seasonality plays a critical role in determining the yearly dietary variability of many nonhuman primates living in tropical and subtropical environments. Much previous research has emphasised the seasonal importance of both preferred resources-eaten whenever available-and fallback foods-eaten during periods of scarcity to compensate for an insufficient availability of preferred resources. However, previous discussions of this dichotomy have often overlooked why different populations of the same taxon may exhibit a different level of engagement with identical resources, especially those that require additional technological investment by virtue of being embedded. Similarly, not enough attention has been given to diachronic trends in the incorporation of novel resources to seasonal consumption patterns among non-human primates. In this paper, we present a systematic framework for understanding the spatio-temporal relationships between preferred and fallback resources, explicitly through the lens of landscape knowledge and seasonal fluctuations in quality and availability among chimpanzee communities. We argue it is the interplay between resource quality and the available knowledge pertaining to its exploitation that will determine the categorisation of a resource. In this regard, the accumulation of further information through encounter, experimentation, and behavioural (including technological( innovation enables resources with high nutritional potential to attain preferred status. We end with an exploration of the gradual consolidation of the hominin carnivory niche in the Early Pleistocene of East Africa, to demonstrate the utility of our framework-specifically the interplay between seasonality and the concept of landscape knowledge-for understanding behavioural change in the archaeological record.

This paper presents an updated view on the morphological and functional significance of the human respiratory system in the context of human evolutionary anatomy. While usually the respiratory system is treated either from a craniofacial perspective, mostly in the context of nasal evolution and air-conditioning, or from a postcranial perspective featuring on overall thoracic shape changes, here we pursue a holistic perspective on the form, function, integration, and evolutionary change of the entire organismal system in hominins. We first present a brief review of the most important morphological structures, their function, and its potential integration and interaction with the nasal cavity and thoracic skeleton. This is followed by an overview of the most important improvements in methods for the comparative study in recent humans and fossil hominins. We then overview and list a compendium of hominin fossil material currently available for the study. We propose four functional categories of hominin respiratory system configurations that differ potentially with respect to size, shape, biomechanics and/or bioenergetics. Finally, we discuss these and speculate on possible ways for future research into an anatomical system that, despite its under-investigated status, is central to the understanding of the form and functions of the hominin organism and its paleobiology.

The present-day diversity of southern African populations was shaped by the confluence of three major pre-historic settlement layers associated with distinct linguistic strata: i) an early occupation by foragers speaking languages of the Kx'a and Tuu families; ii) a Late Stone Age migration of pre-Bantu pastoralists from eastern Africa associated with Khoe-Kwadi languages; iii) the Iron Age expansion of Bantu-speaking farmers from West-Central Africa who reached southern Africa from the western and eastern part of the continent. Uniting data and methodologies from linguistics and genetics, we review evidence for the origins, migration routes and internal diversification patterns of all three layers. By examining the impact of admixture and sex-biased forms of interaction, we show that southern Africa can be characterized as a zone of high contact between foraging and food-producing communities, involving both egalitarian interactions and socially stratified relationships. A special focus on modern groups speaking languages of the Khoe-Kwadi family further reveals how contact and admixture led to the generation of new ethnic identities whose diverse subsistence patterns and cultural practices have long puzzled scholars from various disciplines.

In this contribution I will review the development of ideas about a recent African origin for our species over the last 50 years, starting from the time of my PhD in the early 1970s. I will examine the instructive and quite different interpretations placed on the 1979 discovery of a partial Neanderthal skeleton associated with a Châtelperronian industry at the rock shelter of St-Césaire in France, and then focus on the crucial years from 1987-1989, including the so-called 'Human Revolution' conference of 1987, and my 1988 Science paper with Peter Andrews: 'Genetic and Fossil Evidence for the Origin of Modern Humans'. Following the historical review, I will assess the status of five proposed models for the evolution of derived Homo sapiens: Recent African Origin (RAO); RAO and Hybridisation (RAOH); Assimilation (AM); Multiregional Evolution (MRE); and Braided Stream (BS). I conclude that a recent African origin model with hybridization (RAOH) is the best supported from the fossil and genetic evidence.