The role of hominins in the hunting and processing of megafauna is one of the most contentious topics in Pleistocene archaeology. Determining whether the remains of very large mammals (>800 kg) were shaped by human activity or carnivore modification requires robust neotaphonomic frameworks. Systematic patterns of carcass consumption by carnivores have been established, particularly for long bones. Yet research on megafauna, and especially on their cranial remains, are scarce despite their recurrent presence in archaeological contexts. This study investigates bone surface modification patterns on rhinoceros (Ceratotherium simum) and elephant (Loxodonta africana) cranial elements consumed by free-ranging spotted hyenas (Crocuta crocuta) in the Timbavati Private Nature Reserve, South Africa. The sample includes 20 cranial specimens (MNE = 20; 12 rhinoceros, 8 elephant) from individuals of different age groups that died naturally and were subsequently scavenged. Analyses using 40x hand lenses and Dino-Lite digital microscope to document bone surface modifications. Distinct patterns emerged: in rhinoceroses, near-complete destruction of mandibular condyles, coronoid processes, nuchal crest, and maxilla; in elephants, pronounced furrowing on mandibular condyles and the symphyseal region. Integrating these observations with published taphonomic analyses of Eurasian assemblages, a five-stage sequence of cranial exploitation is proposed, paralleling models established for long bone consumption. These results highlight prey species-specific differences in modification intensity linked to bone density, tissue distribution, and feeding strategies. More broadly, they provide comparative criteria for distinguishing hyena from hominin-induced modifications, and the paleoecological significance of megafauna in Pleistocene ecosystems.
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