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Palynology, palynofacies and palaeoenvironment of the Serpukhovian Tazlourt Formation (Tinghir region, southern margin of the Variscan Zone, SE Morocco) 摩洛哥东南部Variscan带南缘Tinghir地区Serpukhovian Tazlourt组孢粉学、孢粉相及古环境
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-11-01 Epub Date: 2025-06-19 DOI: 10.1016/j.revpalbo.2025.105396
Amine Talih , Daniel Ţabără , Hamid Slimani , Ralph Thomas Becker , Imad Tmimne , Salma Aboutofail , Hicham El Asmi , Soukaina Jaydawi
The microfloristic study of limestones from the Tazlourt Formation in the Tinghir region, led to the first discovery of miospores of herbaceous and sub-arborescent lycopsids small fern and sphenopsids belonging to both non-forest mire and forest mire. The palynological assemblage also includes rare marine phytoplankton (i.e. Lophosphaeridium spp.). Among the continental palynomorphs, significant marker taxa, such as Grandispora spinosa, Dictyotriletes bireticulatus, Cirratriradites cf. C. rarus, Schulzospora cf. S. campyloptera and Punctatisporites gretensis enable a confident attribution to the upper Serpukhovian stage (the subtriquetra–ornatus or SO miospore Zone of the Western Europe). This proves the presence of uppermost Mississippian deposits at the southern front of the Moroccan Variscan chain. The kerogen released by the analyzed samples is predominantly of continental origin, mainly composed of opaque phytoclasts with different sizes and shapes, while translucent phytoclasts and miospores are less represented. The analysis of microfacies, as well as the qualitative and quantitative parameters of the various components of the palynofacies, indicates a shallow to moderately deep neritic environment and supports the existence of a connection to the “Variscan Sea” between the Anti-Atlas domain, located on the northwestern margin of the stable Gondwana craton, and the Moroccan Hercynides, situated at the southwestern margin of the Variscides. The miospores are characterized by significant alteration in texture and color, with Spore Color Index (SCI) values ranging from 7 to 9 and advanced silicification. This is likely related to the combination of tectonic events that affected the study area during the Meso- and Post-Variscan phases.
通过对Tinghir地区Tazlourt组灰岩的微区系研究,首次发现了草本和亚乔木石松类、小蕨类和桫椤类的小孢子,它们属于非森林沼泽和森林沼泽。孢粉组合还包括稀有的海洋浮游植物(如Lophosphaeridium spp.)。在大陆的孢子类中,有重要的标志分类群,如大孢子虫、双网三叶虫、Cirratriradites cf. C. rarus、Schulzospora cf. S. campyloptera和Punctatisporites gretensis,可以肯定地归属于Serpukhovian的上阶段(西欧的亚三孢子带或SO小孢子带)。这证明了在摩洛哥瓦里斯坎链的南部前沿存在密西西比最上层的矿床。分析样品释放的干酪根以陆源干酪根为主,主要由不同大小和形状的不透明植物碎屑组成,而半透明植物碎屑和微孢子较少。微相分析及孢粉相各组分的定性和定量参数表明,该区为浅海至中深浅海环境,支持位于稳定的冈瓦纳克拉通西北缘的反阿特拉斯域与位于瓦里西底斯西南缘的摩洛哥海西尼德之间存在与“瓦里西坎海”的联系。孢子颜色指数(Spore color Index, SCI)在7 ~ 9之间,呈高度硅化。这可能与中、后瓦里斯坎期影响研究区的构造事件的组合有关。
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引用次数: 0
Rare species in past pollen records and herbarium specimens: Linnaea borealis L. lived in north-eastern lowlands in Italy during the Neolithic 过去花粉记录和植物标本室标本中的稀有物种:新石器时代生活在意大利东北部低地的北方林草
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-11-01 Epub Date: 2025-06-21 DOI: 10.1016/j.revpalbo.2025.105398
Jessica Zappa , Paola Torri , Giovanni Astuti , Lorenzo Lastrucci , Anna Maria Mercuri , Assunta Florenzano
This paper emphasises the role of detailed pollen identification in past records and the importance of historical herbaria in palaeoecological research on plant distribution over the last millennia. Palynological analyses of a sediment core, covering approximately the last 10,000 years, from the wetland surrounding the UNESCO archaeological site of Palù di Livenza, in the lowlands of Friuli-Venezia Giulia, revealed the unexpected presence of Linnaea borealis during the Mid-Holocene (about 8600–4200 cal yr BP) in north-eastern Italy. This species is a rare small suffruticose plant which grows among mosses in conifer and mixed forests, between 1200 and 2100 m a.s.l. In Italy, due to the restricted climatic requirements, its current distribution is limited to a few high-mountain stands and to four northern regions (Val d'Aosta, Piedmont, Trentino-Alto Adige, Lombardy). Pollen grains of L. borealis found in past samples from the Friuli-Venezia Giulia region raised interest because this species was not recorded so far in this region either at present or in the past. To confirm the identification, a morphological study was carried out on pollen from flowers collected from University Herbaria (Florence and Pisa). Dried pollen extracted from anthers was acetolysed and observed with a Digital Optical Microscope. Polar axis, equatorial axis, exine thickness, equatorial diameters, mesocolpium, and distance among apices of colpi were measured in current and past pollen grains. Morphological analysis confirms the presence of L. borealis in the sedimentary archive studied, adding an important insight into the knowledge of past and present biodiversity of the area.
本文强调了花粉详细鉴定在过去记录中的作用,以及历史植物标本库在近千年植物分布古生态研究中的重要性。在联合国教科文组织考古遗址Palù di Livenza周围的湿地,在Friuli-Venezia Giulia的低地,对沉积物岩心进行了大约10000年的孢粉学分析,揭示了在意大利东北部全新世中期(约8600-4200 cal yr BP), Linnaea borealis的意外存在。本种是一种罕见的小型湿性植物,生长在海拔1200米至2100米之间的针叶林和混交林中的苔藓中。在意大利,由于气候要求有限,其目前的分布仅限于少数高山林分和四个北部地区(Val d'Aosta, Piedmont, Trentino-Alto Adige,伦巴第)。在弗留利-威尼斯-朱利亚地区过去的样品中发现的北方松花粉粒引起了人们的关注,因为该地区目前和过去都没有记录到该物种。为了证实这一鉴定,对从佛罗伦萨和比萨大学植物标本室收集的花粉进行了形态学研究。从花药中提取干花粉进行乙酰化,用数码光学显微镜观察。测定了现在和过去花粉粒的极轴、赤道轴、外壁厚度、赤道直径、中胚轴和胚轴尖间距离。形态学分析证实了L. borealis在研究的沉积档案中的存在,为该地区过去和现在的生物多样性知识增加了重要的见解。
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引用次数: 0
A sedimentary record of fire evolution history and its response to climate change and human activities during the last 7000 years from Lake Qionghai, southeastern Tibetan Plateau 青藏高原东南部琼海近7000年火的沉积记录及其对气候变化和人类活动的响应
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-11-01 Epub Date: 2025-06-13 DOI: 10.1016/j.revpalbo.2025.105394
Haozhen Yin , Xingqi Liu , Huanyu Sun , Xin Mao , Jun Yang , Guoxiang Li
The occurrence of fire is related to climate, vegetation, and human activity. The Qionghai Lake catchment, situated on the southeastern margin of the Tibetan Plateau, exhibits well-developed vegetation and a high fire frequency and is located in Xichang City—a settlement with a long-standing historical legacy. It is therefore one of the most suitable areas to study the fire-climate-vegetation-human connection. In this study, we reconstruct the fire evolution history of Lake Qionghai over the last 7000 years using AMS 14C dating, along with an analysis of charcoal accumulation rate (CHAR) and major pollen taxa. Our results reveal two distinct phases in Holocene fire dynamics: (1) During the period of 7000–620 cal yr BP, fire activity exhibited a sustained decrease, which may have been linked to the reduction in lightning due to the temperature drop during that period. (2) From 620 cal yr BP to the present‌, a marked increase in fire activity has been accompanied by a pronounced decline in woody plants, coinciding with the proliferation of anthropophilic plant—Poaceae. We propose that the increased fire activity in the catchment area has become increasingly influenced by human activities, particularly through population growth, settlement expansion, and the adoption of slash-and-burn agricultural practices that collectively intensified fire regime.
火灾的发生与气候、植被和人类活动有关。琼海流域位于青藏高原东南边缘,植被发达,火灾频发,地处历史悠久的西昌市。因此,这里是研究火-气候-植被-人类关系最适宜的地区之一。本研究利用AMS 14C测年技术重建了琼海湖区近7000年的火灾演化史,并对炭积累速率(CHAR)和主要花粉类群进行了分析。研究结果揭示了全新世火灾动态的两个不同阶段:(1)在7000-620 cal yr BP期间,火灾活动呈现持续减少的趋势,这可能与该时期气温下降导致闪电减少有关。(2)从620 cal yr BP至今,森林火灾活动的显著增加伴随着木本植物的显著减少,与亲人植物——禾科的大量繁殖相一致。我们认为,汇水区火灾活动的增加受到人类活动的影响越来越大,特别是人口增长、定居点扩张和采用刀耕火种的农业做法,这些都加剧了火灾的发生。
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引用次数: 0
The bees tell the story of disappeared landscapes: The case of the apiaries of Ürünlü/Kite (Bursa, Turkey) 蜜蜂讲述了消失的景观的故事:Ürünlü/Kite (Bursa,土耳其)的养蜂场的案例
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-11-01 Epub Date: 2025-06-18 DOI: 10.1016/j.revpalbo.2025.105397
Güven Gümgüm , Aycan Tosunoglu
This study covers the ethnoarchaeological research on the hives abandoned 200 years ago, which evaluates the historical significance of studied apiaries and melissopalynological research, which addresses the change of floral diversity through palynological analysis in comparison with contemporary data. Conducted in the village of Ürünlü (Kite), Turkey, the study explores the effects of beekeeping, honey production, urbanization, and agricultural expansion on the flora of ancient Bithynia in recent centuries.
Extensive construction, new buildings, and landscape changes contributed to the obsolescence of these traditional apiaries. The abandonment of crude brick hive structures marked the shift to modern beekeeping. Melissopalynological analyses reveal that honey bees serve not only as honey producers but also as recorders of floristic change. Their foraging patterns reflect human-induced changes in floristic composition over time.
Traces of this unique apicultural practice persist in Turkey, particularly in a village near Bursa (ancient Prusa), south of the Sea of Marmara and were contributed to its strategic location along the Silk Road. Surrounding villages, including Ürünlü, still retain remnants of historical agricultural and pastoral activities. By examining these abandoned hives by a multidisciplinary approach, this study sheds light on the interplay between traditional beekeeping practices and environmental transformations. It underscores the importance of historical apiaries in understanding both past and present floristic diversity, offering valuable insights into the ecological and cultural history of the region.
本研究涵盖了对200年前遗弃的蜂箱的民族考古研究,评估了所研究的蜂房和孢粉学研究的历史意义,并通过孢粉学分析与当代数据进行了比较,解决了花卉多样性的变化。该研究在土耳其Ürünlü (Kite)村进行,探讨了近几个世纪以来养蜂、蜂蜜生产、城市化和农业扩张对古代比提尼亚植物群的影响。大规模的建设、新建筑和景观变化导致了这些传统养蜂场的过时。原始砖房结构的废弃标志着向现代养蜂的转变。蜂蜜学分析表明,蜜蜂不仅是蜂蜜的生产者,而且是植物区系变化的记录者。它们的觅食模式反映了人类引起的植物区系组成随时间的变化。这种独特的养蜂方式的痕迹在土耳其一直存在,特别是在马尔马拉海以南的布尔萨(古代普鲁萨)附近的一个村庄,并有助于其沿丝绸之路的战略位置。周围的村庄,包括Ürünlü,仍然保留着历史上农业和畜牧活动的遗迹。通过多学科方法检查这些废弃的蜂箱,本研究揭示了传统养蜂实践与环境转变之间的相互作用。它强调了历史上的养蜂场对了解过去和现在的植物多样性的重要性,为了解该地区的生态和文化历史提供了宝贵的见解。
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引用次数: 0
Cupressaceous woods in Jurassic geothermal deposits of the Deseado Massif, Patagonia, Argentina with evidence of nurse-log growth strategy 阿根廷巴塔哥尼亚Deseado地块侏罗纪地热矿床中的柏木与护士原木生长策略的证据
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-11-01 Epub Date: 2025-06-29 DOI: 10.1016/j.revpalbo.2025.105400
Sebastian Alejandro Molano , Giovanni Cristian Nunes , Josefina Bodnar , Ignacio Hernán Escapa , Juan Leandro García Massini , Diego Guido
The fossil wood record of Cupressaceae is extensive across the Mesozoic and Cenozoic, with a wide distribution in both hemispheres. However, the affinities of some fossil woods are uncertain due to incomplete descriptions, preservation issues, and anatomical traits of ambiguous interpretation. This study describes the anatomy and discusses the affinities of well-preserved Jurassic silicified woods. Samples were collected from geothermal deposits at the ‘Claudia’ locality in the Deseado Massif, Santa Cruz Province, Southern Patagonia, Argentina. The material corresponds to pycnoxylic and homoxylic secondary xylem characterized by a distinctive combination of traits, including mixed pitting on tracheids, cupressoid cross-field pits arranged in a cupressoid pattern, and abundant diffuse axial parenchyma. These characteristics are compatible with Protocupressinoxylon and suggest affinities with Cupressaceae. Additionally, chert blocks preserving fragments of these woods contain vegetative organs (e.g., leaves, roots) with affinities with this family, supporting the suggested affiliation. The studied woods are assigned to the open nomenclature Protocupressinoxylon-type wood, following the IAPT Fossil Committee's recommendations concerning the nomenclature problems of this name, while acknowledging the extensive historical and current use of it. Nurse logs are also recorded based on the presence of several stems and rootlets anchored to some of the studied woods, which expands the fossil record of this growth strategy.
柏科木材化石记录广泛分布于中生代和新生代,在两个半球分布广泛。然而,由于描述不完整、保存问题和解剖特征解释不明确,一些化石木材的亲缘关系尚不确定。本研究描述了保存完好的侏罗纪硅化木的解剖结构,并讨论了它们的亲缘关系。样品采集自阿根廷南巴塔哥尼亚圣克鲁斯省Deseado地块“Claudia”地区的地热矿床。其次生木质部为强氧木质部和均氧木质部,具有管胞上的混合麻点、以柏木形态排列的柏木交叉坑和丰富的弥漫性轴向薄壁组织等特征。这些特征与原柏树(Protocupressinoxylon)一致,表明其与柏科有亲缘关系。此外,保存这些木材碎片的燧石块含有与这个家族有亲缘关系的营养器官(如叶子、根),支持了所建议的联系。根据IAPT化石委员会关于该名称命名问题的建议,所研究的木材被分配到开放命名的protocupressinoxylon型木材,同时承认它在历史和当前的广泛使用。护士原木也被记录下来,这是基于一些被研究的树木上固定的一些茎和根的存在,这扩展了这种生长策略的化石记录。
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引用次数: 0
Resilient gymnosperms: reassessing floral dynamics at the permian–triassic extinction in Meishan 弹性裸子植物:重新评估梅山二叠纪-三叠纪物种灭绝的花卉动态
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-10-01 Epub Date: 2025-05-13 DOI: 10.1016/j.revpalbo.2025.105373
Elke Schneebeli-Hermann , Francesca Galasso
The impact of the Permian–Triassic mass extinction on terrestrial plant diversity and ecosystems is still controversially discussed. While for numerous plant fossil records, particularly those from terrestrial sites, adequate dating is unfeasible, for plant microfossil records, deposited in marine environments, independent biostratigraphic dating is usually possible. Since the ratification of the Global Stratotype Section and Point (GSSP) for the Permian–Triassic boundary in Meishan, a plethora of not only palynological data but also precise geochronological ages, high-resolution geochemical and biostratigraphic datasets are available. Here, we aim to revise the palynological data published by Ouyang & Utting in 1990 in the current geochronological framework. Their three initially described assemblage zones yield a wealth of information with respect to depositional environment and vegetation composition- from an off-shore Permian setting to a more proximal setting in the Triassic. To date, the comparison of the palynological data with the carbon isotope record, current lithostratigraphy, and U/Pb ages indicates that gymnosperm-dominated vegetation thrived during and after the Permian–Triassic extinction event in the continental area near Meishan.
二叠纪-三叠纪大灭绝对陆生植物多样性和生态系统的影响仍存在争议。虽然对于许多植物化石记录,特别是来自陆地遗址的植物化石记录,不可能进行适当的定年,但对于沉积在海洋环境中的植物微化石记录,通常可以进行独立的生物地层定年。自从眉山二叠纪-三叠纪界线的全球层型剖面和点(GSSP)被批准以来,不仅有大量的孢粉学数据,而且有精确的年代学年龄,高分辨率的地球化学和生物地层数据集。在此,我们拟对欧阳等人发表的孢粉学资料进行修订。在目前的地质年代学框架下是在1990年。他们最初描述的三个组合带提供了关于沉积环境和植被组成的丰富信息——从沿海的二叠纪环境到更近的三叠纪环境。迄今为止,将孢粉学资料与碳同位素记录、当前岩石地层和U/Pb年龄进行比较,表明梅山附近大陆地区在二叠纪-三叠纪灭绝事件期间和之后,裸子植物为主的植被繁盛。
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引用次数: 0
Qualitative LM and SEM study of the Messel palynoflora: Part II. Fabales to Caryophyllales 禾草的定性LM和SEM研究(二)。寓言属到石蕊属
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-10-01 Epub Date: 2025-04-22 DOI: 10.1016/j.revpalbo.2025.105349
Johannes M. Bouchal , Christian Geier , Silvia Ulrich , Volker Wilde , Olaf K. Lenz , Reinhard Zetter , Friðgeir Grímsson
This second part of the re-investigation of the Eocene Messel palynoflora, using combined light- and scanning electron microscopy (single-grain method), comprises pollen of the Fabales to Caryophyllales. During our study, we discovered one Fabales pollen type, eight Rosales, 20 Fagales, ten Malpighiales, two Myrtales, twelve Sapindales, eleven Malvales, three Santalales, and a single Caryophyllales pollen type. New taxa in the Messel palynoflora includes Erythrina (Fabaceae) and Moraceae. Continuing to compare the palynoflora and meso/macroflora, we notice that additional six families only occur in the microflora, and six other families are represented only by leaf/fruit/seed fossils. Pursuing our assessment of the pros and cons of the combined versus classical quantitative light microscopy studies, we show that the combined method continues to provide a higher diversity of pollen taxa from the plant groups encountered. We report a similar pollen diversity for the orders Fabales to Charyophyllales from a single sample, as previously reported from conventional LM analyses of multi-sampled drill cores. Our combined LM/SEM approach revealed several new floral elements and enabled us to revise several previous botanical affinities based on additional morphological characters observed with SEM.
第二部分是对始新世麦塞尔孢粉植物的再调查,采用光和扫描电子显微镜(单粒法)联合研究,包括fabale到caryophyllale的花粉。在研究过程中,我们发现了1个Fabales花粉类型、8个Rosales花粉类型、20个Fagales花粉类型、10个Malpighiales花粉类型、2个Myrtales花粉类型、12个Sapindales花粉类型、11个malvalales花粉类型、3个Santalales花粉类型和1个Caryophyllales花粉类型。禾草植物的新分类群包括赤藓属(豆科)和桑科。继续比较孢粉区系和中/大区系,我们注意到另外6个科只存在于微生物区系中,另外6个科只存在于叶/果/种子化石中。我们对联合定量光学显微镜研究与经典定量光学显微镜研究的利弊进行了评估,结果表明,联合方法继续从所遇到的植物群中提供更高多样性的花粉分类群。我们报告了一个单一样本中Fabales目与Charyophyllales目的花粉多样性,正如之前报道的对多个采样岩心的常规LM分析一样。我们的LM/SEM结合方法揭示了一些新的花元素,并使我们能够根据SEM观察到的其他形态特征修改先前的一些植物亲和性。
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引用次数: 0
Almost forgotten fossil wood points to the existence of an overlooked group of Mesozoic Gondwanan gymnosperms 几乎被遗忘的木材化石表明存在一个被忽视的中生代冈瓦纳裸子植物群
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-10-01 Epub Date: 2025-05-22 DOI: 10.1016/j.revpalbo.2025.105383
Marc Philippe , Mike Pole , Pierre Maurizot , Lilian Alizert , Gendry Damien
The wood Araucarioxylon australe Crié was published in 1889. Despite the many nomenclatural and taxonomic issues involved, it has never been revised. We found type material in Rennes University (France) palaeontological collection. Syntypes are described and illustrated, and a lectotype is designated. A new species of Planoxylon Stopes, P. toitoii n.sp. is recognised. A similar wood, Planoxylon hectorii Stopes is also revised. Taxonomical relationship between Mixoxylon Chernomorets et Sakala and Planoxylon Stopes are discussed and a key to Planoxylon species proposed. This renewed taxonomical framework allows a clearer palaeobiogeographical image to emerge from Gondwana high-palaeolatitude Mesozoic fossil wood-record.
木材Araucarioxylon australe cri出版于1889年。尽管涉及许多命名和分类学问题,但它从未被修订。我们在法国雷恩大学的古生物收藏中发现了模式材料。对模式进行了描述和说明,并指定了一个选型。标题平梭梭(Planoxylon Stopes)一新种,P. toitoii n.sp。是认可的。一种类似的木材,Planoxylon hectorii Stopes也经过修订。讨论了扶桑(Mixoxylon Chernomorets et Sakala)与扶桑(Planoxylon Stopes)的分类学关系,并提出了扶桑的种键。这种新的分类框架使得冈瓦纳高古纬度中生代木材化石记录的古生物地理图像更加清晰。
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引用次数: 0
An illustrated protocol for extracting palynomorphs from Early Pleistocene pollen-poor sediments using LST Fastfloat 利用LST Fastfloat从早更新世花粉含量较低的沉积物中提取孢异形的说明方案
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-10-01 Epub Date: 2025-05-15 DOI: 10.1016/j.revpalbo.2025.105372
Valérie Andrieu , Belinda Gambin
This method for extracting palynomorphs is specifically dedicated for Early Pleistocene pollen-poor carbonate or siliceous sediments in the Mediterranean, which are characterised by prolonged summer droughts. It utilises LST Fastfloat, a non-toxic, low viscosity sodium polytungstate heavy liquid. Due to its physical qualities, particularly its low viscosity, LST Fastfloat enables more efficient pollen concentration compared to other methods, including those that rely on alternative dense liquids.
This approach was applied to fluviatile and lacustrine deposits in both the western (Marseille) and eastern (Acıgöl, Kocabaş) Mediterranean. Unlike standard method, LST Fastfloat produced highly concentrated pollen residues and clear pollen slides, reducing analysis time, which is an important advantage in a labour-intensive discipline such as palynology. The primary drawback of this method is its high cost, although it is partially recyclable.
Our article is designed to be didactic and richly illustrated, featuring figures and photographs to facilitate its application, particularly for beginners. The positive results presented here may encourage researchers to sample sediments previously considered pollen-poor or devoid of palynomorphs, especially from understudied regions of major biogeographical or archaeological interest, such as the arid regions of the tropics and the Mediterranean.
这种提取地貌的方法专门用于地中海早更新世缺乏花粉的碳酸盐或硅质沉积物,其特征是夏季长期干旱。它采用LST Fastfloat,一种无毒,低粘度的多钨酸钠重质液体。由于其物理特性,特别是其低粘度,与其他方法(包括那些依赖于替代致密液体的方法)相比,LST Fastfloat可以更有效地浓缩花粉。这种方法应用于地中海西部(马赛)和东部(Acıgöl,科卡巴伊)的流质和湖泊沉积物。与标准方法不同,LST Fastfloat产生高度浓缩的花粉残留物和清晰的花粉载玻片,减少了分析时间,这在孢粉学等劳动密集型学科中是一个重要的优势。这种方法的主要缺点是成本高,尽管它是部分可回收的。我们的文章被设计成教学和丰富的插图,以数字和照片,以方便其应用,特别是对初学者。这里提出的积极结果可能会鼓励研究人员对以前被认为缺乏花粉或没有花粉形态的沉积物进行取样,特别是来自主要生物地理或考古兴趣的研究不足的地区,如热带和地中海的干旱地区。
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引用次数: 0
Tenellisporites capillaris sp. nov., a new dispersed lycopsid megaspore from the Middle–Upper Triassic Badong Formation, Hunan Province, China 湖南中上三叠统巴东组新分散石松类大孢子Tenellisporites capillaris sp. nov.
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-10-01 Epub Date: 2025-05-21 DOI: 10.1016/j.revpalbo.2025.105384
Hong-Xiao Zhan , Qun Sui , Han Wu , Weiyu Lu , Jianbo Chen , Stephen McLoughlin , Zhuo Feng
Dispersed megaspore fossils provide crucial insights into the diversity of heterosporous lycopsids through geological time. However, detailed systematic studies on Middle–Late Triassic dispersed lycopsid megaspores of the South China Block are lacking. Here, we report a unique dispersed lycopsid megaspore species, Tenellisporites capillaris sp. nov., from the Middle–Upper Triassic Badong Formation in Hunan Province, China. This trilete megaspore species is characterised by narrow, elevated, membraneous laesurae with ragged finger-like projections; a flattened equatorial zona; and single or rarely bifurcate spines with sharp tips on both the proximal and distal surfaces. These spines are connected basally to form a fine reticulum and gradually increase in thickness and length towards the equator. Ultrastructural analysis reveals that the megaspore wall consists of two layers: 1) a very thin and dense foot layer, 2) a thick, dense outer layer with parallel sporopollenin units. The outermost sporopollenin units transition to vertically arranged units where forming the spinous ornamentation. The morphological and ultrastructural characteristics indicate that the new megaspore species belongs to Isoetales. Our finding represents the first fossil record of Tenellisporites in Asia, enhancing our understanding of the palaeogeographic distribution and diversity of isoetalean lycopsids during the Middle–Late Triassic.
分散的大孢子化石为了解不同地质时期异孢子石松类的多样性提供了重要的见解。然而,对华南地块中晚三叠世分散石松大孢子的详细系统研究尚缺乏。本文报道了湖南中上三叠统巴东组的一种独特的分散石松类大孢子——Tenellisporites capillaris sp. nov.。这三节大孢子种的特征是狭窄,升高,膜质laesurae具粗糙的指状突起;平坦的赤道带;单根或很少分叉的刺在近端和远端表面都有锋利的尖端。这些刺基部相连,形成一个细小的网状结构,并逐渐向赤道方向增加厚度和长度。超微结构分析表明,大孢子壁由两层组成:1)薄而致密的足部层,2)厚而致密的外层,平行的孢子粉粒单元。最外层的孢子孢粉单元过渡到垂直排列的单元,在那里形成刺状纹饰。形态和超微结构特征表明该大孢子新种属于等生门。该发现是亚洲第一个Tenellisporites化石记录,增强了我们对中晚三叠世等纯石松类古地理分布和多样性的认识。
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Review of Palaeobotany and Palynology
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