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New data on medullosalean foliage Odontopteris schlotheimii from the uppermost Carboniferous–lowermost Permian of central and eastern Europe 中欧和东欧石炭纪上端-二叠纪下端的髓状叶Odontopteris schlotheimii的新资料
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-09-17 DOI: 10.1016/j.revpalbo.2025.105450
Zbyněk Šimůnek , Christopher J. Cleal
The stomatal structure of the medullosalean frond species Odontopteris schlotheimii is documented for the first time. This supports the view that it is taxonomically distinct from the morphologically similar but rather older frond-species Odontopteris cantabrica. Evidence is also presented that supports the view that the types of Neuropteris pseudoblissii are conspecific with O. schlotheimii.
本文首次报道了齿形齿形植物(Odontopteris schlotheimii)的气孔结构。这支持了它在分类上不同于形态相似但更古老的叶类Odontopteris cantabrica的观点。也提出了证据,支持的观点,即类型的伪布氏神经翅虫与O. schlotheimii同源。
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引用次数: 0
Effect of steep climate gradient on palynological assemblages on the Arabian part of the Tethys shore in two Carboniferous-Permian ice age time slices 陡峭的气候梯度对两个石炭纪-二叠纪冰期片中特提斯海岸阿拉伯部分孢粉组合的影响
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-09-15 DOI: 10.1016/j.revpalbo.2025.105448
Michael H. Stephenson , Shuzhong Shen , Junxuan Fan , Linshu Hu , Jin Qi
Palaeoclimate modelling of two time slices, Serpukhovian to Bashkirian (Carboniferous), and Gzhelian to early Sakmarian (latest Carboniferous to Early Permian), indicates marked changes in palynological assemblages from 30°S and 50°S across the Gondwana and Tethys margin. Both time slices indicate a distinct cold climate assemblage dominated by cheilocardioid spores, while a warmer climate with mean annual temperature above 19 °C appears to encourage the growth of plants that produced Euramerican coal belt spores such as Vestispora fenestrata and Reticulatisporites polygonalis. It is possible that the parent plants of cingulicamerate spores such as Lundbladispora spp., though tolerant of cold conditions, were not tolerant of the coldest conditions (for example below 0 °C).
Serpukhovian—Bashkirian(石炭世)和Gzhelian—早Sakmarian(石炭世晚期—早二叠世早期)两个时间片的古气候模拟表明,在30°S和50°S范围内,Gondwana和Tethys边缘的孢粉组合发生了显著变化。这两个时间切片都显示了一个明显的寒冷气候组合,主要是cheilocardioid孢子,而年平均温度高于19°C的温暖气候似乎促进了产生欧美煤带孢子的植物的生长,如Vestispora fenestrata和Reticulatisporites polygonalis。有可能钩孢孢子的亲本植物,如Lundbladispora spp.,虽然能耐受寒冷条件,但不能耐受最冷的条件(例如低于0°C)。
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引用次数: 0
First record of Cyathocarpus (Marattiales) in Gondwana: A new species from the Permian of Patagonia, Argentina 冈瓦纳地区Cyathocarpus (Marattiales)的首次记录:阿根廷巴塔哥尼亚二叠纪的一新种
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-09-11 DOI: 10.1016/j.revpalbo.2025.105447
Gonzalo Rodriguez Rizk, Bárbara Cariglino
Marattialean ferns were key components of Paleozoic wetland ecosystems, yet their biogeographic history in Gondwana remains poorly resolved. While Cyathocarpus is well documented from Euramerica and Cathaysia, no species had been recognized in Gondwana—until now. We describe Cyathocarpus polinensis sp. nov. from the Guadalupian La Golondrina Formation of Patagonia, Argentina, representing the first Gondwanan record of the genus. The species exhibits a distinctive arrangement of synangial rows that do not contact the midvein or pinnule margins and are consistently absent from the apical and basal regions of the pinnules. The midvein is conspicuous, straight, and divides fanwise near the apex; lateral veins bifurcate once, close to the midvein. This discovery extends the paleogeographic range of Cyathocarpus and supports a dispersal scenario from Euramerica into southern Gondwana during the Cisuralian. These findings call for a critical reassessment of comparable Gondwanan taxa and underscore the role of southwestern Gondwana as a refugium and center of marattialean diversification during postglacial climatic recovery.
摘要蕨类植物是古生代湿地生态系统的重要组成部分,但其在冈瓦纳的生物地理历史仍未得到很好的解决。虽然Cyathocarpus在欧洲和华夏都有很好的记录,但直到现在,冈瓦纳还没有发现任何物种。我们描述了来自阿根廷巴塔哥尼亚Guadalupian La Golondrina组的Cyathocarpus polinensis sp. 11 .,代表了该属的第一个冈瓦纳记录。该种表现出一种独特的排列,不接触中脉或小穗边缘的同质行,并且始终不存在于小穗的顶端和基部区域。中脉明显,直,并在先端附近扇形分开;侧脉分叉一次,靠近中脉。这一发现扩大了Cyathocarpus的古地理范围,并支持了一种从欧洲到冈瓦纳南部的分布情况。这些发现要求对冈瓦纳的类群进行批判性的重新评估,并强调冈瓦纳西南部在冰川后气候恢复期间作为避难所和物种多样化中心的作用。
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引用次数: 0
Paleobotanical insights from the Santa Clara Arriba Formation (Anisian, Middle Triassic, Cuyana Basin, West Gondwana) 圣克拉拉Arriba组古植物学研究(西冈瓦纳奎亚纳盆地中三叠世安尼西亚期)
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-09-11 DOI: 10.1016/j.revpalbo.2025.105449
Tomas Ezequiel Pedernera , Cecilia Andrea Benavente
The Santa Clara Arriba Formation is an Anisian unit representing a lacustrine deltaic depositional setting that forms part of the El Peñasco Group Santa Clara sub-basin (Cuyana Basin, Mendoza, Argentina). The objective of this contribution is to present new plant fossil findings and a preliminary taphonomic evaluation of the Santa Clara Arriba Formation. It also includes a comparison and analysis of the paleobotanical record of the Santa Clara Arriba Formation with other units of the El Peñasco Group and Anisian units in Argentina. The plant fossil assemblage of the Santa Clara Arriba Formation comprises Equisetales (Equisetales indet.), Osmundales (Cladophlebis spp.), Marattiales (Danaeopsis sp.), Umkomasiales (Fanerotheca cf. papilioformis), Peltaspermales (Peltaspermum spp.), Coniferales (Pagiophyllum sp.), and gymnosperms of uncertain affinity (undeterminated leaves, ?Cordaicarpus sp., ovules/seeds, and strobili). Remains were recorded in delta-plain facies, and their preservation is attributed to rapid sedimentation and a persistent high-water table. The plant paleocommunities are interpreted as forests dominated by gymnosperms, and an understory comprising Osmundales, and Marattiales, while Equisetales formed riparian communities along water margins. The taxonomic composition of the assemblage is consistent with other Triassic floras from the region. The Santa Clara Arriba Formation provides a valuable opportunity to elucidate the relationship between Anisian vegetation and climate in Gondwana, owing to its paleogeographic setting, well-preserved fossil record, and robust geochronological framework.
圣克拉拉Arriba组是一个代表湖泊三角洲沉积环境的阿尼西亚单元,是El Peñasco群圣克拉拉子盆地(阿根廷门多萨圭亚那盆地)的一部分。这篇文章的目的是介绍新的植物化石发现和对圣克拉拉阿里巴组的初步地语学评价。本文还将圣克拉拉Arriba组的古植物学记录与阿根廷El Peñasco组和Anisian组的其他单元进行了比较和分析。圣克拉拉Arriba组的植物化石组合包括Equisetales (Equisetales indet.), Osmundales (Cladophlebis spp.), Marattiales (Danaeopsis sp.), Umkomasiales (Fanerotheca c.papilioformis), Peltaspermales (Peltaspermum spp.), Coniferales (Pagiophyllum sp.)和不确定亲和力的裸子植物(未确定的叶子,?心形花序,胚珠/种子,和球果)。在三角洲-平原相中记录了遗迹,它们的保存归因于快速沉积和持续的高水位。植物古群落被解释为以裸子植物为主的森林,以及一个由裸子植物和马属植物组成的林下植物群落,而马属植物沿着水边形成了河岸群落。该组合的分类组成与该地区其他三叠纪植物区系一致。圣克拉拉Arriba组由于其古地理环境、保存完好的化石记录和健全的年代学框架,为阐明冈瓦纳地区阿尼西亚植被与气候之间的关系提供了宝贵的机会。
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引用次数: 0
The role of agroforestry yards in the diet of Melipona (Melikerria) interrupta Latreille, 1811 (Hymenoptera, Apidae) in a terra-firme environment near the confluence of the Amazon and Tapajós Rivers 在亚马逊河和Tapajós河汇合处的陆地环境中,农林业工场对Melipona (Melikerria) interrupta Latreille, 1811(膜翅目,蜂科)食性的作用
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-09-10 DOI: 10.1016/j.revpalbo.2025.105430
Alyne Daniele Alves Pimentel , Cristiane Krug , Vanessa Holanda Righetti De Abreu , Marcos Gonçalves Ferreira
This study aimed to identify the botanical origin of Melipona interrupta honey produced in agroforestry backyards (AB) by traditional communities near the confluences of the Amazon and Tapajós Rivers. For comparison purposes, stingless beehives were installed in a secondary forest (SF) environment as well. In all, forty-four honey samples were analyzed from four meliponaries in two areas (AB1 and AB2; SF1 and SF2) between July 2021 and June 2022 in Arapixuna. In AB1, 25 pollen types were found in the honey samples and, in AB2, 22 were found. More than 50% of the pollen types found in the honey samples in the ABs were related to plants present only in the SF, indicating that the bees left the ABs and collected pollen in the SF. The main resources collected by M. interrupta for honey composition were Protium heptaphyllum present in 70% of the samples, followed by Triplaris type (68%), Psidium guajava (52%), Bellucia imperialis and Eugenia biflora (38%) and Tapirira guianensis (34%). The survey of botanical species in the ABs indicated that most plant species were fruit trees; however, the bees visited a low percentage of these plants. When analyzing the SF samples, a behavioral pattern of foraging was observed which was similar to those of the ABs. These data indicate that the two environments studied have melliferous characteristics that favor beekeeping. Moreover, ABs also have advantageous characteristics in relation to the SF, such as improved protection of colonies against invaders/theft of honey and the ease of maintenance of the colonies.
本研究旨在确定亚马逊河和Tapajós河汇合处附近传统社区农林业后院(AB)生产的Melipona interrupta蜂蜜的植物来源。为了比较,在次生林(SF)环境中也安装了无刺蜂箱。在2021年7月至2022年6月期间,总共分析了Arapixuna两个地区(AB1和AB2; SF1和SF2)四个蜜源的44份蜂蜜样本。AB1中有25种花粉类型,AB2中有22种花粉类型。在ABs中发现的蜂蜜样品中超过50%的花粉类型与仅存在于SF中的植物相关,表明蜜蜂离开ABs并在SF中收集花粉。对蜂蜜成分采集的主要资源为七叶Protium(70%),其次为Triplaris(68%)、Psidium guajava(52%)、Bellucia imperialis和Eugenia biflora(38%)和Tapirira guianensis(34%)。植物种类调查表明,植物种类以果树为主;然而,蜜蜂造访这些植物的比例很低。结果表明,这两种环境均具有有利于养蜂的蜜蜂特征。此外,ABs还具有相对于SF有利的特性,例如更好地保护蜂群免受入侵者/盗窃蜂蜜和易于维护蜂群。
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引用次数: 0
Terrestrial end-Guadalupian crisis constrained by mid-latitude Permian palynological data from Jimsar, Junggar Basin, China 准噶尔盆地吉木萨尔中纬度二叠纪孢粉资料约束的瓜达鲁普末期陆相危机
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-09-05 DOI: 10.1016/j.revpalbo.2025.105445
Xueqin Wang , Yuewu Sun , Xiao Shi
The end-Permian mass extinction and the end-Guadalupian crisis are the most remarkable events in the Permian terrestrial ecosystem. To better understand the end-Guadalupian event in mid-latitude of Pangea in the Northern Hemisphere, palynological investigations are conducted on the terrestrial Permian deposits in the Dalongkou profile of Jimsar, northern Xinjiang, China. Two palynological assemblages are recognized in stratigraphic ascending order, the Cordaitina subrotataStriatoabieites lipidusProtowelwitschiapollis exolescus assemblage from the uppermost Lucaogou, Hongyanchi and the lower Quanzijie formations, and the Kraeuselisporites spinulosusTuberculatosporites homotubercularisPotonieisporites turpanensis assemblage from the upper Quanzijie and lowermost Wutonggou formations. Based on the palynological data, the uppermost Lucaogou and Hongyanchi formations are assigned to the Roadian stage, and the lower Quanzijie Formation to the Capitanian stage, and the upper Quanzijie and lowermost Wutonggou formations to the Wuchiapingian stage. Thus, a Wordian-age hiatus is confirmed between the lower Quanzijie Formation and the Hongyanchi Formation on the basis of palynological evidence. The palynofloral succession shows a smooth transition between the Guadalupian and Lopingian palynofloras in the Dalongkou profile of Jimsar, in the Junggar Basin. The end-Guadalupian crisis resulted in a 17% loss of pollen species, indicating that the extinction was on the background level.
二叠纪末的大灭绝和瓜达鲁普期末的危机是二叠纪陆地生态系统中最显著的事件。为了更好地了解北半球盘古大陆中纬度的瓜德鲁普末期事件,对新疆北部吉木萨尔大龙口剖面的陆相二叠系沉积物进行了孢粉学研究。从地层升序上可识别出两个孢粉组合,即最上层芦草沟组、红盐池组和最下层泉子沟组的Cordaitina subrotata-Striatoabieites lipidus-Protowelwitschiapollis exolescus组合,以及最上层泉子沟组和最下层梧桐沟组的Kraeuselisporites spinulosus-Tuberculatosporites homotubercularis-Potonieisporites turpanensis组合。根据孢粉资料,将芦草沟组上部和红盐池组划分为路甸期,将泉子街组下部划分为capitian期,将泉子街组上部和梧桐沟组下部划分为五家坪期。因此,根据孢粉学证据,确定了全子街组下段与红盐池组之间存在一个世界间隙。准噶尔盆地吉木萨尔大龙口剖面的孢粉区系在瓜达鲁普系和洛平系之间有一个平稳的过渡。瓜达鲁普末期的危机导致17%的花粉物种损失,表明灭绝是在背景水平上。
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引用次数: 0
An 11,000-year vegetation and climate record from a subalpine peatland in the Luoxiao Mountains, subtropical Southeast China 中国东南部亚热带罗霄山亚高山泥炭地11000年植被和气候记录
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-09-01 DOI: 10.1016/j.revpalbo.2025.105442
Xin Zhong , Caifeng Cen , Cong Chen , Aiyan Han , Yongjie Tang , Kangyou Huang
The evolution of broadleaved forests and their responses to climate changes in subtropical Southeast China remain largely unclear, partly because of the controversies regarding the history of climate (i.e., temperature and monsoonal precipitation) changes during the Holocene. In this study, we integrated pollen and other proxies (including loss on ignition, humification degree, and stable carbon isotope) from a subalpine peatland in the Luoxiao Mountains, to reveal the predominant patterns of vegetation and climate dynamics in mountainous Southeast China. Vegetation at the study site was dominated by Quercus-evergreen and Fagus with relatively flourishing herbs during the early Holocene. A transformation from mixed evergreen–deciduous broadleaved forests to evergreen broadleaved forests occurred between 8200 and 1850 cal. yr BP. The lag of the transformation in response to monsoonal rainfall changes in mountainous regions during the early Holocene can be attributed to thermal limitation. Since 1850 cal. yr BP, broadleaved pollen substantially declined with the expansion of Poaceae and pines, likely reflecting the impact of intensive anthropogenic activities. The expansion of evergreen broadleaved forests from different sites revealed a gradual warming in Southeast China during the Holocene. Additionally, monsoon precipitation in the study area underwent substantial changes, showing a pattern of wetting, sustained humid conditions, subsequent drying, and wetting. Overall, temperature and precipitation during the Holocene were decoupled at the study site, as supported by widely distributed records in mountainous subtropical Southeast China. This may be attributed to the influence of different driving factors on temperature and precipitation.
中国亚热带东南部阔叶林的演变及其对气候变化的响应在很大程度上是不清楚的,部分原因是对全新世气候(即温度和季风降水)变化的历史存在争议。通过对罗霄山亚高山泥炭地的花粉和其他指标(包括着火损失、腐殖化程度和稳定碳同位素)的综合分析,揭示了东南山区植被和气候动态的主要模式。全新世早期植被以常绿栎树和蕨类植物为主,草本植物相对丰富。8200 ~ 1850 cal. yr BP之间发生了常绿落叶混交林向常绿阔叶林的转变。全新世早期山区对季风降水变化响应的滞后可归因于热限制。自1850 cal. yr BP以来,阔叶花粉随着禾科和松科的扩张而大幅下降,可能反映了强烈的人类活动的影响。不同地点常绿阔叶林的扩张揭示了全新世以来中国东南部气候逐渐变暖的趋势。此外,研究区的季风降水经历了实质性的变化,呈现出湿润、持续湿润、随后干燥和湿润的模式。总体而言,全新世温度与降水在研究点呈解耦关系,华南副热带山区分布广泛。这可能与不同驱动因素对温度和降水的影响有关。
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引用次数: 0
Ilex subg. Prinos (Aquifoliaceae) from the upper Miocene of Beihai, low-latitude China: New insights for phylogeny 冬青属植物subg。中国低纬度北海中新世上部水仙花属:系统发育的新认识
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-09-01 DOI: 10.1016/j.revpalbo.2025.105444
Bingshan Niu , Qijia Li , Hui Jia , Jianhua Jin , Cheng Quan
The holly family (Aquifoliaceae) is a monogeneric taxon comprising highly diverse species of Ilex (> 600 species), predominantly distributed in warm, humid regions. However, the phylogeny of Ilex and the evolutionary differentiation of its subgenera, i.e., Ilex subg. Ilex, I. subg. Prinos, and I. subg. Byronia, remain challenging partly due to limited fossil constraints of each subgenus. Previous phylogenetic study was only anchored by the of Ilex subg. Ilex, which is not sufficient to calibrate the differentiation of all subgenera. Here, we report the fossil leaves of Ilex subg. Prinos from the late Miocene Foluo Formation of Beihai, Guangxi, low-latitude China. The fossils are characterized by elliptic leaf in shape with slight serrations on the leaf margin except for the leaf base. The venation is pinnate with secondary veins of 7–8 pairs, while the tertiaries are mixed percurrent. Anatomically, the leaves are hypostomatic with wart-like structures and cyclocytic stomata. Importantly, the anticlinal walls are straight in both upper and lower epidermises. This is the earliest known fossil Ilex subg. Prinos so far, providing a new insight for molecular clock analyses.
冬青科是冬青属的一个单属分类单元,由高度多样化的冬青种组成(约600种),主要分布在温暖潮湿的地区。然而,对冬青的系统发育及其亚属(即冬青亚属)的进化分化进行了研究。Ilex, i, subg。普林诺斯和我。由于每个亚属的化石限制有限,拜伦亚属仍然具有挑战性。以往的系统发育研究仅以冬青亚群为基础。这不足以校准所有亚属的分化。本文报道了冬青属植物的叶化石。中国低纬度广西北海晚中新世洛罗组中的巨象。叶形椭圆形,除叶基外叶缘有轻微锯齿。脉脉为羽状,次生脉7 ~ 8对,三级脉为混流脉。解剖上,叶下气孔,有疣状结构和环状气孔。重要的是,背斜壁在上下表皮上都是直的。这是已知最早的冬青亚属化石。到目前为止,为分子钟分析提供了新的见解。
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引用次数: 0
Soil phytolith assemblages reflect palm community composition in western Amazonia 西部亚马逊地区土壤植物岩组合反映了棕榈群落组成
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-08-28 DOI: 10.1016/j.revpalbo.2025.105443
Crystal N.H. McMichael , Daniel Guerra , Nina H. Witteveen , Britte M. Heijink , Annabel Zwarts , Gabriela Zuquim , Henrik Balslev , Kalle Ruokolainen , Hanna Tuomisto
Humans have domesticated and used palms in tropical forests for millennia. The extent to which past human palm use affects modern tropical systems remains relatively unexplored due to lack of a good fossil record. However, palms produce an array of phytoliths, which are siliceous cell structures that preserve well in the soil even after the organic plant material has decayed. Phytoliths vary in morphology and can be identified to group level and sometimes even to species. We analyzed if palm phytolith assemblages sampled from the soil reflect palm species compositions and abundances in 63 western Amazonian transects. Principal coordinate analysis (PCoA) and Mantel tests showed that palm communities and phytolith assemblages had similar turnover patterns. Negative binomial generalized linear models indicated that the abundances of nine of the 13 common palm genera were significantly related to a combination of soil cation concentrations, grass phytolith percentages, and palm phytolith PCoA axis scores. These results suggest that phytoliths have potential as quantitative indicators of changes in palm abundances in paleoecological and archaeological reconstructions.
几千年来,人类一直在热带森林中驯化和使用棕榈树。由于缺乏良好的化石记录,过去人类使用棕榈树对现代热带系统的影响程度仍然相对未被探索。然而,棕榈树会产生一系列植物岩,这是一种硅质细胞结构,即使在有机植物物质腐烂后,它们也能很好地保存在土壤中。植物岩在形态上各不相同,可以在群水平上识别,有时甚至可以在种水平上识别。我们分析了63个亚马逊西部样带的棕榈植物岩组合是否反映了棕榈物种的组成和丰度。主坐标分析(PCoA)和Mantel试验表明,棕榈群落和植物岩组合具有相似的更替模式。负二项广义线性模型表明,13个常见棕榈属中有9个属的丰度与土壤阳离子浓度、草植体百分比和棕榈植体PCoA轴评分的组合呈显著相关。这些结果表明,植物岩具有作为古生态和考古重建中棕榈丰度变化的定量指标的潜力。
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引用次数: 0
Searching for Neoproterozoic meteorite impact combusted carbon cenospheres in palynological records 孢粉记录中寻找新元古代陨石撞击燃烧碳微球
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-08-25 DOI: 10.1016/j.revpalbo.2025.105439
Morten Smelror
Carbon cenospheres comparable to those derived from incomplete combustion of coal or fuel-oil droplets appear to be present in Neoproterozoic post-impact sediments in the Gardnos meteorite crater in central Norway and in possible time-equivalent strata in East Finnmark, northern Norway. The Neoproterozoic cenospheres resemble carbon cenospheres found in K-Pg boundary sediments around the world linked to the Chicxulub meteorite impact on Yucatan and in sediments from the Early Cretaceous Mjølnir impact in the Barents Sea. A likely source of the Late Neoproterozoic carbonized cenospheres is hydrocarbons combusted from organic-rich sediments when the bolide hit the shallow marine impact site of the Gardnos meteorite crater.
在挪威中部加诺斯陨石坑的新元古代撞击后沉积物中,以及挪威北部东芬马克的可能与时间相当的地层中,似乎存在与煤或燃料油滴不完全燃烧产生的碳微球。新元古代的微微球与世界各地与尤卡坦的希克苏鲁伯陨石撞击有关的K-Pg边界沉积物和巴伦支海早白垩世Mjølnir撞击的沉积物中发现的碳微微球相似。晚新元古代碳化微球粒的一个可能来源是,当陨石撞击加诺斯陨石坑浅海撞击点时,富含有机物的沉积物燃烧出碳氢化合物。
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