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Cisuralian plant fossils from Khenifra (Central Morocco): Palaeoenvironmental and palaeobiogeographical reconstructions 来自 Khenifra(摩洛哥中部)的 Cisuralian 植物化石:古环境和古生物地理重建
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2024-08-30 DOI: 10.1016/j.revpalbo.2024.105186
Aimad Mazouar , Habiba Aassoumi , Evelyn Kustatscher , Abdelkarim Ait Baha , Maria Mazouar , Hans Kerp

This study focuses on newly collected plant fossils from the Cisuralian successions of the Khenifra Basin in Central Morocco, revealing a diverse assemblage dominated by conifers, pteridosperms, and sphenophytes. The assemblage includes eleven taxa of vegetative and reproductive organs preserved as compressions and impressions. Two pteridosperm species taxa (Neurodontopteris auriculata, Odontopteris subcrenulata) are recorded for the first time from Central Morocco, whereas other taxa are known from the early Permian deposits of Bou Achouch and Tiddas basins. Contrary to previous interpretations, the flora is does not exhibit a mixed character, but instead exhibits a distinct Euramerican affinity, despite the absence of the widespread callipterids Autunia conferta and A. naumannii. These similarities between this flora and coeval Euramerican floras highlight the significant biogeographical connections across Pangea during the Cisuralian. The plant fossil assemblage is dominated by meso- to xerophilous species adapted to drought, with rare hygrophilous elements adapted to humid habitats, reflecting a heterogeneous landscape with various vegetation types.

这项研究的重点是从摩洛哥中部凯尼弗拉盆地的西苏拉利亚演替中新采集的植物化石,揭示了一个以针叶树、翼手目和鞘翅目植物为主的多样化组合。该集合体包括 11 个以压痕和印痕形式保存的无性和生殖器官类群。摩洛哥中部首次记录到两个翼手目类群(Neurodontopteris auriculata 和 Odontopteris subcrenulata),而其他类群则是在布阿舒奇盆地(Bou Achouch)和蒂达斯盆地(Tiddas)的二叠纪早期沉积物中发现的。与之前的解释相反,尽管没有发现广泛分布的胼胝体 Autunia conferta 和 A. naumannii,但该植物群并没有表现出混合特征,而是表现出明显的欧亚亲缘关系。该植物群与共生的欧亚大陆植物群之间的这些相似性凸显了西苏拉时代整个潘吉亚大陆之间重要的生物地理联系。植物化石群以适应干旱的中生至嗜湿物种为主,也有极少数适应潮湿生境的嗜湿物种,反映了不同植被类型的异质景观。
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引用次数: 0
New pollen taxon Syncolpraedapollis angolensis nov. gen. sp. nov.: A noteworthy discovery reported in the preliminary investigation of the latest Eocene-latest Oligocene deposits in the Kwanza Basin, Angola 新的花粉分类群 Syncolpraedapollis angolensis nov:安哥拉宽扎盆地最新始新世-最新渐新世矿床初步调查中的一个值得注意的发现
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2024-08-17 DOI: 10.1016/j.revpalbo.2024.105178
Márcia Mendes , Zélia Pereira , Cristina Rodrigues , Pedro Claude Nsungani

A palynostratigraphic study of the upper Cunga and lower Quifangondo deposits in the Cabo de São Brás section, Kwanza Basin, Angola, has revealed a new pollen named Syncolpraedapollis angolensis nov. gen. sp. nov. This finding was part of a wider survey in the upper Cunga and lower Quifangondo deposits of the Cabo de São Brás section, Kwanza Basin, Angola, covering the latest Eocene to the latest Oligocene.

The novel pollen features unique characteristics, notably a 3-syncolporate structure with well-defined and distinctive pores. The pollen grain is adorned with a free but irregular reticulum with an irregular polygonal mesh. Syncolpraedapollis angolensis nov. gen. sp. nov. is sporadically but consistently observed within the latest Eocene-latest Oligocene interval, conspicuously absent in the underlying Eocene sediments (lower Cunga Formation) and occurring infrequently in the latest Oligocene sediments (lower Quifangondo Formation). Thus, it is plausible to infer a limited age range, likely restricted to the latest Eocene-latest Oligocene, as previous studies in the Kwanza Basin have not documented this pollen.

对安哥拉宽扎盆地圣布拉什卡博段上部Cunga和下部Quifangondo沉积进行的一项古地层学研究发现了一种新的花粉,被命名为nov.gen.sp.nov.。这一发现是对安哥拉宽扎盆地圣布拉什卡博段的上昆加和下基凡贡多矿床进行的更广泛调查的一部分,调查范围涵盖了始新世晚期到渐新世晚期。
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引用次数: 0
Fossil record of Meliolaceae from India sheds new insight into its taxonomy and life cycle 印度楝科植物的化石记录为了解其分类和生命周期提供了新的视角
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2024-08-17 DOI: 10.1016/j.revpalbo.2024.105177
Sampa Kundu, Mahasin Ali Khan

The asexual morph of Meliolaceae has long been a focus of attention for mycologists because there has been a lack of research to establish the asexual morph cycle of Meliolaceae. Besides asexual morph, only a few research works have also been done on the setae (mainly ascomatal setae) of Meliolaceae. In this connection, the use of reliable fossil evidence in evolutionary studies is obligatory. The present study fills this lacuna by reporting the in-situ occurrence of a new enigmatic fossil-genus and species Palaeomeliola indica Kundu and Khan, gen. et sp. nov. with mycelial setae, ascomata, ascomatal setae, ascomatal conoid cells, 4-septate ascospores, phialides, and phialoconidia on the cuticle fragments of a compressed disease-symptomatic monocot leaf recovered from the middle Siwalik (Late Miocene; ca. 12–8 Mya) of Himachal Pradesh, western Himalaya. Palaeomeliola indica is the first fossil-species of Meliolaceae having all sexual and asexual morphs. From this point of view, our new fossil fungus is very distinctive and establishes the greatest preserved fossil record of Meliolaceae. The preserved sexual and asexual morph data sheds new insight into the systematics, life cycle, and evolutionary lineages of Meliolaceae. Furthermore, the findings of this study will form the basis for any future life cycle reconstruction and morphological character evolutionary study of Meliolaceae.

长期以来,由于缺乏对 Meliolaceae 无性形态周期的研究,Meliolaceae 的无性形态一直是真菌学家关注的焦点。除无性形态外,对瓜子黄杨科植物刚毛(主要是气孔刚毛)的研究也为数不多。因此,在进化研究中必须使用可靠的化石证据。本研究报告了一个神秘的新化石属、新种 Palaeomeliola indica Kundu and Khan,gen. et sp.在喜马拉雅山脉西部喜马偕尔邦中斯瓦利克(中新世晚期;约 12-8 Mya)的一片压缩的有疾病症状的单子叶的角质层碎片上发现了菌丝刚毛、子囊、子囊刚毛、子囊球细胞、4 septate ascospores、phialides 和 phialoconidia。Palaeomeliola indica 是第一个具有所有有性和无性形态的 Meliolaceae 化石物种。从这一角度来看,我们的新化石真菌非常独特,并建立了保存最完整的瓜子黄杨科化石记录。保存下来的有性和无性形态数据为研究瓜子黄杨科的系统学、生命周期和进化谱系提供了新的视角。此外,本研究的发现将为今后对瓜子黄杨科进行生命周期重建和形态特征进化研究奠定基础。
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引用次数: 0
Fossil pollen data can reconstruct robust spatial patterns of biodiversity in the past 化石花粉数据可重建过去生物多样性的强大空间模式
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2024-08-16 DOI: 10.1016/j.revpalbo.2024.105175
Kuber P. Bhatta , Xianyong Cao , Vivian A. Felde , John-Arvid Grytnes , Hilary H. Birks , H. John B. Birks

Fossil pollen data are valuable for understanding past spatio-temporal variations in plant diversity. However, deducing robust patterns of fossil pollen-assemblage properties is challenging due to factors related to pollen production, dispersal, and fossilisation; sediment sampling; and nomenclature and taxonomy of the fossil pollen grains reflecting the vegetation that produced them. If the composition pattern of the most recent fossil pollen assemblage concords with those in modern or surface pollen assemblages from the same location(s), the pattern can be regarded as robust.

Using recent (50–500 years) fossil pollen and surface pollen assemblages of angiosperms compiled from the same geographic area of Central Asia, we analyse spatial patterns in compositional turnover and phylogenetic dispersion. These are widely used to quantify different aspects of plant diversity.

Despite differences in the data regarding spatial coverage, number of pollen records, and number of pollen taxa, the spatial pattern in the modern and fossil pollen-assemblage properties across climate zones is broadly similar. Especially, the latitudinal patterns of pollen-assemblage properties are strikingly similar between both data types. However, the degree of concordance varies according to the diversity metric being analysed, and the magnitudes of the metric may differ between both data types. When the data are compiled, processed, and standardised following consistent criteria and analysed using appropriate metrics and methods, spatial patterns in pollen-assemblage properties of fossil pollen data are as robust as those in surface pollen data (∼ modern vegetation). Therefore, regional patterns of compositional turnover and phylogenetic dispersion can be confidently inferred from older fossil assemblages through the Holocene.

化石花粉数据对于了解过去植物多样性的时空变化非常有价值。然而,由于花粉的产生、传播和化石化;沉积物取样;反映产生花粉的植被的化石花粉粒的命名和分类等相关因素,要推断出化石花粉组合特性的可靠模式具有挑战性。如果最近的花粉化石集合体的组成模式与同一地点的现代或地表花粉集合体的组成模式一致,则可认为该模式是稳健的。
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引用次数: 0
Leaf phenology and paleoclimatic and paleoenvironmental insights derived from Yiwupitys elegans of the Yiwu Jurassic Forest, Xinjiang, China 从中国新疆伊吾侏罗纪森林中的Yiwupitys elegans得出的叶片物候以及古气候和古环境启示
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2024-08-16 DOI: 10.1016/j.revpalbo.2024.105176
Xu-Dong Gou , Qun Sui , Hai-Bo Wei , Ji-Yuan Yang , Yu Zhou , Zhuo Feng

Many permineralized conifer stems and trunks have been documented from the Jurassic strata of the Xinjiang Uygur Autonomous Region (Xinjiang), Northwest China. Xylem anatomical studies have showed that these stems and trunks belonged to a diverse range of plants. However, the leaf phenology and the paleoclimatic and paleoenvironmental significance of these fossil plants have rarely been studied. We quantitatively analyzed the anatomical characteristics of a permineralized conifer stem—Yiwupitys elegans Gou et Feng 2023—from the Middle Jurassic Xishanyao Formation in northeastern Xinjiang, which has distinct and wide growth rings in the secondary xylem. Analyses of the Mean Sensitivity and Ring Markedness Index of the growth rings suggest that a stable paleoclimate with regular seasonal fructifications likely prevailed during the Middle Jurassic in this area. The statistical analyses of cell diameters and anatomical features of leaf traces in six successive growth rings indicate that this fossil plant was deciduous. Our study elucidates the ecophysiological characteristics of a Middle Jurassic conifer plant and provides new insights into the corresponding paleoclimate and paleoenvironment based on quantitative cellular analysis of the fossil stem.

在中国西北部新疆维吾尔自治区(新疆)的侏罗纪地层中,记录了许多针叶树的茎干。木质部解剖学研究表明,这些茎干属于多种植物。然而,对这些化石植物的叶片物候学及其古气候和古环境意义的研究却很少。我们定量分析了新疆东北部中侏罗世西山窑地层中的一株过矿化针叶树茎干-Yiwupitys elegans Gou et Feng 2023的解剖特征,该茎干的次生木质部具有明显而宽的生长年轮。对生长年轮的平均灵敏度和年轮标记度指数的分析表明,该地区在中侏罗世时期可能存在着稳定的古气候和有规律的季节性果化。对六个连续生长年轮的细胞直径和叶痕解剖特征的统计分析表明,这种化石植物是落叶植物。我们的研究阐明了中侏罗世针叶植物的生态生理特征,并根据对化石茎干的定量细胞分析,对相应的古气候和古环境提供了新的见解。
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引用次数: 0
Composition of Paleocene forests from Antarctica based on fossil wood 基于化石木材的南极古新世森林的构成
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2024-08-15 DOI: 10.1016/j.revpalbo.2024.105174
Laura Jane Tilley

A new assemblage of fossil wood of Paleocene age from Seymour Island, Antarctic Peninsula, is described. Conifer species have been identified, belonging to the fossil genera Agathoxylon, Phyllocladoxylon, Protophyllocladoxylon, and Podocarpoxylon. Angiosperm fossil wood species are assigned to Nothofagoxylon, Caldcluvioxylon, and Myrceugenellites. New wood types have been described with possible affinity to Atherospermataceae and Asteraceae (daisy family). A newly identified fossil angiosperm species, Aextoxicoxylon jacksius, has been recorded from Antarctica for the first time. It shares traits with modern Aextoxicon punctatum and other Aextoxicoxylon wood from South America but is considered a separate species due to significant differences in vessel density, ray height, and frequency. This study indicates, in accordance with previous studies that conifers were the most common elements in the Paleocene Antarctic vegetation. In particular, Agathoxylon being the most dominant wood type. Nothofagoxylon was the most common angiosperm wood type. Overall the composition of the Antarctic Paleocene vegetation deduced from the presence of fossil wood resembles that of the modern warm to cool temperate forests of Tasmania, New Zealand and southern South America. Possible upland and lowland forest types have been identified, with the lowland forests likely most similar to the modern mixed cool temperate forests on South Island, New Zealand, and Tasmania. The upland forests were similar to the modern open canopy Araucaria-Nothofagus forests on the high Andes today.

本文描述了南极半岛西摩岛出土的一批新的古新世时期的木化石。化石中的针叶树物种已被确认,分别属于Agathoxylon、Phyllocladoxylon、Protophyllocladoxylon和Podocarpoxylon化石属。被子植物的木材化石种类被归入 Nothofagoxylon、Caldcluvioxylon 和 Myrceugenellites。新描述的木材类型可能与 Atherospermataceae 和 Asteraceae(菊科)有亲缘关系。南极洲首次记录了一个新发现的被子植物化石物种--Aextoxicoxylon jacksius。它与现代的 Aextoxicon punctatum 和南美洲的其他 Aextoxicoxylon 木材具有相同的特征,但由于在血管密度、射线高度和频率方面存在显著差异,因此被认为是一个独立的物种。这项研究表明,与之前的研究一致,针叶树是古新世南极植被中最常见的植物。特别是,Agathoxylon 是最主要的木材类型。Nothofagoxylon是最常见的被子植物木材类型。总体而言,根据化石木材的存在推断出的南极古新世植被组成与塔斯马尼亚、新西兰和南美洲南部的现代暖温带至寒温带森林相似。已经确定了可能的高地和低地森林类型,其中低地森林可能与南岛、新西兰和塔斯马尼亚岛上的现代凉温带混交林最为相似。高地森林类似于今天安第斯山脉高地的现代开阔树冠Araucaria-Nothofagus森林。
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引用次数: 0
A microfossil-based object biogeography of a set of stocks from the Rijksmuseum in Amsterdam 基于微化石的阿姆斯特丹里克斯博物馆藏品生物地理学
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2024-08-10 DOI: 10.1016/j.revpalbo.2024.105172
Hans Piena , Bas van Geel , Tom Hakbijl , Arie J. Kalis , Pim (W.O.) van der Knaap , Jacqueline F.N. van Leeuwen , Kees Nooren

In 2019 the Rijksmuseum in Amsterdam acquired a set of oak stocks about which little was known. Radiocarbon wiggle-matching revealed that the tree used to make the stocks was likely felled around 1800 C.E., while ancient DNA indicated the tree's growing zone to be Central Europe. The question remained where and how these stocks were used. To answer these questions, traces of former use at the surface of the stocks were studied and sediment from cracks and holes in the stocks was analyzed for pollen, fungal spores, diatoms and other algae, phytoliths, and insects. The biogeographical information of the recorded taxa shows that the stocks had been used in the western Iberian Peninsula. Although the sediments could have entered in various ways and at various moments, a dungeon seems the most likely context in which these stocks have been used.

2019 年,阿姆斯特丹的里克斯博物馆获得了一套橡木托,而人们对这套橡木托知之甚少。通过放射性碳比对发现,用于制作橡木托的树木很可能是在公元前 1800 年左右被砍伐的,而古老的 DNA 表明该树木的生长区域位于中欧。问题仍然是这些存货在哪里以及如何使用。为了回答这些问题,我们研究了树种表面以前使用过的痕迹,并对树种裂缝和孔洞中的沉积物进行了分析,以寻找花粉、真菌孢子、硅藻和其他藻类、植物石和昆虫。所记录分类群的生物地理信息表明,这些种群曾在伊比利亚半岛西部使用过。虽然沉积物可能以各种方式在不同时间进入,但地宫似乎是这些种群最有可能的使用环境。
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引用次数: 0
A new species of Neocalamites from the Upper Buntsandstein (Anisian) of Üdingen (Rur Eifel, Germany) 来自于Üdingen(德国Rur Eifel)上Buntsandstein(Anisian)的Neocalamites新种
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2024-08-10 DOI: 10.1016/j.revpalbo.2024.105173
Iris I. Kuipers , Johanna H.A. van Konijnenburg-van Cittert , Friederike Wagner-Cremer

Well preserved Early-Middle Triassic plant fossils from Northwest Europe are relatively rare due to the environmental conditions within the Germanic basin during this time period. The only sphenophyte species commonly described in the floras that are found in the area, is Equisetites mougeotii. However, only stems without leaf sheaths were ever reported. Here, we describe sphenophyte stems with free standing leaves from the Upper Buntsandstein of Üdingen, Germany, which we attribute to the genus Neocalamites based on this characteristic and the difference, mainly in stem size, of the Üdingen material to E. mougeotii. Most of the material consists of internodal, infertile stems that are unbranched and vary in width. A few of these stems are three-dimensionally preserved. Due to the difference of the material to the other, rarely present, Neocalamites species reported from the Anisian of Western Europe, we propose the plant fossils from Üdingen to be a new species called Neocalamites vanderburghii n.sp.

由于这一时期日耳曼盆地的环境条件,西北欧保存完好的早中三叠世植物化石相对稀少。在该地区发现的植物区系中,唯一常见的海绵状植物物种是Equisetites mougeotii。不过,只有无叶鞘的茎被报道过。在这里,我们描述了来自德国于丁根(Üdingen)上布恩桑德斯坦(Upper Buntsandstein)的具有独立叶片的茎叶植物,根据这一特征以及于丁根材料与 E. mougeotii 的差异(主要是茎的大小),我们将其归于 Neocalamites 属。大部分材料都是节间不孕茎,不分枝,宽度不一。其中少数茎干保存有三维结构。由于这些材料与西欧安尼西亚报告的其他很少出现的新卡米蒂(Neocalamites)物种不同,我们建议将于丁根发现的植物化石作为一个新物种,名为 Neocalamites vanderburghii n.sp.。
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引用次数: 0
How many pollen grains should we count? – A basic statistical view 我们应该计算多少花粉粒?- 基本统计观点
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2024-08-02 DOI: 10.1016/j.revpalbo.2024.105126
Chengyu Weng , Henry Hooghiemstra

In pollen analysis, one question is often being asked: how many pollen grains should be counted for one sample? The proper count number should guarantee the reliability of the abundance (usually expressed as percentage) of the pollen taxa concerned in the sample. Practically, this number is usually more than 100 grains, but seldom over 1000. Most people tend to count at least 300 pollen grains (often of terrestrial plants)for each sample. However, this number is more likely based on the experience of earlier palynologists rather than on theoretical consideration. In this paper, we apply a simple statistical theory to evaluate the reliability of the observed pollen abundance with different total count number. The reliability is evaluated based on the error range. The results show that the reliability depends on both the true pollen abundance of the taxon concerned and the total count number: a low abundant pollen taxon needs a high count number to obtain a reliable data set. The conclusion is that there is no universal “proper” total count number for each sample. The total count number should be determined by both the abundance of the interested taxa and the research objective. The commonly used 300 grains may be good enough for most general vegetation reconstructions and for the accurate survey of pollen taxa with more than 10% abundance. For less abundant taxa, higher count numbers are suggested.

在花粉分析中,人们经常会问这样一个问题:一个样本应该计数多少花粉粒?适当的计数数量应能保证样本中相关花粉类群丰度(通常以百分比表示)的可靠性。实际上,这个数量通常超过 100 粒,但很少超过 1000 粒。大多数人倾向于为每个样本计数 300 粒花粉(通常是陆生植物的花粉)。不过,这个数字更可能是基于早期古生物学家的经验,而非理论上的考虑。在本文中,我们运用简单的统计理论来评估不同计数总数下观察到的花粉丰度的可靠性。可靠性是根据误差范围来评估的。结果表明,可靠性取决于相关类群的真实花粉丰度和总计数:低丰度的花粉类群需要高计数才能获得可靠的数据集。结论是,对于每个样本而言,并不存在通用的 "适当 "总计数。总计数应该根据相关类群的丰度和研究目标来确定。对于大多数一般植被重建和准确调查丰度超过 10%的花粉类群来说,常用的 300 粒可能就足够了。对于丰度较低的分类群,建议使用更高的计数粒数。
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引用次数: 0
Taxonomic and environmental significance of Poaceae and Cyperaceae phytoliths from the Northern Territory, Australia 澳大利亚北部地区诗歌科和桧柏科植物岩石的分类和环境意义
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2024-07-30 DOI: 10.1016/j.revpalbo.2024.105169
Kelsey C. Boyd , Carlos E. Cordova , Haidee R. Cadd , Cassandra Rowe , Tim J. Cohen

Phytoliths are a valuable tool for reconstructing past grassy ecosystems. However, they are not commonly utilised as a palaeoecological proxy in Australia due to a lack of modern reference material from plants, particularly grasses. This study analyses phytolith reference material from 49 grass and 4 sedge species from the Northern Territory, Australia. This is used to develop a detailed classification scheme and multivariate morphospace analysis to examine taxonomic and ecological patterns of grass silica short cell phytolith (GSSCP) production within the Poaceae (grass) family. The results of detrended correspondence analysis show that grass subfamilies, tribes, and genera in this region can be differentiated through phytolith assemblages, and that these assemblages reflect the environmental conditions associated with various grass taxa. The developed detailed classification system and examination of co-occurrence patterns reduces redundancy of phytolith morphotypes across subfamilies, improving taxonomic and palaeoecological resolution of phytolith assemblages. Finally, based on the results of this study, a detailed scheme is presented for palaeoecological reconstructions in grassy ecosystems of northern Australia.

植物残片是重建过去草地生态系统的重要工具。然而,由于缺乏植物(尤其是草类)的现代参考材料,它们在澳大利亚并没有被普遍用作古生态学的替代物。本研究分析了澳大利亚北部地区 49 种草和 4 种莎草的植物岩石参考材料。通过详细的分类方案和多变量形态空间分析,研究了禾本科(Poaceae)内禾本科硅质短细胞植物岩石(GSSCP)的分类和生态模式。去趋势对应分析的结果表明,该地区的禾本科亚科、科和属可通过植硅体组合加以区分,这些组合反映了与各种禾本科类群相关的环境条件。详细的分类系统和共同出现模式的研究减少了亚科之间植物形态的冗余,提高了植物群落的分类学和古生态学分辨率。最后,根据本研究的结果,提出了澳大利亚北部草地生态系统古生态重建的详细方案。
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