Review of Palaeobotany and Palynology最新文献_第5页

Palynotaxonomy of species and genera of the Cyanocladus and Trichocladus clades (Lamiaceae: Hyptidinae) 青枝和毛枝的种属分类（紫枝科：棘枝科）

IF 1.7 3区地球科学 Q2 PALEONTOLOGY

Review of Palaeobotany and Palynology

Pub Date : 2026-01-01 Epub Date: 2025-08-17 DOI: 10.1016/j.revpalbo.2025.105428

Renata Jacomo Paixão de Carvalho , Raymond Mervey Harley , José Floriano Barêas Pastore , Cláudia Barbieri Ferreira Mendonça , Vania Gonçalves-Esteves

This study analyzed the pollen morphology of 22 taxa of the Cyanocladus and Trichocladus clades. The aim was to describe and characterize pollen grains and assess whether pollen morphology corroborates the recent taxonomy of the group. Only genera occurring in Brazil were studied, including Cyanocephalus (Pohl ex Benth.) Harley & J.F.B.Pastore (7 spp.), the endemic genus Eplingiella Harley & J.F.B.Pastore (3 spp.), Marsypianthes Mart. ex Benth. (4 spp.), Martianthus Harley & J.F.B.Pastore (3 spp.), and Medusantha Harley & J.F.B.Pastore (5 spp.). Acetolyzed pollen grains were measured, described, and photomicrographed. Non-acetolyzed pollen grains were examined by scanning electron microscopy. All taxa analyzed have pollen grains shed in monads, medium to large in size, isopolar, (4)-6-(8–10)colpate, ranging from suboblate to prolate, with a small to very small polar area. The colpi are long to very long, broad, with ornamented membranes and an acute apex, and lack distinct margins. An operculum was observed in all species of Cyanocephalus. The sexine is bireticulated in all other species and genera. The results showed that the pollen morphology of the studied genera was informative in classifying taxa and distinguishing genera, demonstrating some notable parallels with current taxonomic opinion. In principal component analysis and hierarchical cluster analysis, morphometric characteristics approximated and grouped, respectively, species of the genera Eplingiella and Martianthus. These genera exhibited great variation in pollen shape.

本研究分析了青枝门和毛枝门22个分类群的花粉形态。目的是描述和表征花粉粒，并评估花粉形态是否证实了该群体的最新分类。只研究了发生在巴西的属，包括Cyanocephalus （Pohl ex Benth.）。Harley &； J.F.B.Pastore（7种），特有属Eplingiella Harley &； J.F.B.Pastore（3种），Marsypianthes Mart。Benth交货。（4席）、马田苏斯·哈利& j.f.b.帕斯托雷（3席）和梅杜莎·哈利& j.f.b.帕斯托雷（5席）。对乙酰化的花粉粒进行测量、描述和显微照相。用扫描电镜观察未乙酰化的花粉粒。所分析的所有分类群花粉粒均以单元体形式脱落，大小从中到大，等极性，(4)-6-(8-10)colpate，从近卵圆形到长形，极面积很小到很小。鞘长到非常长，宽，具装饰的膜和一个锐尖的先端，并且缺乏明显的边缘。在所有种类的蓝头科植物中都观察到有盖。在所有其他物种和属中，性器官都是双网状的。结果表明，所研究属的花粉形态对分类群的划分和属的区分具有重要的参考价值，与目前的分类观点有显著的相似之处。在主成分分析和层次聚类分析中，分别对Eplingiella属和Martianthus属的形态特征进行了近似和分组。这些属在花粉形态上表现出很大的差异。

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引用次数: 0

Monitoring seasonal pollen deposition in an arid environment using Tauber traps: ecological and aerobiological insights from Kashan, central Iran 利用陶伯捕集器监测干旱环境中的季节性花粉沉积：来自伊朗中部卡尚的生态和有氧生物学见解

IF 1.7 3区地球科学 Q2 PALEONTOLOGY

Review of Palaeobotany and Palynology

Pub Date : 2026-01-01 Epub Date: 2025-10-22 DOI: 10.1016/j.revpalbo.2025.105462

Elham Mahlooji , Morteza Djamali , Hossein Akhani , Emmanuel Gandouin , Hossein Batooli , Marzieh Heidarzadeh Arani , Mohammad-Reza Mozdianfard

Monitoring atmospheric pollen deposition with Tauber traps has long supported palaeoecological studies in temperate regions of Europe, but their application in arid regions and their potential in aerobiology remain underexplored. We installed Tauber traps at three sites in Kashan and neighboring towns on the western margin of the central Iranian desert, with weekly and monthly (4-weekly) collections over one year, to assess their efficiency in capturing seasonal airborne pollen. The traps reliably reflected both regional desert-steppe vegetation and urban or peri-urban plantations. Comparison with clinical allergy data shows that Chenopodiaceae (s. str.) and Artemisia, together with Morus alba, Olea europaea, and Cupressus, are the major airborne allergens in Kashan. Our findings stress that the allergenic importance of Chenopodiaceae and Artemisia—dominant in summer and autumn pollen rains—has been underestimated in regional aerobiological surveys and clinical testing, which typically rely on a narrow set of non-native extracts. We propose that Tauber traps, as a simple and inexpensive tool, could complement aerobiological monitoring in remote drylands and inform both ecological and medical studies.

利用陶伯捕集器监测大气花粉沉积长期以来一直支持欧洲温带地区的古生态学研究，但其在干旱地区的应用及其在空气生物学中的潜力仍未得到充分探索。我们在伊朗中部沙漠西部边缘的卡尚和邻近城镇的三个地点安装了陶伯捕集器，在一年的时间里每周和每月（4周）收集一次，以评估它们捕捉季节性空气传播花粉的效率。捕集器可靠地反映了区域荒漠草原植被和城市或城郊人工林。与临床变态反应资料比较发现，喀山地区空气中主要的变态反应原为藜科（s.s str.）和青蒿科，以及桑、油橄榄和柏树。我们的研究结果强调，Chenopodiaceae和蒿属植物在夏季和秋季花粉雨中占主导地位，在区域空气生物学调查和临床试验中被低估了，这些调查和临床试验通常依赖于一组狭窄的非本地提取物。我们建议，陶伯捕集器作为一种简单而廉价的工具，可以补充偏远旱地的空气生物学监测，并为生态和医学研究提供信息。

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引用次数: 0

New fossil evidence from southwestern China sheds light on the formation of modern distribution of Toddalia (Rutaceae) 中国西南地区新发现的化石证据揭示了龙牙亚（龙牙科）现代分布的形成

IF 1.7 3区地球科学 Q2 PALEONTOLOGY

Review of Palaeobotany and Palynology

Pub Date : 2026-01-01 Epub Date: 2025-10-16 DOI: 10.1016/j.revpalbo.2025.105460

Jiang-Min Chen , Xue-Ping Ji , Chun-Xia Zhang , Yu Ji , Jun Yang , Huan-Chong Wang , Yun-Heng Ji , Yong-Jiang Huang

Toddalia Juss. is a genus in Rutaceae, with a modern geographical range that extends from West and East Africa to South, East and Southeast Asia. The genus has a good fossil record in Europe, where it has become extinct, but is poorly represented in the geological record of Asia, where it flourishes today. This study reports the second fossil record of Toddalia in Asia, based on three-dimensional seed remains recovered from the latest Miocene Baoshan Basin in western Yunnan, southwestern China. The fossil seeds are characterized by their reniform shape, smooth surface, large hilar scar, unique raphe excrescence and layered seed coat. Following morphological and anatomical comparisons with both extant and other fossil species, the fossil seeds were assigned to T. nanlinensis, a species first reported from the Early-Middle Miocene Longchuan Basin in southwestern Yunnan. This new fossil finding suggests that the Baoshan Basin probably had forest vegetation in a warm, humid climate at some time during the latest Miocene. This is in line with previous suggestions based on other fossils, such as hominoids (Lufengpithecus sp.) and other mammals, found from the same sedimentary layer of the basin. Together with the previous record, our fossil discovery mirrors continuous presence of Toddalia in southwestern China since at least the Early-Middle Miocene. This provides insight into how the modern distribution range of the genus formed in this region and even in Asia.

Toddalia汁液。是芸香科的一个属，其现代地理分布范围从西非和东非延伸到南亚、东亚和东南亚。该属在欧洲有很好的化石记录，在那里它已经灭绝了，但在亚洲的地质记录中却很少有代表，在那里它今天很繁荣。本研究报告了在滇西宝山盆地中新世晚期发现的三维种子化石，这是亚洲第二次对幼童龙的化石记录。化石种子具有肾形、表面光滑、门瓣瘢痕大、独特的中缝赘肉和层状种皮等特点。通过与现存和其他化石种的形态和解剖比较，化石种子归属于滇西南龙川盆地早中新世-中中新世首次报道的一种T. nanlinensis。这一新的化石发现表明，在中新世晚期的某个时候，在温暖潮湿的气候下，宝山盆地可能有森林植被。这与先前在盆地同一沉积层中发现的其他化石（如人猿（陆丰猿人）和其他哺乳动物）的建议一致。与之前的记录相结合，我们的化石发现反映了至少从中新世早期到中中新世以来，中国西南部就一直存在着托德利亚。这为了解该属的现代分布范围如何在该地区甚至在亚洲形成提供了见解。

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引用次数: 0

First record of Cyathocarpus (Marattiales) in Gondwana: A new species from the Permian of Patagonia, Argentina 冈瓦纳地区Cyathocarpus （Marattiales）的首次记录：阿根廷巴塔哥尼亚二叠纪的一新种

IF 1.7 3区地球科学 Q2 PALEONTOLOGY

Review of Palaeobotany and Palynology

Pub Date : 2026-01-01 Epub Date: 2025-09-11 DOI: 10.1016/j.revpalbo.2025.105447

Gonzalo Rodriguez Rizk, Bárbara Cariglino

Marattialean ferns were key components of Paleozoic wetland ecosystems, yet their biogeographic history in Gondwana remains poorly resolved. While Cyathocarpus is well documented from Euramerica and Cathaysia, no species had been recognized in Gondwana—until now. We describe Cyathocarpus polinensis sp. nov. from the Guadalupian La Golondrina Formation of Patagonia, Argentina, representing the first Gondwanan record of the genus. The species exhibits a distinctive arrangement of synangial rows that do not contact the midvein or pinnule margins and are consistently absent from the apical and basal regions of the pinnules. The midvein is conspicuous, straight, and divides fanwise near the apex; lateral veins bifurcate once, close to the midvein. This discovery extends the paleogeographic range of Cyathocarpus and supports a dispersal scenario from Euramerica into southern Gondwana during the Cisuralian. These findings call for a critical reassessment of comparable Gondwanan taxa and underscore the role of southwestern Gondwana as a refugium and center of marattialean diversification during postglacial climatic recovery.

摘要蕨类植物是古生代湿地生态系统的重要组成部分，但其在冈瓦纳的生物地理历史仍未得到很好的解决。虽然Cyathocarpus在欧洲和华夏都有很好的记录，但直到现在，冈瓦纳还没有发现任何物种。我们描述了来自阿根廷巴塔哥尼亚Guadalupian La Golondrina组的Cyathocarpus polinensis sp. 11 .，代表了该属的第一个冈瓦纳记录。该种表现出一种独特的排列，不接触中脉或小穗边缘的同质行，并且始终不存在于小穗的顶端和基部区域。中脉明显，直，并在先端附近扇形分开；侧脉分叉一次，靠近中脉。这一发现扩大了Cyathocarpus的古地理范围，并支持了一种从欧洲到冈瓦纳南部的分布情况。这些发现要求对冈瓦纳的类群进行批判性的重新评估，并强调冈瓦纳西南部在冰川后气候恢复期间作为避难所和物种多样化中心的作用。

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引用次数: 0

Pollen morphology of Pseudanthus (Picrodendraceae) 拟蕨科植物花粉形态研究

IF 1.7 3区地球科学 Q2 PALEONTOLOGY

Review of Palaeobotany and Palynology

Pub Date : 2026-01-01 Epub Date: 2025-08-13 DOI: 10.1016/j.revpalbo.2025.105426

Angelika Till , Silvia Ulrich , David J. Cantrill , Friðgeir Grímsson

This is the first comprehensive investigation on the pollen morphology of Pseudanthus, a small genus within the Picrodendraceae, comprising nine species all endemic to Australia. This study presents the pollen morphology of eight out of the nine currently accepted Pseudanthus species using light and scanning electron microscopy. Pollen from five of the species (incl. subspecies) are described here for the first time. Pollen from different Pseudanthus species appears alike, sharing many overlapping features, including sculpture patterns observed with both LM and SEM. However, detailed examination reveals species-specific differences such as size and aperture number, that aid in discriminating pollen of the individual species. In general, Picrodendraceae pollen is mostly isodiametric to oblate, spherical to spheroidal in shape, and either exhibits a pantoporate or stephanoporate aperture configuration, as well as echinate sculpture. Although, Pseudanthus pollen shares these morphological features, being isodiametric to slightly oblate, pantoporate, and echinate, the pollen of this genus can be differentiated from other closely related Picrodendraceae (Kairothamnus, Micrantheum, Neoroepera, Scagea, and Stachystemon) based on set of traits observed with combined LM and SEM.

本文首次对拟花属（Pseudanthus）花粉形态进行了全面的研究。拟花属是拟花科（Picrodendraceae）中的一个小属，由澳大利亚特有的9个品种组成。本文利用光镜和扫描电镜研究了目前公认的9种假花属植物中8种的花粉形态。其中5种（包括亚种）的花粉为首次报道。不同假药属植物的花粉看起来很相似，具有许多重叠的特征，包括用LM和SEM观察到的雕刻图案。然而，详细的研究揭示了物种特有的差异，如大小和孔数，这有助于区分单个物种的花粉。一般来说，Picrodendraceae花粉的形状大多为等径至扁圆，球形至球形，并表现出宽孔或深孔的孔径结构，以及刺状雕刻。虽然假花属的花粉具有等径到略平球形、泛孔状和棘刺状的形态特征，但根据LM和SEM结合观察到的一组性状，该属的花粉可以与其他密切相关的Picrodendraceae （Kairothamnus、Micrantheum、Neoroepera、Scagea和Stachystemon）区分开来。

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引用次数: 0

Late Holocene vegetation dynamics and Indian Summer Monsoon evolution from the Core Monsoon Zone, India 印度核心季风带的晚全新世植被动态与印度夏季风演化

IF 1.7 3区地球科学 Q2 PALEONTOLOGY

Review of Palaeobotany and Palynology

Pub Date : 2026-01-01 Epub Date: 2025-10-06 DOI: 10.1016/j.revpalbo.2025.105455

Nagendra Prasad , Mohammad Firoze Quamar , Paulramasamy Morthekai , Maneesha Muraleedharan ET , Pooja Tiwari , Biswajeet Thakur , Anupam Sharma

Understanding the vegetation response to the Late Holocene Indian Summer Monsoon (ISM) variability from the Core Monsoon Zone (CMZ), India, during the Late Holocene, is the principal aim of the present study. Pollen analysis of a lacustrine sediment core from the Korba District of Chhattisgarh State (CMZ) demonstrated that between ca. 3560 and 2860 cal yr BP, savannah vegetation dominated the landscape in a cool-dry climate, suggesting reduced ISM rainfall. A shift in the vegetation from savannah to wooded savannah during ca. 2860 to 1960 cal yr BP was observed in a moderately strengthened ISM rainfall regime. From ca. 1960 to 890 cal yr BP, the region experienced a transformation from the wooded savannah vegetation to mixed tropical deciduous forests in a warm-humid climate, reflecting a comparatively increased ISM rainfall. From ca. 890 to 225 cal yr BP (CE1060–1725), dense mixed tropical deciduous forests came into being in a warm and relatively more humid climate, indicating intensified ISM rainfall. The initial part (CE 1060–1400) of this phase corresponds with the global Medieval Climate Anomaly (MCA; CE 700–1400). Agricultural practices and other human activities were suggested, based on the presence of Cerealia pollen, and other cultural pollen taxa, respectively.

了解印度核心季风带（CMZ）晚全新世期间植被对印度夏季风（ISM）变率的响应是本研究的主要目的。对Chhattisgarh邦Korba地区湖泊沉积物岩心的花粉分析表明，在约3560 - 2860 calyr BP之间，热带稀树草原植被在干冷气候中占主导地位，表明ISM降雨量减少。在中等强度的ISM降雨条件下，在约2860 ~ 1960 cal yr BP期间，植被从草原向森林草原转变。约1960 ~ 890 calyr BP，该地区经历了由树木繁茂的草原植被向热带落叶混交林的转变，气候温暖湿润，ISM降水相对增加。约890 ~ 225 cal yr BP (CE1060-1725)，在温暖且相对湿润的气候中形成了茂密的热带混交林，表明ISM降水加剧。这一阶段的初始部分（CE 1060-1400）与全球中世纪气候异常（MCA; CE 700-1400）相对应。根据谷类花粉和其他文化花粉分类群的存在，分别提出了农业实践和其他人类活动的建议。

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引用次数: 0

Taphonomic Completeness: A new metric for assessing fossil leaf preservation using the Spitsbergen Cenozoic fossil flora 埋藏完整性：利用斯匹次卑尔根新生代化石区系评估化石叶片保存的新指标

IF 1.7 3区地球科学 Q2 PALEONTOLOGY

Review of Palaeobotany and Palynology

Pub Date : 2026-01-01 Epub Date: 2025-10-30 DOI: 10.1016/j.revpalbo.2025.105464

Harrison O'Rourke , Eimear Nic Lughadha , Karen L. Bacon

Leaves are the most common type of plant macrofossil found in the plant fossil record; however, comparing leaf preservation quality across plant groups, sites, and time periods is challenging. A new metric was devised that allowed the quality of leaf preservation to be quantitatively measured for leaf fossils preserved from different localities, timepoints and plant life forms, based on a suite of leaf traits. The metric produces a score between 6 (high quality preservation) and 0 (low quality preservation). The leaf traits visible in the Spitsbergen Cenozoic fossil flora were studied to analyse the effect of leaf traits on preservation potential. Non-metric multidimensional scaling indicated that plant life form strongly influenced the leaf traits preserved within the Spitsbergen fossil flora, while stratigraphy had a much weaker effect, indicating that depositional environment and climate influenced preservation less than plant life form. The ‘Taphonomic Completeness’ metric indicated that the gymnosperms and ferns had a significantly higher leaf preservation quality than the woody angiosperms and the herbaceous angiosperms, with the latter group having the lowest quality preservation overall. This result suggests that there may be a bias against the leaves of herbaceous species being preserved in the fossil record, which has implications for interpreting the plant fossil record and how paleo ecosystems functioned. Additionally, the development of this quantitative ‘Taphonomic Completeness’ metric provides a standardised approach that can be widely applied to fossil leaf assemblages from other sites and time periods, allowing robust, large-scale comparisons of preservation quality across fossil floras.

叶子是植物化石记录中最常见的植物大化石类型；然而，比较不同植物类群、地点和时间段的叶片保存质量是具有挑战性的。设计了一种新的度量标准，可以根据一系列叶片特征，对不同地点、时间点和植物生命形式保存的叶片化石的叶片保存质量进行定量测量。该度量产生的分数在6（高质量保存）和0（低质量保存）之间。研究了斯匹次卑尔根新生代植物区系中可见的叶片性状，分析了叶片性状对保存潜力的影响。非度量多维尺度表明，植物生命形式对斯匹次卑尔根化石区系中保存的叶片性状影响较大，而地层的影响较弱，表明沉积环境和气候对保存的影响小于植物生命形式。结果表明，裸子植物和蕨类植物的叶片保存质量显著高于木本被子植物和草本被子植物，草本被子植物的叶片保存质量最低。这一结果表明，化石记录中保存的草本物种的叶子可能存在偏见，这对解释植物化石记录和古生态系统的功能具有重要意义。此外，这种定量的“埋藏完整性”指标的发展提供了一种标准化的方法，可以广泛应用于其他地点和时期的化石叶片组合，允许对化石区系的保存质量进行可靠的大规模比较。

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引用次数: 0

An illustrated key for the identification of pollen from Rajaji National Park, India 印度拉贾吉国家公园花粉鉴定的图解钥匙

IF 1.7 3区地球科学 Q2 PALEONTOLOGY

Review of Palaeobotany and Palynology

Pub Date : 2026-01-01 Epub Date: 2025-08-22 DOI: 10.1016/j.revpalbo.2025.105438

Ruchita Rawat , Prabhawati Tiwari , Shivani Negi , Dinesh Singh Rawat

Accurate identification of pollen grains is fundamental to diverse fields such as palaeobotany, melissopalynology, aerobiology, and forensic science. This study presents a comprehensive illustrated key for the identification of pollen from Rajaji National Park (RNP). The park is situated in the Shivalik foothills of the Garhwal Himalaya, a region renowned for its rich floral diversity and significant ecological value. Extensive field surveys were conducted from 2021 to 2024, and polliniferous material from 302 plant species, representing 229 genera and 67 families, was collected. Pollen grains were prepared following the standard acetolysis protocol and subsequently examined using both light microscopy (LM) and scanning electron microscopy (SEM). Based on dispersal units and aperture type, the taxa were classified into 24 distinct pollen classes. An illustrated pollen identification key was developed, comprising a general key to differentiate pollen into distinct classes, followed by separate keys for each class. This key serves as a foundational reference for palynological research in Himalayan foothills. The work addresses a critical gap in regional palynological studies and contributes to strengthening pollen databases in India.

花粉粒的准确鉴定是古植物学、微生物学、空气生物学和法医学等多个领域的基础。本研究提出了一种鉴定Rajaji国家公园（RNP）花粉的综合图解密钥。该公园位于加尔瓦尔喜马拉雅山脉的Shivalik山麓，该地区以其丰富的植物多样性和重要的生态价值而闻名。从2021年到2024年进行了广泛的实地调查，收集了67科229属302种植物的传粉材料。按照标准的乙酰解方案制备花粉粒，随后使用光学显微镜（LM）和扫描电子显微镜（SEM）进行检查。根据花粉的散布单位和孔型，将其划分为24个不同的花粉类。开发了一个图解的花粉识别键，包括一个将花粉区分为不同类别的通用键，然后是每个类别的单独键。该钥匙可作为喜马拉雅山麓孢粉学研究的基础参考。这项工作解决了区域孢粉学研究中的一个关键空白，并有助于加强印度的花粉数据库。

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引用次数: 0

Chemosystematic studies of Devonian coal-forming plants: Barzas Orestovia-like plants compressions 泥盆纪造煤植物的化学系统研究：Barzas orestovia样植物压缩

IF 1.7 3区地球科学 Q2 PALEONTOLOGY

Review of Palaeobotany and Palynology

Pub Date : 2026-01-01 Epub Date: 2025-09-23 DOI: 10.1016/j.revpalbo.2025.105452

Lyudmila Kocheva , Anatoly Karmanov , Olga Telnova , Alexey Gomankov , Vladimir Lutoev

Orestovia-like plants are of great scientific and practical interest for various fields of natural science, forming the most ancient unique Devonian Barzas coals in the Kuznetsk Basin, Russia. However their natural affinity (higher plants or algae) remains disputable. This work studies Barzas compressions of Orestovia-like plants from the standpoint of chemosystematics. We proposed to use the aromatic biopolymer lignin, which is included in all higher terrestrial plants, as a marker of organic matter. To prove the presence of lignin and establish its structure, a complex of physical and chemical methods was used: FTIR and EPR spectroscopy, pyrolytic gas chromatography–mass spectrometry, scanning electron microscopy, X-ray fluorescence and X-ray phase analysis. Based on the obtained results, we found the presence of transformation products of compositionally homogeneous p-coumaric protolignin in the substance of plant remains. The obtained data do not exclude the validity of the hypothesis that the Devonian Orestovia-like plants belong to higher plants.

orestova类植物在自然科学的各个领域具有重要的科学和实用价值，在俄罗斯库兹涅茨克盆地形成了最古老独特的泥盆纪Barzas煤。然而，它们的天然亲和力（高等植物或藻类）仍有争议。本文从化学系统学的角度研究了orestovia类植物的Barzas压缩。我们建议使用所有高等陆生植物中都含有的芳香族生物聚合物木质素作为有机质的标记物。为了证明木质素的存在并确定其结构，采用了一系列物理和化学方法：FTIR和EPR光谱、热解气相色谱-质谱、扫描电镜、x射线荧光和x射线相分析。在此基础上，我们发现在植物残体物质中存在组成均质的对香豆素原木质素转化产物。获得的数据并不排除泥盆纪orestovia类植物属于高等植物的假设的有效性。

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引用次数: 0

Cupule-bearing cones of Jarudia (Doyleales) and associated leaves from the Early Cretaceous of the New Siberian Islands, Arctic Russia 俄罗斯北极地区新西伯利亚群岛早白垩世的刺叶属植物的带对偶的球果及其伴生叶

IF 1.7 3区地球科学 Q2 PALEONTOLOGY

Review of Palaeobotany and Palynology

Pub Date : 2026-01-01 Epub Date: 2025-09-20 DOI: 10.1016/j.revpalbo.2025.105451

Natalya Nosova , Peter R. Crane , Gongle Shi , Ksenia Domogatskaya

A new species of cupule-bearing seed cone assigned to Jarudia (Doyleales) is described from the Lower Cretaceous (Aptian–Albian) Balyktakh Formation of Kotelnyi Island in the New Siberian Islands. The new species, Jarudia borealis sp. nov. is an elongated, cylindrical seed cone consisting of a main axis with helically arranged, densely spaced bract-cupule axis complexes, each of which terminates in a single incurved cupule. Cupule-bearing stalks are thin proximally where they are fused with a narrow, elongated bract. Above, they are free from the bract, gradually widen toward the tip and are curved adaxially relative to the main axis. Each cupule is formed by the wide flattened distal part of the cupule stalk, one median and two lateral flaps, and is quadrangular in transverse section. The flattened cupule stalk and three flaps are fused at the apex of the cupule but are free toward the base. Micro-CT scans reveal that most cupules are empty, but maceration of remaining carbonaceous material produced pieces of the cupule cuticle, as well as fragments of the nucellar cuticle and megaspore membrane. Stomata, occasionally present in the outer epidermis of the cupules, are haplocheilic, with 5–8 subsidiary cells. The megaspore membrane is two layered comprising a foot layer and branched, interconnected bacula. The cupule-bearing cones of Jarudia borealis sp. nov. occur closely associated with numerous remains of linear leaves described as Phoenicopsis arcticus sp. nov., as well as rare leaf fragments of Pityophyllum sp. and Ginkgoites sp.

描述了新西伯利亚群岛Kotelnyi岛下白垩世（Aptian-Albian） Balyktakh组Jarudia （Doyleales）的一新种。这个新种，Jarudia borealis sp. nov.是一个细长的圆柱形种子球果，由一个主轴和螺旋状排列的密距苞片-偶联轴组成，每个轴在单个弯曲的偶联中终止。具羽对的茎薄的近端在那里它们与一狭窄的，拉长的苞片融合。在上面，它们脱离苞片，逐渐向尖端加宽，并且相对于主轴向正面弯曲。每一羽节由羽节柄的宽而扁平的远端部、一个中瓣和两个侧瓣组成，横切面呈四角形。扁平的羽对柄和三个瓣在羽对顶端融合，但向基部自由。微ct扫描显示，大多数小泡是空的，但剩余的碳质物质浸渍产生了小泡角质层的碎片，以及珠心角质层和大孢子膜的碎片。偶有气孔，偶有于茎梗的外表皮，单倍体，有5-8个附属细胞。大孢子膜是两层的，包括一个足层和分枝的、相互连接的杆状体。北方Jarudia borealis sp. 11 .的有双苞的球果与许多被称为Phoenicopsis arcticus sp. nov.的线状叶片的遗迹以及Pityophyllum sp.和Ginkgoites sp.的罕见叶片碎片密切相关。

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