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First report of Hirnantian (Late Ordovician) palaeostomate bryozoans and associated biota of the northern Arabian Plate in southeastern Türkiye 土耳其东南部阿拉伯板块北部Hirnantian(晚奥陶世)古口苔藓虫及其伴生生物群首次报道
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-09-18 DOI: 10.1016/j.palwor.2025.201026
Andrej Ernst , İzzet Hoşgör , Olev Vinn
The Halevikdere Formation of southern Türkiye yields an under-explored archive of Late Ordovician marine ecosystems. Our knowledge about Ordovician bryozoans in Türkiye is restricted due to the poor preservation of material. The first report of the Hirnantian high-latitude north Gondwanan bryozoans from southeastern Türkiye (Mardin-Derik area, Halevikdere Formation) is presented here. The Late Ordovician (Sandbian–Hirnantian) predominantly fine-grained succession of Derik is among the most remarkable Early Palaeozoic successions in southeastern Türkiye, but its precise geological age is poorly constrained. New bryozoan-rich sequence of glacio-marine related deposits contains indeterminate crinoids and brachiopods. Here we describe a new fauna of bryozoans from the ice-distal glaciomarine succession of the Halevikdere Formation, corresponding to the northern margin of the Arabian Plate. Three palaeostomate bryozoan species were identified: the esthonioporate Esthonioporata sp. indet., cryptostomes Graptodictya sp., and Ptilodictyina sp. indet. (aff. Proavella proava). In this study, a new trace fossil assemblage comprising several ichnotaxa is also documented from the upper part of the bryozoan-bearing sequence of the Hirnantian Halevikdere Formation. The identified bryozoans are related to the typical Ordovician taxa, but their presence in the assemblage, previously reported only from the Upper Ordovician of the Baltic region, is noticeable. These bryozoans and ichnofossils are typically associated with shallow marine environments, reflecting the conditions that prevailed during the time of their formation.
rkiye南部的Halevikdere组提供了一个未被探索的晚奥陶世海洋生态系统档案。由于材料保存较差,我们对基耶省奥陶系苔藓虫的认识有限。本文首次报道了在 rkiye东南部(Mardin-Derik地区,Halevikdere组)发现的Hirnantian高纬度北部Gondwanan苔藓虫。晚奥陶世(Sandbian-Hirnantian)是基耶耶东南地区最显著的早古生代演替之一,但其确切的地质年代尚不清楚。新的富含苔藓虫的冰川期海洋相关沉积物序列包含不确定的海贝壳类和腕足类。在这里,我们描述了一种新的苔藓虫动物群,它们来自于与阿拉伯板块北缘相对应的Halevikdere组的冰-远端冰川海洋演替。鉴定出3种古口苔藓虫:estoniopate esthoniopata sp. indet;、隐口虫、隐口虫和隐口虫。(aff. Proavella proava)。本研究还在Hirnantian Halevikdere组苔藓虫序列的上半部分记录了一个新的由几个鱼类群组成的微量化石组合。发现的苔藓虫与典型的奥陶系分类群有关,但它们的存在是值得注意的,以前只报道过来自波罗的海地区的上奥陶系。这些苔藓虫和鱼类化石通常与浅海环境有关,反映了它们形成时的普遍条件。
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引用次数: 0
Palynological succession in Gzhelian No. 8 Thick Coal of the Taiyuan Formation, Ordos Basin, North China 鄂尔多斯盆地太原组格家连8号厚煤孢粉演替
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-09-18 DOI: 10.1016/j.palwor.2025.201025
Hui Zhang , Yun-He Shi , Peng-Shuai Jiao , Bo Pan , Feng Liu , Ming-Li Wan , Jing-Kun Zhang , Hui-Ping Peng
The No. 8 Thick Coal in the Taiyuan Formation, correlated with the Gzhelian (Pennsylvanian, Carboniferous) stage, extends throughout the Ordos Basin and reaches up to 30 m in thickness according to basin-wide seismic profiles analysis. This thick coal bed was formed as long-residence histosols, primarily in ombrotrophic mires with minor development in rheotrophic mires. Analysis of miospore taxa percentages across the thick coal seams identified three consistent bulk characterizing species (bcs): Torispora spp., Laevigatosporites spp., and Florinites spp. These species occur in predictable sequences throughout all No. 8 Thick Coals at both the Palougou outcrop and the Huo10 Borehole. Detailed examination revealed three distinct palynomorph associations in the No. 8 Thick Coal from the Huo10 Borehole: Latosporites globosus Association at the base, Laevigatosporites spp. Association in the middle, and Florinites spp. Association near the top. These associations share similar taxa with the Striatosporites clatratus Biozone and Pachetisporites kaipingensis Biozone previously identified in Carboniferous–Permian transitional deposits at the Palougou Section. This similarity confirms that the No. 8 Thick Coal was deposited simultaneously across the northern Ordos Basin and maintains a consistent, widespread distribution. The widespread presence of trilete and monolete miospores throughout the No. 8 Thick Coal suggests that peat accumulation kept pace with basin subsidence during the Gzhelian in the Ordos Basin. This balance created stable soil conditions that allowed tree ferns to form thick coal layers with consistent bcs. The bcs model in No. 8 Thick Coal differs from the previous Euramerican models in two key aspects: it exhibits prolonged dominance of trilete and monolete miospores and lacks both Lycospora and Densosporites spp. bcs at the coal base and roof. These differences reflect the distinct climate and vegetation of the Cathaysian Palaeofloristic Province during the Pennsylvanian (late Carboniferous), where Psaronius dominated the peat-forming swamp with minor Cordaites presence along its margins.
根据全盆地地震剖面分析,太原组8号厚煤与格芝连(宾夕法尼亚石炭系)相对应,延伸至整个盆地,厚度可达30 m。这种厚煤层是作为长期存在的组织溶胶形成的,主要是在营养型煤中形成的,在流变型煤中也有少量发育。通过对厚煤层中微孢子类群百分比的分析,确定了三个一致的块状特征物种(bcs): Torispora spp、Laevigatosporites spp和Florinites spp,这些物种在Palougou露头和Huo10钻孔的所有8号厚煤中均以可预测的顺序出现。火10井8号厚煤中有3个明显的孢型组合:底部的Latosporites globosus Association,中部的Laevigatosporites spp. Association,顶部附近的Florinites spp. Association。这些组合与先前在帕洛沟剖面石炭-二叠纪过渡沉积中发现的Striatosporites clatratus Biozone和Pachetisporites kaipingensis Biozone具有相似的分类群。这种相似性证实了8号厚煤在鄂尔多斯盆地北部同时沉积,且分布一致、广泛。鄂尔多斯盆地8号厚煤中普遍存在三联体和单联体小孢子,表明盆地格日连期泥炭成藏与盆地沉降同步。这种平衡创造了稳定的土壤条件,使蕨类植物能够形成具有一致bcs的厚煤层。8号厚煤的bcs模式与以前的欧美模式有两个关键的不同:它在煤底和煤顶表现出三孢子和单孢子的长期优势,缺乏Lycospora和Densosporites两种bcs。这些差异反映了宾夕法尼亚纪(晚石炭世)华夏古植物区省独特的气候和植被,在那里,Psaronius占泥炭形成沼泽的主导地位,Cordaites在其边缘存在少量。
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引用次数: 0
Multi-proxy climatic reconstruction of the Siwalik (Middle Miocene) Balason River area, Darjeeling, eastern Himalaya 喜马拉雅东部大吉岭中中新世巴拉松河地区Siwalik(中中新世)多指标气候重建
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-09-17 DOI: 10.1016/j.palwor.2025.201017
Sumana Mahato , Robert A. Spicer , Sandip More , Ai Song , Mahasin Ali Khan
No single palaeoclimate proxy is perfect, so a multiproxy approach is always desirable for reconstructing past environments. Here, using the quantitative methods known as the Coexistence Approach, and Climate Leaf Analysis Multivariate Program (CLAMP), along with qualitative assessments based on the nearest living relative, leaf cuticle analysis, palynology, and herbivory reconstructions, we explore the palaeoclimate of the Middle Miocene Darjeeling (Balason) Siwalik, eastern Himalaya. This multiproxy intercomparison provides an important cross-validation of qualitative and quantitative climate proxies in an Indian Cenozoic context. We find that all proxies give similar palaeoclimate outcomes and show that the Balason location experienced a tropical, warm, humid climate during the Middle Miocene Siwalik deposition.
没有单一的古气候代理是完美的,所以多代理方法总是重建过去环境的理想方法。本文采用共存法和气候叶片分析多元程序(CLAMP)等定量方法,以及基于近缘生物、叶片角质层分析、孢粉学和草食重建的定性评估,对喜马拉雅东部中新世中吉岭(Balason) Siwalik的古气候进行了研究。这种多代理相互比较为印度新生代气候代理的定性和定量提供了重要的交叉验证。我们发现所有的代用指标都给出了相似的古气候结果,并表明Balason地区在中中新世Siwalik沉积时期经历了热带、温暖、潮湿的气候。
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引用次数: 0
A new genus of Neoliodidae (Acariformes: Oribatida) from Early Cretaceous Lebanese amber and Azarliodes andreneli n. comb. 文章标题早白垩世黎巴嫩琥珀和阿扎利亚蛛蛛一新属(鞘翅目:翅虫目)。
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-09-15 DOI: 10.1016/j.palwor.2025.201016
Antonio Arillo, Mónica Gutiérrez
Neoliodidae is a family of oribatid mites including four extant genera with a worldwide distribution from temperate to tropical environments. It is considered the sister group of the rest of the Brachypilina. A new genus, Azarliodes n. gen., is proposed to include the species Neoliodes andreneli from the Early Cretaceous Lebanese amber, as Azarliodes andreneli n. comb., based on a new specimen of the same species and the re-examination of the holotype. The new genus preserves both rostral and lamellar setae. Accordingly, the loss of a pair of prodorsal setae in Neoliodes and Poroliodes, considered to be the lamellar ones, might have been a misinterpretation and the lost pair of setae could correspond in fact to the rostral ones. Furthermore, the taxon Neoliodini n. parvorder is erected to include the superfamily Neoliodoidea, considering the rest of the Brachypylina as a monophyletic group.
新甲螨科是甲螨科的一个科,包括现存的四个属,分布在温带至热带环境。它被认为是Brachypilina其余部分的姐妹组。提出了一个新属,Azarliodes n. gen.,包括来自早白垩世黎巴嫩琥珀的Neoliodes andreneli,作为Azarliodes andreneli n. comb。,根据同一物种的新标本和重新检查的全型。新属保留吻侧和片层刚毛。因此,在新科和孔科中被认为是片层的一对前卧刚毛的丢失可能是一种误解,而丢失的一对刚毛实际上可能对应于吻侧刚毛。此外,还建立了一个分类单元Neoliodini n. parvorder,包括了超科neoliodo总科,将其余的Brachypylina视为一个单系类群。
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引用次数: 0
Lebanese amber: A time capsule from the dawn of modern ecosystems 黎巴嫩琥珀:现代生态系统的时间胶囊
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-09-15 DOI: 10.1016/j.palwor.2025.201015
Sibelle Maksoud , Haig Tabakian
Lebanon has played a pivotal role in the advancement of palaeoentomology and the broader understanding of insect evolution and palaeobiodiversity. This contribution is evidenced by its 39 Cretaceous fossiliferous outcrops, encompassing 32 Early Cretaceous amber localities and several lithological sites (including marl, limestone, and dysodile) bearing fossil insects, in addition to a Jurassic amber occurrence. Despite its relatively small geographic extent, Lebanon constitutes a major palaeohotspot for Mesozoic entomofauna. Its distinctive geological evolution and palaeogeographic setting during the Mesozoic Era have rendered it a region of exceptional continental palaeontological significance. In this study, we provide a revised simplified geological section of the Late Jurassic and Early Cretaceous amber-bearing deposits containing biological inclusions, along with a comprehensive catalogue of all fossil insects and associated biological inclusions described to date from Lebanese amber.
黎巴嫩在古昆虫学的进步和对昆虫进化和古生物多样性的更广泛理解方面发挥了关键作用。这一贡献可以通过其39个白垩纪化石露头得到证明,其中包括32个早白垩纪琥珀地点和几个带有化石昆虫的岩性地点(包括泥灰岩、石灰石和异质岩),此外还有一个侏罗纪琥珀产地。尽管黎巴嫩的地理范围相对较小,但它构成了中生代昆虫动物的主要古热点。其独特的中生代地质演化和古地理环境使其成为一个具有特殊大陆古生物意义的地区。在这项研究中,我们提供了晚侏罗世和早白垩世含有生物包裹体的琥珀矿床的修订简化地质剖面,以及迄今为止从黎巴嫩琥珀中描述的所有化石昆虫和相关生物包裹体的综合目录。
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引用次数: 0
Psychodid flies (Diptera: Psychodidae) from the Lower Cretaceous Lebanese amber 下白垩纪黎巴嫩琥珀中的精神病蝇(双翅目:精神病蝇科)
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-09-15 DOI: 10.1016/j.palwor.2025.201014
Haig Tabakian
Psychodid flies had a great diversity in Early Cretaceous and were remarkably preserved in the lower Barremian amber of Lebanon. The first two species from Lebanese amber were described by Hennig in 1972, and 20 more species have been added later to the family during the past two decades. In this paper, a review of all the described Psychodidae from the Lebanese amber and an identification key for determination are proposed.
精神病蝇在早白垩纪有很大的多样性,并在黎巴嫩的巴雷米亚琥珀中得到了显著的保存。黎巴嫩琥珀的前两个物种是由Hennig在1972年描述的,在过去的20年里,又有20个物种被添加到这个家族中。本文综述了黎巴嫩琥珀中所描述的所有精神病科,并提出了鉴定的关键字。
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引用次数: 0
Siliceous sponge fossils from the early Cambrian Qingxi Formation in South China 华南早寒武世青溪组硅质海绵化石
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-09-11 DOI: 10.1016/j.palwor.2025.201013
Xin-Yuan Cao , Hai-Zhou Wang , Wen-Zhe Liu , Jia-Yue Wang , Jun-Ying Ding , Guo-Guang Wang , Qing Tang
The emergence of biomineralized sponge spicules in the early Cambrian provides critical insights into the origin of spiculogenesis and animal biomineralization. While sponge spicules proliferated in the early Cambrian, their original chemical composition is rarely preserved due to diagenetic alterations, hampering understanding on the mineralogy of the early spicules. Here we report abundant disarticulated fossil spicules from the early Cambrian Qingxi Formation in northern Guangxi Zhuang Autonomous Region (hereafter referred to as Guangxi), South China. Using reflected light microscopy, scanning electron microscopy (SEM), energy-dispersive X-ray spectroscopy (EDS), and Raman spectroscopy, our study reveals that the Qingxi spicules are predominately derived from Hexactinellida and are preserved in various modes, including pyritization, mold, and silica preservation. Notably, many spicules in silica preservation are composed of microcrystalline quartz with conchoidal fractures in cross-sectional view, suggesting minimal diagenetic alteration and retention of original chemical compositions. This study represents the first systematic investigation on the paleontological record of the Qingxi Formation in Guangxi, highlighting the exceptional potential of Cambrian shales and mudstones to preserve sponge spicules with primary silica composition.
早寒武纪生物矿化海绵针状体的出现为针状体形成和动物生物矿化的起源提供了重要的认识。虽然海绵针状体在早寒武纪大量出现,但由于成岩作用的改变,其原始化学成分很少被保存下来,阻碍了对早期针状体矿物学的认识。本文报道了在中国南方广西壮族自治区北部(以下简称广西)发现的大量早寒武世青溪组分离的化石针状体。利用反射光显微镜、扫描电镜(SEM)、能量色散x射线能谱(EDS)和拉曼光谱分析表明,清溪针状体主要来源于Hexactinellida,并以黄铁矿、霉菌和二氧化硅等多种保存方式保存。值得注意的是,在石英保存中,许多针状体是由微晶石英组成的,在剖面上具有贝壳状裂缝,表明成岩蚀变很小,原始化学成分保留。本研究首次对广西清溪组的古生物记录进行了系统的研究,强调了寒武系页岩和泥岩保存原生硅质海绵针状体的特殊潜力。
{"title":"Siliceous sponge fossils from the early Cambrian Qingxi Formation in South China","authors":"Xin-Yuan Cao ,&nbsp;Hai-Zhou Wang ,&nbsp;Wen-Zhe Liu ,&nbsp;Jia-Yue Wang ,&nbsp;Jun-Ying Ding ,&nbsp;Guo-Guang Wang ,&nbsp;Qing Tang","doi":"10.1016/j.palwor.2025.201013","DOIUrl":"10.1016/j.palwor.2025.201013","url":null,"abstract":"<div><div>The emergence of biomineralized sponge spicules in the early Cambrian provides critical insights into the origin of spiculogenesis and animal biomineralization. While sponge spicules proliferated in the early Cambrian, their original chemical composition is rarely preserved due to diagenetic alterations, hampering understanding on the mineralogy of the early spicules. Here we report abundant disarticulated fossil spicules from the early Cambrian Qingxi Formation in northern Guangxi Zhuang Autonomous Region (hereafter referred to as Guangxi), South China. Using reflected light microscopy, scanning electron microscopy (SEM), energy-dispersive X-ray spectroscopy (EDS), and Raman spectroscopy, our study reveals that the Qingxi spicules are predominately derived from Hexactinellida and are preserved in various modes, including pyritization, mold, and silica preservation. Notably, many spicules in silica preservation are composed of microcrystalline quartz with conchoidal fractures in cross-sectional view, suggesting minimal diagenetic alteration and retention of original chemical compositions. This study represents the first systematic investigation on the paleontological record of the Qingxi Formation in Guangxi, highlighting the exceptional potential of Cambrian shales and mudstones to preserve sponge spicules with primary silica composition.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"35 1","pages":"Article 201013"},"PeriodicalIF":1.7,"publicationDate":"2025-09-11","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145929109","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
引用次数: 0
A new early Barremian fossiliferous amber outcrop at Beqaa Kafra (Northern Lebanon) Beqaa Kafra(黎巴嫩北部)新发现的早期巴雷米亚化石琥珀露头
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-09-10 DOI: 10.1016/j.palwor.2025.201012
Dany Azar , Eduardo Barrón , Denise Iskandar-Tabib , Sibelle Maksoud
A new outcrop of fossiliferous amber discovered at a site in the village of Beqaa Kafra (Bcharreh District, Northern Lebanon) during recent geological surveys, is described herein. This site represents the 32nd known Lebanese amber outcrop containing biological inclusions. A palynological study was conducted to investigate the botanical diversity of the surrounding palaeoenvironment. Fourier Transform Infrared spectroscopy was performed on amber samples from the newly discovered site to depict their chemical composition and evaluate potential correlations with other Lebanese amber outcrops. The discovery significantly contributes to our understanding of Early Cretaceous palaeobiodiversity along the northeastern margin of Gondwana, particularly during the early Barremian.
在最近的地质调查中,在Beqaa Kafra村(Bcharreh区,黎巴嫩北部)的一个地点发现了一个新的化石琥珀露头。该遗址是已知的第32个含有生物包裹体的黎巴嫩琥珀露头。通过孢粉学研究,探讨了其周围古环境的植物多样性。对新发现地点的琥珀样品进行了傅里叶变换红外光谱分析,以描述其化学成分,并评估与其他黎巴嫩琥珀露头的潜在相关性。这一发现对我们了解冈瓦纳东北缘早白垩世古生物多样性,特别是巴雷米亚早期的生物多样性有重要意义。
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引用次数: 0
New and revised middle Villafranchian canid material from Dafnero-3 (Greece): Implications to Nyctereutes systematics and zoogeography 希腊Dafnero-3中Villafranchian犬科动物资料的新修订:对Nyctereutes系统学和动物地理学的启示
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-09-04 DOI: 10.1016/j.palwor.2025.201011
Anna Savvidou, Dimitris S. Kostopoulos
New cranial, mandibular and dental canid material from Dafnero-3 site, in northern Greece confirms previous reports of Nyctereutes megamastoides and Vulpes sp. from either Dafnero-3 or/and Dafnero-1 sites but allows a reassignment of specimens previously referred to as Nyctereutes tingi from Dafnero-3 to Nyctereutes vulpinus. The results further support the previous hypothesis of a close phylogenetic link between N. tingi and N. vulpinus, revise the intraspecific diversity of the latter, and allow the establishment of a working phylogeographic hypothesis. Nyctereutes vulpinus is noted for the first time in southeast Europe, expanding its geographical range, confirming the simultaneous presence of two distinct Nyctereutes species in the Balkan area as in China.
来自希腊北部Dafnero-3遗址的新颅、下颌和牙类材料证实了之前在Dafnero-3或/和Dafnero-1遗址发现的巨齿Nyctereutes megamastoides和Vulpes,但允许将以前被称为tingnyctereutes tingi的标本从Dafnero-3重新分配到vulpinus Nyctereutes。该研究结果进一步支持了先前关于刺毛螨与狐毛螨之间存在密切系统发育联系的假设,修正了后者的种内多样性,并允许建立一个有效的系统地理学假设。在欧洲东南部首次发现夜蛾,扩大了其地理分布范围,证实了在巴尔干地区和中国同时存在两种不同的夜蛾。
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引用次数: 0
Middle–late Cambrian acritarchs from the Llanos Basin in Colombia: Implications for biostratigraphy and palaeobiogeography 哥伦比亚Llanos盆地中晚寒武世树栖类:生物地层学和古生物地理学意义
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-09-04 DOI: 10.1016/j.palwor.2025.201009
Claudia V. Rubinstein , María C. Vargas , Felipe de la Parra
A palynological investigation of a Cambrian sedimentary succession drilled in well A, located in the Llanos Basin of Colombia, revealed moderately preserved and relatively diverse acritarchs. This analysis allows for the identification of two distinct acritarch assemblages. Based on the presence of key diagnostic species and correlation with established middle–late Cambrian acritarch biozones, the lower acritarch assemblage is dated to the middle Cambrian, corresponding to the Drumian–Guzhangian stages of the Miaolingian Series. The upper acritarch assemblage is dated as late Cambrian, corresponding to the Jiangshanian–Stage 10 of the Furongian Series on the Global Chronostratigraphic Scale. The presence of globally-occurring species such as Eliasum llaniscum, Cristalinium cambriense, Ninadiacrodium dumontii, Timofeevia phosphoritica, Vulcanisphaera mougnoana, and V. africana supports the cosmopolitanism of middle and late Cambrian acritarchs. The identification of middle and upper Cambrian strata in the Llanos Basin’s Palaeozoic sequence helps to establish a more accurate biostratigraphic framework for the Palaeozoic of Colombia.
对哥伦比亚Llanos盆地A井寒武系沉积演替进行的孢粉学研究显示,该地层中存在保存较好且种类相对丰富的关键种。这种分析允许识别两种不同的主柱头组合。根据关键诊断种的存在,并与已建立的中晚寒武统树栖生物带进行对比,下树栖生物组合的年代为中寒武统,对应于庙岭系的鼓店—古张期。上支岩组合为晚寒武世,对应于全球年代地层尺度上的芙蓉系江山期—第10期。Eliasum llaniscum、Cristalinium cambriense、Ninadiacrodium dumontii、Timofeevia phosphoritica、Vulcanisphaera mougnoana和V. africana等全球存在的物种的存在支持了中晚寒武纪树栖生物的世界主义。通过对Llanos盆地古生代层序中、上寒武统地层的识别,有助于建立更为准确的哥伦比亚古生代生物地层格架。
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引用次数: 0
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