Pub Date : 2025-05-26DOI: 10.1016/j.palwor.2025.200964
Wu-Jun Xu , Zuo-Yu Sun , Shuang Liu , Cheng Ji , Hao Lu , Zhao-Liang Ma , Fei-Yang Wu , Shou-Ming Chen , Jian-Dong Huang
The apparatus of the coniform conodont genus Cornudina remains equivocal because the previous results were derived from discrete element collections rather than natural assemblages. Four exceptionally preserved natural assemblages collected from bed 719 of the upper Nanlinghu Formation in Majiashan quarry, Chaohu of Hefei City, Anhui Province, East China, were imaged using high-resolution X-ray microtomography. The consistent and repeated patterns of element juxtaposition suggest an apparatus of 15 elements, conforming to the classical 2M-9S-4P skeletal template that comprises three basic morphologies: two breviform dolabrate elements, nine elongate bipennate elements, and four angulate elements. The angulate P1 and P2 elements have a distinct cusp, short anterior and posterior processes and the P2 element is twisted crescent-shaped, which supports a taxonomic assignment to Cornudina breviramulis sensu stricto Koike. Multielement comparisons among Cornudina, Neostrachanognathus, and Hindeodus apparatuses suggest Cornudina and Neostrachanognathus are closely related and possibly derived from Hindeodus. The multi-element comparison between Ordovician ‘complex’ conodont apparatuses (e.g., Paracordylodus and Phragmodus) and Triassic ‘coniform’ apparatuses (e.g., Cornudina and Neostrachanognathus) suggests that the low variation in the elemental forms of the latter is probably a secondary ‘evolution’.
锥形牙形刺属的装置仍然是模棱两可的,因为以前的结果是来自离散元素集合而不是自然组合。利用高分辨率x射线显微层析成像技术,对安徽省合肥市巢湖马家山采石场南陵湖上组719层采集的4个保存特别完好的自然组合进行了成像。元素并置的一致和重复的模式表明了一个由15个元素组成的装置,符合经典的2M-9S-4P骨架模板,包括三种基本形态:两个短形dolabate元素,九个细长的bipenate元素和四个角形元素。P1和P2呈角状,尖部明显,前后突短,P2呈弯月牙状,属于corudina breviramulis sensu stricto Koike。Cornudina、Neostrachanognathus和Hindeodus器械的多元素比较表明Cornudina和Neostrachanognathus亲缘关系密切,可能来源于Hindeodus。奥陶系“复杂”牙形器(如Paracordylodus和Phragmodus)与三叠系“形形”器(如Cornudina和Neostrachanognathus)的多元素比较表明,后者元素形式的低变化可能是二次“进化”。
{"title":"Multielement conodont apparatus (Cornudina) from the Lower Triassic of China, revealed by high-resolution X-ray microtomography","authors":"Wu-Jun Xu , Zuo-Yu Sun , Shuang Liu , Cheng Ji , Hao Lu , Zhao-Liang Ma , Fei-Yang Wu , Shou-Ming Chen , Jian-Dong Huang","doi":"10.1016/j.palwor.2025.200964","DOIUrl":"10.1016/j.palwor.2025.200964","url":null,"abstract":"<div><div>The apparatus of the coniform conodont genus <em>Cornudina</em> remains equivocal because the previous results were derived from discrete element collections rather than natural assemblages. Four exceptionally preserved natural assemblages collected from bed 719 of the upper Nanlinghu Formation in Majiashan quarry, Chaohu of Hefei City, Anhui Province, East China, were imaged using high-resolution X-ray microtomography. The consistent and repeated patterns of element juxtaposition suggest an apparatus of 15 elements, conforming to the classical 2M-9S-4P skeletal template that comprises three basic morphologies: two breviform dolabrate elements, nine elongate bipennate elements, and four angulate elements. The angulate P<sub>1</sub> and P<sub>2</sub> elements have a distinct cusp, short anterior and posterior processes and the P<sub>2</sub> element is twisted crescent-shaped, which supports a taxonomic assignment to <em>Cornudina breviramulis</em> sensu stricto Koike. Multielement comparisons among <em>Cornudina</em>, <em>Neostrachanognathus</em>, and <em>Hindeodus</em> apparatuses suggest <em>Cornudina</em> and <em>Neostrachanognathus</em> are closely related and possibly derived from <em>Hindeodus</em>. The multi-element comparison between Ordovician ‘complex’ conodont apparatuses (e.g., <em>Paracordylodus</em> and <em>Phragmodus</em>) and Triassic ‘coniform’ apparatuses (e.g., <em>Cornudina</em> and <em>Neostrachanognathus</em>) suggests that the low variation in the elemental forms of the latter is probably a secondary ‘evolution’.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"34 5","pages":"Article 200964"},"PeriodicalIF":1.7,"publicationDate":"2025-05-26","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145018784","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-05-20DOI: 10.1016/j.palwor.2025.200962
Paola Zuliani , Alexandra Crisafulli , Ricardo Melchor , Laura Vallejos Leiz
The Carapacha Formation consists of two members: the Calencó (lower) and the Urre-Lauquen (upper), dated as Permian based on its glossopterid flora. The formation outcrops in La Pampa Province, near the locality of Puelches, with sediments of continental origin, fluvio-lacustrine in type. This study describes two gymnosperm woods species. One, from the Calencó Member, is a Cordaitalean assigned to Scleromedulloxylon aveyronense found in the Permian of Aveyron, France, characterized by having a heterocellular pith with islands of sclerenchymatic cells and unorganized lacunae in transverse section. The other specimen, from the Urre-Lauquen Member, is a conifer of uncertain affinity, Zalesskioxylon uniseriatum, previously reported from the Raniganj (India), Yaguarí (Uruguay), and Tacuary (Paraguay) formations. Both woods exhibit secondary xylem features typical of Paleozoic gymnosperms, particularly concerning the radial system and the arrangement of pits in the tracheid walls. These xylological records confirm that the Carapacha Formation comprises another Permian sedimentological sequences that contain woods with diaphragm-solenoid pith such as the taxa found in the Melo (Uruguay), the Río Bonito, and the Irati (Brazil), the White Band (Africa), the Barakar, and the Raniganj (India) formations. This palaeofloristic association is part of the Australoxylon xylotaphoflora characteristic of the Gondwanan Permian. Finally, this study allows us to infer that the environmental conditions in this formation were warm with low humidity and seasonally dry.
{"title":"New findings in the Permian xyloflora of Carapacha Formation (Argentina)","authors":"Paola Zuliani , Alexandra Crisafulli , Ricardo Melchor , Laura Vallejos Leiz","doi":"10.1016/j.palwor.2025.200962","DOIUrl":"10.1016/j.palwor.2025.200962","url":null,"abstract":"<div><div>The Carapacha Formation consists of two members: the Calencó (lower) and the Urre-Lauquen (upper), dated as Permian based on its glossopterid flora. The formation outcrops in La Pampa Province, near the locality of Puelches, with sediments of continental origin, fluvio-lacustrine in type. This study describes two gymnosperm woods species. One, from the Calencó Member, is a Cordaitalean assigned to <em>Scleromedulloxylon aveyronense</em> found in the Permian of Aveyron, France, characterized by having a heterocellular pith with islands of sclerenchymatic cells and unorganized lacunae in transverse section. The other specimen, from the Urre-Lauquen Member, is a conifer of uncertain affinity, <em>Zalesskioxylon uniseriatum</em>, previously reported from the Raniganj (India), Yaguarí (Uruguay), and Tacuary (Paraguay) formations. Both woods exhibit secondary xylem features typical of Paleozoic gymnosperms, particularly concerning the radial system and the arrangement of pits in the tracheid walls. These xylological records confirm that the Carapacha Formation comprises another Permian sedimentological sequences that contain woods with diaphragm-solenoid pith such as the taxa found in the Melo (Uruguay), the Río Bonito, and the Irati (Brazil), the White Band (Africa), the Barakar, and the Raniganj (India) formations. This palaeofloristic association is part of the <em>Australoxylon</em> xylotaphoflora characteristic of the Gondwanan Permian. Finally, this study allows us to infer that the environmental conditions in this formation were warm with low humidity and seasonally dry.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"34 5","pages":"Article 200962"},"PeriodicalIF":1.7,"publicationDate":"2025-05-20","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145018780","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-05-20DOI: 10.1016/j.palwor.2025.200959
Qian-Qi Zhang , Lin Mu , Lei Zhang , Xu-Heng Du , Hai-Chun Zhang
As the earliest record of the family Curvicubitidae Hong, 1984, Sinocurvicubitus qingjiangensis n. gen. n. sp. is established based on a cicada tegmen collected from the Laoshan Member of the Leping Formation in the Qingjiang Basin, Jiangxi Province, southern China. This new taxon is characterised by vein R forked at basal 1/3rd tegminal length, indentation at basal 2/3rds length, vein M first forked on nodal line, more developed M and less developed CuA. A Wuchiapingian age for this new species is suggested by the co-occurrence of the ammonoids Araxoceras tricarinatus and Pseudogastrioceras sp. The new find not only extends the known palaeogeographical range of curvicubitids in China, but also suggests an origin of this family in the Palaeozoic. This new record, combined with other records of Curvicubitidae, suggests a possibility of simultaneously northward and southward migration and dispersal of this family during the late Permian to the Late Triassic.
Sinocurvicubitus qingjiangensis n. gen. n. sp.根据江西清江盆地乐平组崂山段的蝉甲,于1984年建立,是curvicutidae科最早记录。该新分类群的特征为脉R在基部1/3顶尖处分叉,脉M在基部2/3处凹陷,脉M在节线上先分叉,脉M较发达,脉CuA较不发达。这一新发现不仅扩大了曲丘类动物在中国已知的古地理分布范围,而且表明曲丘类动物起源于古生代。这一新的记录,结合其他的曲丘虫科记录,表明该科在晚二叠世至晚三叠世期间可能同时向北和向南迁移和分散。
{"title":"The oldest known Curvicubitidae (Hemiptera) from the Permian (Wuchiapingian) of Zhangshu City, Jiangxi Province, southern China","authors":"Qian-Qi Zhang , Lin Mu , Lei Zhang , Xu-Heng Du , Hai-Chun Zhang","doi":"10.1016/j.palwor.2025.200959","DOIUrl":"10.1016/j.palwor.2025.200959","url":null,"abstract":"<div><div>As the earliest record of the family Curvicubitidae <span><span>Hong, 1984</span></span>, Sinocurvicubitus qingjiangensis n. gen. n. sp. is established based on a cicada tegmen collected from the Laoshan Member of the Leping Formation in the Qingjiang Basin, Jiangxi Province, southern China. This new taxon is characterised by vein R forked at basal 1/3rd tegminal length, indentation at basal 2/3rds length, vein M first forked on nodal line, more developed M and less developed CuA. A Wuchiapingian age for this new species is suggested by the co-occurrence of the ammonoids <em>Araxoceras tricarinatus</em> and <em>Pseudogastrioceras</em> sp. The new find not only extends the known palaeogeographical range of curvicubitids in China, but also suggests an origin of this family in the Palaeozoic. This new record, combined with other records of Curvicubitidae, suggests a possibility of simultaneously northward and southward migration and dispersal of this family during the late Permian to the Late Triassic.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"34 5","pages":"Article 200959"},"PeriodicalIF":1.7,"publicationDate":"2025-05-20","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144170129","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
At Tagogapu in West Java, the Miogypsinidae larger benthic foraminifera from the Rajamandala Limestone have been studied for biometric trends in evolution. Observed taxa include Neorotalia mecatepecensis, Paleomiogypsina sp., Miogypsinella bornea, Miogypsinoides complanatus, and Miogypsinoides formosensis, with biometric parameters (X, Y, Z, AX, and γ) revealing distinct phylogenetic trends through heterochrony. Neorotalia mecatepecensis morphotype A is identified as the basal species, while Neorotalia mecatepecensis morphotype B diverges through nepionic retardation. The transition to Paleomiogypsina sp. is marked by the development of auxiliary chambers. The transition to Miogypsinella bornea is suggested by the addition of secondary foramen and more auxiliary chambers. The evolution to Miogypsinoides complanatus is characterized by the planispiral nepionic, loss of an apertural lip, and broader auxiliary chamber arrangements. The speciation of Miogypsinoides formosensis is recognized by the peramorphic addition of late adult stage auxiliary chambers. Biostratigraphic analysis and strontium isotopic dating place the studied taxa within the Te 2-3 to Te 4 Letter Stage (late Oligocene). This research highlights the biostratigraphic application of biometric methods in a lineage with heterochronous evolutionary trends, and proposes AX as a robust metric for the descendant peramorphic forms.
{"title":"Phylogenetic evolution of Neorotalia mecatepecensis to Miogypsinidae from Rajamandala Formation, West Java, Indonesia","authors":"Nadila Novandaru , Khoiril Anwar Maryunani , Rubiyanto Kapid , Aswan , Wahyu Budhi Khorniawan","doi":"10.1016/j.palwor.2025.200961","DOIUrl":"10.1016/j.palwor.2025.200961","url":null,"abstract":"<div><div>At Tagogapu in West Java, the Miogypsinidae larger benthic foraminifera from the Rajamandala Limestone have been studied for biometric trends in evolution. Observed taxa include <em>Neorotalia mecatepecensis</em>, <em>Paleomiogypsina</em> sp., <em>Miogypsinella bornea</em>, <em>Miogypsinoides complanatus</em>, and <em>Miogypsinoides formosensis</em>, with biometric parameters (X, Y, Z, AX, and γ) revealing distinct phylogenetic trends through heterochrony. <em>Neorotalia mecatepecensis</em> morphotype A is identified as the basal species, while <em>Neorotalia mecatepecensis</em> morphotype B diverges through nepionic retardation. The transition to <em>Paleomiogypsina</em> sp. is marked by the development of auxiliary chambers. The transition to <em>Miogypsinella bornea</em> is suggested by the addition of secondary foramen and more auxiliary chambers. The evolution to <em>Miogypsinoides complanatus</em> is characterized by the planispiral nepionic, loss of an apertural lip, and broader auxiliary chamber arrangements. The speciation of <em>Miogypsinoides formosensis</em> is recognized by the peramorphic addition of late adult stage auxiliary chambers. Biostratigraphic analysis and strontium isotopic dating place the studied taxa within the Te 2-3 to Te 4 Letter Stage (late Oligocene). This research highlights the biostratigraphic application of biometric methods in a lineage with heterochronous evolutionary trends, and proposes AX as a robust metric for the descendant peramorphic forms.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"34 5","pages":"Article 200961"},"PeriodicalIF":1.7,"publicationDate":"2025-05-20","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145018792","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-05-20DOI: 10.1016/j.palwor.2025.200960
Shi-Qi Wang , Xiao-Yong Zhang , Yi-Kun Li , Xiao-Xiao Zhang , Qi-Gao Jiangzuo
Fossils of Antilopina were often referred to and classified under the genus name “Gazella”, except for some Miocene spiral horned antelopes; however, their true taxonomic attribution is difficult to determine due to the significant morphological variation observed in large fossil samples. Species of two extant genera within Antilopina that are widely distributed in China are Gazella subgutturosa and Procapra spp. Assigning abundant fossil specimens identified as “Gazella” from China to these two extant groups is particularly challenging. Molecular phylogenetic studies have revealed that Gazella and Procapra belong to two distinct clades within Antilopina. Therefore, the study of fossil “Gazella” is crucial for understanding the differentiation and evolutionary history of Antilopina, which includes a diverse array of genera that lived in Eurasia and Africa. In this article, we describe a newly discovered cranium of “Gazella” nihensis from the Pliocene Epoch on the Qinghai-Tibetan Plateau. The new cranium exhibits several distinctive features: the braincase is considerably elongated, with the occipital condyles protruding caudally beyond the paroccipital process; the cranial-facial axis is weakly curved, and the horn cores are strongly inclined caudally. The morphology is distinct from most other members of Antilopina. We testified the view that “Gazella” nihensis is possibly related with the living Procapra because of these specialized features. Interestingly, in “Gazella” nihensis, the elongated braincase that is less bent from the facial axis resembles that of the living Litocranius walleri; and the strongly caudally inclined horn core that of the living Ammodorcas clarkei and Antidorcas marsupialis. These similarities may be the result of parallel evolution, but further research is needed to explore this interesting resemblance.
{"title":"A cranium of “Gazella” nihensis from Pliocene of Qinghai-Tibetan Plateau and the differentiation of early Antilopina","authors":"Shi-Qi Wang , Xiao-Yong Zhang , Yi-Kun Li , Xiao-Xiao Zhang , Qi-Gao Jiangzuo","doi":"10.1016/j.palwor.2025.200960","DOIUrl":"10.1016/j.palwor.2025.200960","url":null,"abstract":"<div><div>Fossils of Antilopina were often referred to and classified under the genus name “<em>Gazella</em>”, except for some Miocene spiral horned antelopes; however, their true taxonomic attribution is difficult to determine due to the significant morphological variation observed in large fossil samples. Species of two extant genera within Antilopina that are widely distributed in China are <em>Gazella subgutturosa</em> and <em>Procapra</em> spp. Assigning abundant fossil specimens identified as “<em>Gazella</em>” from China to these two extant groups is particularly challenging. Molecular phylogenetic studies have revealed that <em>Gazella</em> and <em>Procapra</em> belong to two distinct clades within Antilopina. Therefore, the study of fossil “<em>Gazella</em>” is crucial for understanding the differentiation and evolutionary history of Antilopina, which includes a diverse array of genera that lived in Eurasia and Africa. In this article, we describe a newly discovered cranium of “<em>Gazella</em>” <em>nihensis</em> from the Pliocene Epoch on the Qinghai-Tibetan Plateau. The new cranium exhibits several distinctive features: the braincase is considerably elongated, with the occipital condyles protruding caudally beyond the paroccipital process; the cranial-facial axis is weakly curved, and the horn cores are strongly inclined caudally. The morphology is distinct from most other members of Antilopina. We testified the view that “<em>Gazella</em>” <em>nihensis</em> is possibly related with the living <em>Procapra</em> because of these specialized features. Interestingly, in “<em>Gazella</em>” <em>nihensis</em>, the elongated braincase that is less bent from the facial axis resembles that of the living <em>Litocranius walleri</em>; and the strongly caudally inclined horn core that of the living <em>Ammodorcas clarkei</em> and <em>Antidorcas marsupialis</em>. These similarities may be the result of parallel evolution, but further research is needed to explore this interesting resemblance.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"34 5","pages":"Article 200960"},"PeriodicalIF":1.7,"publicationDate":"2025-05-20","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145018829","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-05-16DOI: 10.1016/j.palwor.2025.200958
Ke-Ying Yue , Sha Li , Tian Jiang , Dang-Peng Xi , Bo-Wen Song , Yu Liang , Li-Xin Sun , Tian-Fu Zhang , Hai-Chun Zhang , Xiao-Qiao Wan , Bo Wang
The Songliao Basin, spanning approximately 260,000 km2 in Northeast China, is one of the largest oil-bearing lacustrine basins. Seawater incursion events have previously been discovered in the Turonian (lower and the uppermost Qingshankou Formation) to the Santonian and early Campanian (the first and second members of the Nenjiang Formation). We newly discovered benthic and planktonic foraminifera in the middle Campanian to Maastrichtian (Fifth Member of the Nenjiang Formation, Sifangtai, and Mingshui formations) in the ZKD-1 borehole. The Planktonic foraminifera include Archaeoglobigerina blowi, A. cretacea, and Globotruncanella sp. The benthic foraminifera include Anomalinoides sp., Gavelinella sp., and Haplophragmoides sp. Among them, A. blowi, A. cretacea, Anomalinoides sp., Gavelinella sp., Haplophragmoides sp. are widely distributed in the marine strata of the Upper Cretaceous. We tested the marine biomarker compounds including C30 24-n-propyl cholestane and C30 24-iso-propyl cholestane, indicating that the Songliao Basin experienced seawater incursion events to varying degrees during the middle to late Campanian. We also found abundant charophyte fossils in the ZKD-1 borehole, including Atopochara trivolvis ulanensis, Latochara yuananensis, Feistiella anluensis, Mesochara biacuta, M. gobica, and M. leiocarpa. Among them, the brackish water charophyte fossils of Feistiella anluensis were first found in Songliao Basin.
{"title":"Middle Campanian to Maastrichtian Foraminifera and biomarkers: Seawater incursion events in the Songliao Basin, Northeast China","authors":"Ke-Ying Yue , Sha Li , Tian Jiang , Dang-Peng Xi , Bo-Wen Song , Yu Liang , Li-Xin Sun , Tian-Fu Zhang , Hai-Chun Zhang , Xiao-Qiao Wan , Bo Wang","doi":"10.1016/j.palwor.2025.200958","DOIUrl":"10.1016/j.palwor.2025.200958","url":null,"abstract":"<div><div>The Songliao Basin, spanning approximately 260,000 km<sup>2</sup> in Northeast China, is one of the largest oil-bearing lacustrine basins. Seawater incursion events have previously been discovered in the Turonian (lower and the uppermost Qingshankou Formation) to the Santonian and early Campanian (the first and second members of the Nenjiang Formation). We newly discovered benthic and planktonic foraminifera in the middle Campanian to Maastrichtian (Fifth Member of the Nenjiang Formation, Sifangtai, and Mingshui formations) in the ZKD-1 borehole. The Planktonic foraminifera include <em>Archaeoglobigerina blowi</em>, <em>A</em>. <em>cretacea</em>, and <em>Globotruncanella</em> sp. The benthic foraminifera include <em>Anomalinoides</em> sp., <em>Gavelinella</em> sp., and <em>Haplophragmoides</em> sp. Among them, <em>A</em>. <em>blowi</em>, <em>A</em>. <em>cretacea</em>, <em>Anomalinoides</em> sp., <em>Gavelinella</em> sp., <em>Haplophragmoides</em> sp. are widely distributed in the marine strata of the Upper Cretaceous. We tested the marine biomarker compounds including C<sub>30</sub> 24-<em>n</em>-propyl cholestane and C<sub>30</sub> 24-<em>iso</em>-propyl cholestane, indicating that the Songliao Basin experienced seawater incursion events to varying degrees during the middle to late Campanian. We also found abundant charophyte fossils in the ZKD-1 borehole, including <em>Atopochara trivolvis ulanensis</em>, <em>Latochara yuananensis</em>, <em>Feistiella anluensis</em>, <em>Mesochara biacuta</em>, <em>M</em>. <em>gobica</em>, and <em>M</em>. <em>leiocarpa</em>. Among them, the brackish water charophyte fossils of <em>Feistiella anluensis</em> were first found in Songliao Basin.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"34 5","pages":"Article 200958"},"PeriodicalIF":1.7,"publicationDate":"2025-05-16","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144170130","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-05-14DOI: 10.1016/j.palwor.2025.200957
André Mbabi Bitchong , Serge Alberto Bitjong , Serge Parfait Koah Na Lebogo , Glwadys Ngo Mandeng , Cecile Olive Mbesse , Soukaina Obad , Gilbert-François Ngon Ngon , Simon Ngos III , Thierry Adatte
The West and Central African Rifts System (WCARS) is well-known for its Cretaceous deposits. Still, the reliable dating of sedimentary successions, especially in the central part, and the mechanisms causing environmental and climate change over time remained less explored. Two sections of exposed deposits were investigated within the Babouri-Figuil Basin, specifically along the Mayo-Figuil (MF) and Mayo-Dakmoune (MD) rivers. The deposits are primarily composed of wackes formed from felsic and intermediate parent rocks, perhaps originating from the adjacent basement rocks, although there are also mafic components represented by an iron-rich shale layer at the MF section. Relative dating has constrained the MF sediment age to the Albian–Cenomanian and the MD sediment age to the Campanian–Maastrichtian. Using whole-rock organic matter geochemistry and trace element concentrations, we discovered anoxic conditions, well-preserved organic matter, and marine salinity water occurrence in the Albian–Cenomanian deposits, probably related to the first Trans-Saharan seaway. Oxic conditions, poor-preserved organic matter, and brackish salinity water occurrence were established during the Campanian–Maastrichtian. The Hg/TOC spike recorded during the Albian–Cenomanian could be the consequence of proximal volcanism, most likely associated with enhanced rifting tectonic activity, whereas the maximum Hg/TOC ratios observed during the Campanian–Maastrichtian could be the product of distal volcanism. Globally, weathering was slow to moderate, and the climate was interpreted as semi-arid throughout the middle to Late Cretaceous in the central WCARS.
{"title":"Paleoenvironmental records across middle to Upper Cretaceous in central part of the West and Central African Rift System: Palynological and geochemical inferences","authors":"André Mbabi Bitchong , Serge Alberto Bitjong , Serge Parfait Koah Na Lebogo , Glwadys Ngo Mandeng , Cecile Olive Mbesse , Soukaina Obad , Gilbert-François Ngon Ngon , Simon Ngos III , Thierry Adatte","doi":"10.1016/j.palwor.2025.200957","DOIUrl":"10.1016/j.palwor.2025.200957","url":null,"abstract":"<div><div>The West and Central African Rifts System (WCARS) is well-known for its Cretaceous deposits. Still, the reliable dating of sedimentary successions, especially in the central part, and the mechanisms causing environmental and climate change over time remained less explored. Two sections of exposed deposits were investigated within the Babouri-Figuil Basin, specifically along the Mayo-Figuil (MF) and Mayo-Dakmoune (MD) rivers. The deposits are primarily composed of wackes formed from felsic and intermediate parent rocks, perhaps originating from the adjacent basement rocks, although there are also mafic components represented by an iron-rich shale layer at the MF section. Relative dating has constrained the MF sediment age to the Albian–Cenomanian and the MD sediment age to the Campanian–Maastrichtian. Using whole-rock organic matter geochemistry and trace element concentrations, we discovered anoxic conditions, well-preserved organic matter, and marine salinity water occurrence in the Albian–Cenomanian deposits, probably related to the first Trans-Saharan seaway. Oxic conditions, poor-preserved organic matter, and brackish salinity water occurrence were established during the Campanian–Maastrichtian. The Hg/TOC spike recorded during the Albian–Cenomanian could be the consequence of proximal volcanism, most likely associated with enhanced rifting tectonic activity, whereas the maximum Hg/TOC ratios observed during the Campanian–Maastrichtian could be the product of distal volcanism. Globally, weathering was slow to moderate, and the climate was interpreted as semi-arid throughout the middle to Late Cretaceous in the central WCARS.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"34 5","pages":"Article 200957"},"PeriodicalIF":1.7,"publicationDate":"2025-05-14","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144138653","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-05-14DOI: 10.1016/j.palwor.2025.200955
Jiao Huang , Hui Jia , Rui-Feng Yan , Xiang-Ning Meng , Zhuo-Chen Han , Tian-Qi Dong , Jian Pan , Cheng Quan
Ostrya Scopoli, family Betulaceae, is mainly distributed in the Northern Hemisphere, with a rich fossil record in the Miocene. This study presents a new record of Ostrya involucres and nutlet fossils from the upper Eocene Bailuyuan Formation in Weinan City, Shaanxi, China. The fossil involucres are saccate, ellipsoid or ovate, with reticulate veins; the nutlet is ellipsoid-ovate, one side surface has five ribs, and is wingless. Based on morphological comparisons, a new species, Ostrya parajaponica Huang and Jia, n. sp. is proposed, which represents the earliest fossil record of Ostrya involucres globally. In this study, the historical distribution of Ostrya is investigated, and it is speculated that Ostrya originated in Asia before the late Eocene. During the late Eocene, Ostrya may have spread from Asia to North America via the Bering Land Bridge and then spread from North America to Europe via the North Atlantic Land Bridge. During the Oligocene, the closure of the Turgai Straits may have facilitated widespread exchange of Ostrya across the Eurasian continent. During the Miocene, during warm climates, Ostrya was widely distributed across the American and Eurasian continents. Subsequently, the cooling and drying of climates beginning in the Pliocene shaped the current geographical distribution pattern of Ostrya.
桦树科桐树属(Ostrya Scopoli)主要分布于北半球,在中新世有丰富的化石记录。本文报道了陕西渭南市始新统上白鹿园组的珍珠岩总壳和坚果化石的新记录。化石总苞囊状、椭球状或卵形,具网状脉;坚果为椭圆体卵形,一侧表面有五棱,无翅。在形态学比较的基础上,提出了一个新种——Ostrya parajaponica Huang and Jia, n. sp,它代表了全球最早的Ostrya involucres化石记录。本文研究了奥斯特拉属的历史分布,推测奥斯特拉属起源于晚始新世以前的亚洲。在始新世晚期,奥斯特拉可能通过白令陆桥从亚洲传播到北美,然后通过北大西洋陆桥从北美传播到欧洲。渐新世期间,图尔盖海峡的关闭可能促进了奥斯特利亚在欧亚大陆上的广泛交换。在中新世温暖的气候下,奥斯特亚广泛分布在美洲和欧亚大陆。随后,上新世开始的气候变冷和干燥形成了奥斯特亚现在的地理分布格局。
{"title":"Fossil involucres and a nutlet of Ostrya (Betulaceae) from the upper Eocene of Shaanxi and their biogeographic implications","authors":"Jiao Huang , Hui Jia , Rui-Feng Yan , Xiang-Ning Meng , Zhuo-Chen Han , Tian-Qi Dong , Jian Pan , Cheng Quan","doi":"10.1016/j.palwor.2025.200955","DOIUrl":"10.1016/j.palwor.2025.200955","url":null,"abstract":"<div><div><em>Ostrya</em> Scopoli, family Betulaceae, is mainly distributed in the Northern Hemisphere, with a rich fossil record in the Miocene. This study presents a new record of <em>Ostrya</em> involucres and nutlet fossils from the upper Eocene Bailuyuan Formation in Weinan City, Shaanxi, China. The fossil involucres are saccate, ellipsoid or ovate, with reticulate veins; the nutlet is ellipsoid-ovate, one side surface has five ribs, and is wingless. Based on morphological comparisons, a new species, <em>Ostrya parajaponica</em> Huang and Jia, n. sp. is proposed, which represents the earliest fossil record of <em>Ostrya</em> involucres globally. In this study, the historical distribution of <em>Ostrya</em> is investigated, and it is speculated that <em>Ostrya</em> originated in Asia before the late Eocene. During the late Eocene, <em>Ostrya</em> may have spread from Asia to North America via the Bering Land Bridge and then spread from North America to Europe via the North Atlantic Land Bridge. During the Oligocene, the closure of the Turgai Straits may have facilitated widespread exchange of <em>Ostrya</em> across the Eurasian continent. During the Miocene, during warm climates, <em>Ostrya</em> was widely distributed across the American and Eurasian continents. Subsequently, the cooling and drying of climates beginning in the Pliocene shaped the current geographical distribution pattern of <em>Ostrya</em>.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"34 5","pages":"Article 200955"},"PeriodicalIF":1.7,"publicationDate":"2025-05-14","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144138649","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-05-14DOI: 10.1016/j.palwor.2025.200956
Qing-Dan Zheng , Xiao-Ju Yang , Ting Li
Terrestrial plants are important indicators of palaeoclimate and palaeoenvironment during the geological history. The Ginkgoales leaves are commonly used to reconstruct the atmospheric palaeo-CO2 concentration (pCO2) during the Jurassic. Baiera, a genus of Ginkgoales, however, has been less reported in the reconstruction of the pCO2 during the Middle Jurassic. In this paper, Baiera hallei Sze from the lower Middle Jurassic (Aalenian) Yan’an Formation in the central Ordos Basin is studied in detail based on abundant materials. Ten well-preserved leaves of B. hallei were selected to calculate stomatal parameters, and their stomatal index was used to estimate the pCO2. The average SI was 6.19 ± 0.38, the pCO2 obtained by the stomatal ratio method was 1121 ± 73 ppm, and the average palaeotemperature reconstructed by McElwain’s empirical formula was 13–15.8°C. The pCO2 roughly coincides with the long-term carbon cycle model GEOCARB II. The palaeoclimate in Yulin district was warm and humid during the Aalenian (early Middle Jurassic).
{"title":"Palaeoenvironmental implications of Baiera hallei Sze from the Middle Jurassic in Shaanxi Province, China","authors":"Qing-Dan Zheng , Xiao-Ju Yang , Ting Li","doi":"10.1016/j.palwor.2025.200956","DOIUrl":"10.1016/j.palwor.2025.200956","url":null,"abstract":"<div><div>Terrestrial plants are important indicators of palaeoclimate and palaeoenvironment during the geological history. The Ginkgoales leaves are commonly used to reconstruct the atmospheric palaeo-CO<sub>2</sub> concentration (<em>p</em>CO<sub>2</sub>) during the Jurassic. <em>Baiera</em>, a genus of Ginkgoales, however, has been less reported in the reconstruction of the <em>p</em>CO<sub>2</sub> during the Middle Jurassic. In this paper, <em>Baiera hallei</em> Sze from the lower Middle Jurassic (Aalenian) Yan’an Formation in the central Ordos Basin is studied in detail based on abundant materials. Ten well-preserved leaves of <em>B</em>. <em>hallei</em> were selected to calculate stomatal parameters, and their stomatal index was used to estimate the <em>p</em>CO<sub>2</sub>. The average SI was 6.19 ± 0.38, the <em>p</em>CO<sub>2</sub> obtained by the stomatal ratio method was 1121 ± 73 ppm, and the average palaeotemperature reconstructed by McElwain’s empirical formula was 13–15.8°C. The <em>p</em>CO<sub>2</sub> roughly coincides with the long-term carbon cycle model GEOCARB II. The palaeoclimate in Yulin district was warm and humid during the Aalenian (early Middle Jurassic).</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"34 5","pages":"Article 200956"},"PeriodicalIF":1.7,"publicationDate":"2025-05-14","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144190024","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-04-29DOI: 10.1016/j.palwor.2025.200954
Han Wang , Zhi-Xin Sun , Hai-Chun Zhang
Eurypterids, commonly known as sea scorpions, were an extinct group of arthropods renowned for their large size and role as apex predators in Early Paleozoic marine ecosystems. Among them, carcinosomatids and pterygotids stand out as two particularly striking groups, both characterized by their large size but differing in morphological specialization. Compared to their diversity and abundance in Laurussia, both groups are rare in the larger Pan-Gondwanan region, which is consistent with the general pattern observed from most eurypterids, especially the Devonian genera. Here, we report a new diverse eurypterid fauna from the Nagaoling Formation (Lochkovian, Devonian) in Guangxi, South China, including Carcinosomatidae, Pterygotidae, and Adelophthalmidae. Tigrisopterus zengi n. gen. n. sp. represents the youngest known carcinosomatid worldwide and the first Devonian member of this family discovered outside Laurussia. Acutiramus sp., with an estimated body length of approximately 1.6–1.8 m, is the first Acutiramus component found in China and one of the youngest representatives of this genus known to date. These two families, along with the generalist Adelophthalmus found in the same bed, suggest a complex ecological structure in the Devonian shallow marine environment of South China. This discovery expands both the temporal and spatial distribution of carcinosomatids and pterygotids, providing new insights into the distribution and evolutionary history of these ancient predators.
广翅类,俗称海蝎子,是一种已灭绝的节肢动物,以其庞大的体型和在古生代早期海洋生态系统中作为顶级捕食者的角色而闻名。其中,癌体虫和翼虫是两个特别引人注目的类群,它们的特点都是体型大,但形态专门化不同。与Laurussia的多样性和丰度相比,这两个类群在更大的Pan-Gondwanan地区是罕见的,这与大多数泛蝶类,特别是泥盆纪属的一般模式一致。本文报道了广西泥盆纪那高岭组(Lochkovian, Devonian)的一种新的泛翅类动物群,包括癌虫科、翼虫科和Adelophthalmidae。Tigrisopterus zengi n. gen. n. sp.代表了世界上已知最年轻的癌体虫,也是该家族在俄罗斯以外发现的第一个泥盆纪成员。acutramus sp.体长约1.6 ~ 1.8 m,是中国发现的第一个acutramus成分,也是该属已知最年轻的代表之一。这两个科,连同在同一层位发现的多面体Adelophthalmus,表明中国南方泥盆纪浅海环境中存在一个复杂的生态结构。这一发现扩大了癌体虫和翼足虫的时空分布,为这些古代食肉动物的分布和进化历史提供了新的见解。
{"title":"New eurypterids from the Lower Devonian Nagaoling Formation of Guangxi, South China","authors":"Han Wang , Zhi-Xin Sun , Hai-Chun Zhang","doi":"10.1016/j.palwor.2025.200954","DOIUrl":"10.1016/j.palwor.2025.200954","url":null,"abstract":"<div><div>Eurypterids, commonly known as sea scorpions, were an extinct group of arthropods renowned for their large size and role as apex predators in Early Paleozoic marine ecosystems. Among them, carcinosomatids and pterygotids stand out as two particularly striking groups, both characterized by their large size but differing in morphological specialization. Compared to their diversity and abundance in Laurussia, both groups are rare in the larger Pan-Gondwanan region, which is consistent with the general pattern observed from most eurypterids, especially the Devonian genera. Here, we report a new diverse eurypterid fauna from the Nagaoling Formation (Lochkovian, Devonian) in Guangxi, South China, including Carcinosomatidae, Pterygotidae, and Adelophthalmidae. <em>Tigrisopterus zengi</em> n. gen. n. sp. represents the youngest known carcinosomatid worldwide and the first Devonian member of this family discovered outside Laurussia. <em>Acutiramus</em> sp., with an estimated body length of approximately 1.6–1.8 m, is the first <em>Acutiramus</em> component found in China and one of the youngest representatives of this genus known to date. These two families, along with the generalist <em>Adelophthalmus</em> found in the same bed, suggest a complex ecological structure in the Devonian shallow marine environment of South China. This discovery expands both the temporal and spatial distribution of carcinosomatids and pterygotids, providing new insights into the distribution and evolutionary history of these ancient predators.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"34 5","pages":"Article 200954"},"PeriodicalIF":1.7,"publicationDate":"2025-04-29","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144069058","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}