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Oryctocephalus indicus from the Hwangju area, North Hwanghae Province, the Democratic People’s Republic of Korea: Implication for the boundary between Series 2 and Miaolingian 朝鲜民主主义人民共和国黄海北道黄州地区的 Oryctocephalus indicus:第 2 系与苗岭系边界的含义
IF 1.7 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2024-04-01 DOI: 10.1016/j.palwor.2024.04.006
Kwang-Sik So, Yong-Su Ju, Jong-Su Ma, Phyong-Hen Kim, Yong-Sun Kim, Pong-Un Kim
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引用次数: 0
Osteomyelitis in the manus of Smilodon populator (Felidae, Machairodontinae) from the Late Pleistocene of South America 南美洲更新世晚期Smilodon种群(Felidae,Machairodotinae)手部的骨髓炎
IF 1.7 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2024-04-01 DOI: 10.1016/j.palwor.2023.05.001
Carlos A. Luna , Roy R. Pool , Marcos D. Ercoli , Nicolás R. Chimento , Fernando H. de S. Barbosa , Alfredo E. Zurita , Pedro Cuaranta

The knowledge of the paleopathologies that affected large mammals during the Pleistocene of South America has increased in the last years, but most of the reported cases belong to the endemic clade Xenarthra. On the contrary, almost no case of diseases is known among representatives of other clades, such as Carnivora. Here we present and describe an inflammatory lesion in a left metacarpal IV assigned to the saber tooth Smilodon populator (Felidae) from the Late Pleistocene (ca. 100 ka; MIS 5) of Northeastern Argentina. The macroscopic and radiologic analyses reveal features consistent with chronic osteomyelitis, which in turn represents the first accurate record of an infectious process in a limb of this predator. This injury presumably caused lameness and loss of toe flexion, and considerably reduced the hunting abilities of this top predator, which used its robust forelimbs, particularly wide forepaws, and powerful back muscles to catch and bring down large prey.

近年来,人们对南美洲更新世时期影响大型哺乳动物的古病理学有了更多的了解,但所报告的大多数病例都属于地方性支系Xenarthra。相反,在其他支系(如食肉目)的代表动物中,几乎没有已知的病例。在这里,我们介绍并描述了阿根廷东北部晚更新世(约 100 ka; MIS 5)剑齿虎(Smilodon populator,鼬科)左掌骨 IV 上的炎症病变。宏观和放射学分析显示了与慢性骨髓炎一致的特征,这也是首次准确记录这种食肉动物肢体的感染过程。这种损伤可能会导致跛行和趾屈曲功能丧失,并大大降低了这种顶级掠食者的捕猎能力,因为这种掠食者利用其强壮的前肢(尤其是宽大的前爪)和强大的背部肌肉来捕捉和捕杀大型猎物。
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引用次数: 0
Two anachoropterid fern rachises from the in situ volcanic ash of the Whetstone Horizon (Kladno Formation, Pennsylvanian), Radnice Basin, Czech Republic 捷克Radnice盆地wheetstone地平线(宾夕法尼亚州Kladno组)原位火山灰中的两个仿翼蕨
IF 1.7 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2024-04-01 DOI: 10.1016/j.palwor.2023.02.003
Wei-Ming Zhou , Josef Pšenička , Jana Votočková Frojdová , Jun Wang , Ming-Li Wan , Zhuo Feng

Two partly anatomically preserved axes parallelly situated on a slab collected from an in situ volcanic ash bed called “Bělka” of the Whetstone Horizon, Kladno Formation (Pennsylvanian), Czech Republic, were studied in detail. Anatomically, both axes possess a C-shaped xylem strand with protoxylem tracheids situated on the convex side, demonstrating an inversicatenalean-type anachoropterid affiliation. They are further suggested to belong to one biological species, as they share a number of similar characteristics and common structures. Systematically, one of the two studied axes retains a primitive form of foliar anatomy with the oldest known anachoropterid plant (Anachoropteris sp.) in having two rather short lateral arms compared to the long median region. Although lacking foliage information, both axes likely belong to the rachis part of Discosoropteris zlatkokvacekii Pšenička, Zhou, Boyce, Votočková Frojdová, Bek and Wang, a fern species that was recently established based on the same slab where the two studied axes were preserved. Such a combination may further indicate the presence of a new family in the late Paleozoic anachoropterid plants. In addition, selected anatomically preserved ferns from the Whetstone Horizon were reviewed, which promotes a better understanding of the anatomical variability of fern species.

研究人员对从捷克共和国克拉德诺地层(宾夕法尼亚)Whetstone地层名为 "Bělka "的原位火山灰床采集的一块石板上平行放置的两根部分解剖结构保存完好的轴进行了详细研究。从解剖学角度来看,两根轴都具有 C 形木质部股,原木质部管胞位于凸面,这表明它们属于逆戟鲸类。由于它们具有许多相似的特征和共同的结构,因此进一步认为它们属于一个生物物种。从系统上看,所研究的两个轴中的一个与已知最古老的鞘翅目植物(Anachoropteris sp.)虽然缺乏叶片信息,但这两根轴很可能属于 Discosoropteris zlatkokvacekii Pšenička, Zhou, Boyce, Votočková Frojdová, Bek 和 Wang 的轴部分。这样的组合可能进一步表明晚古生代achoropterid植物中存在一个新的科。此外,研究人员还对Whetstone地层中部分解剖学上保存完好的蕨类植物进行了回顾,这有助于更好地了解蕨类植物的解剖学变异性。
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引用次数: 0
Upper Oxfordian (Upper Jurassic) radiolarians from Rybaki Section, Moscow Region, Central Russia 俄罗斯中部莫斯科地区Rybaki剖面的上牛津阶(上侏罗纪)放射虫
IF 1.7 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2024-04-01 DOI: 10.1016/j.palwor.2023.03.001
Nikita Bragin, Liubov Bragina, Aleksandr Mironenko

A well-preserved radiolarian assemblage was recovered from the Upper Jurassic section near the Rybaki village, Moscow Region, Central Russia. The radiolarians were found within the Amoeboceras serratum ammonite zone (upper Oxfordian), in phosphatized body chambers of ammonite shells. The radiolarian assemblage is represented by 23 species of 11 genera and displays Boreal affinity. It is characterized by abundance of Praeparvicingula: P. blackhorsensis (Pessagno and Whalen), P. donnae Bragin, P. elementaria (Carter), P. enormis (Yang) and by presence of the characteristic boreal family Echinocampidae (genera Echinocampe and Nordvikella). Five new species are described.

在俄罗斯中部莫斯科地区雷巴基村附近的上侏罗纪地段发现了一批保存完好的放射虫。这些放射虫发现于 Amoeboceras serratum 菊石区(上牛津期),菊石壳的磷化体腔内。放射虫群由 11 个属的 23 个物种组成,具有北方亲缘关系。它的特点是有大量的 Praeparvicingula:P.blackhorsensis(Pessagno 和 Whalen)、P. donnae Bragin、P. elementaria(Carter)、P. enormis(Yang),以及具有北方特征的棘螈科(Echinocampe 和 Nordvikella 属)。描述了五个新物种。
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引用次数: 0
Biostratigraphy of Bajocian–Oxfordian strata based on dinoflagellate cysts and ammonites from the Dalichai Formation, northeastern Iran 基于伊朗东北部Dalichai组甲藻囊肿和菊石的Bajocian-Oxfordian地层生物地层学
IF 1.7 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2024-04-01 DOI: 10.1016/j.palwor.2023.04.003
Ebrahim Ghasemi-Nejad , Parvin Esmaili , Mahmoud Reza Majidifard , Mohammad Sharifi

The Bajocian–Oxfordian Dalichai Formation of the Binalud Mountains (northeastern Iran) consists of marls, marly limestones and limestones that were sampled and studied in detail for their palynomorph and ammonite contents. Palynological studies enabled differentiation of three marine biozones (Cribroperidinium crispum, Dichadogonyaulax sellwoodii, and Ctenidodinium continuum) within the lower interval of the formation. The palynofacies analysis shows a gradual sea-level rise during the depositional period of the rock unit, while terrestrial palynomorphs (spores and pollen grains) revealed a predominantly tropical climate for the time interval. Furthermore, a large number of ammonites were recovered from the formation, based on which 12 families, 22 genera, 30 species and 12 biozones (Zigzag, Aurigerus, Retrocostatum, Discus, Bullatus, Gracilis, Anceps, Coronatum, Athleta, Transversarium, Bifurcatus, and Bimammatum) were differentiated. When comparing the dinoflagellate cyst zones with the ammonite zones, the Dichadogonyaulax sellwoodii dinoflagellate cyst zone was correlated with the ammonite zones of Zigzag, Aurigerus, Retrocostatum, Discus, Bullatus, and Gracilis and the Ctenidodinium continuum dinocyst zone was correlated with the Gracilis and Anceps ammonite zones. The dinocyst and ammonite assemblages of the Binalud Mountains are similar to assemblages described from northern and northeastern Iran (Alborz and Koppeh-Dagh basins) as well as Northwest Europe. This suggests marine connections between these sedimentary basins during the Bajocian–Oxfordian. Consequently, the integrated dinocyst and ammonite zonation established here can be utilized also in other parts of the northern Tethyan Realm. The diachroneity of the upper boundary of the Dichadogonyaulax sellwoodii Zone between the Binalud Mountains and Northwest Europe is however revealed by comparison with the ammonite zones.

对比纳鲁德山(伊朗东北部)的巴约卡期-新石器时代达利柴地层(Bajocian-Oxfordian Dalichai Formation)的泥灰岩、泥灰质灰岩和灰岩进行了取样,并详细研究了其中的古生物和氨虫含量。通过古植物学研究,可以在该地层的下部区间区分出三个海洋生物区(Cribroperidinium crispum、Dichadogonyaulax sellwoodii 和 Ctenidodinium continuum)。古生界分析表明,在该岩石单元的沉积时期,海平面逐渐上升,而陆生古生界(孢子和花粉粒)则显示该时期主要为热带气候。此外,在该岩层中还发现了大量的氨甲虫,并据此划分出 12 科、22 属、30 种和 12 个生物区(Zigzag、Aurigerus、Retrocostatum、Discus、Bullatus、Gracilis、Anceps、Coronatum、Athleta、Transversarium、Bifurcatus 和 Bimammatum)。在比较甲藻胞囊区与鹦鹉螺区时,Dichadogonyaulax sellwoodii 甲藻胞囊区与 Zigzag、Aurigerus、Retrocostatum、Discus、Bullatus 和 Gracilis 的鹦鹉螺区相关,Ctenidodinium continuum 甲藻胞囊区与 Gracilis 和 Anceps 的鹦鹉螺区相关。比纳鲁德山的二叠体和氨石群与伊朗北部和东北部(Alborz 和 Koppeh-Dagh 盆地)以及欧洲西北部的二叠体和氨石群相似。这表明这些沉积盆地在巴约纪-新石器时代有海洋联系。因此,在此建立的综合二叠纪和氨虫区系也可用于泰提安地界北部的其他地区。不过,通过与鹦鹉螺区带的比较,可以发现比纳鲁德山和西北欧之间的 Dichadogonyaulax sellwoodii 区带的上边界具有非同步性。
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引用次数: 0
Growth patterns of palaeoscolecid sclerites from the Furongian (upper Cambrian) Wangcun section, western Hunan, South China 湘西芙蓉系(上寒武统)王村剖面古球藻硬结石生长模式
IF 1.7 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2024-04-01 DOI: 10.1016/j.palwor.2023.03.005
Xiao-Feng Xian , Mats E. Eriksson , Hua-Qiao Zhang

In a strict sense, palaeoscolecids are a group of cycloneuralian worms (ecdysozoans) characterized by the tessellation of trunk sclerites, which are differentiated into plates, platelets, and microplates. The Wangcun section in western Hunan, South China, which represents the Furongian Series (upper Cambrian), has yielded a rich and diverse collection of palaeoscolecids. However, there is uncertainty about how the morphology and arrangement of their sclerites change during ontogeny and the potential taxonomic implications of these changes. Here, we report new phosphatized trunk fragments from the Wangcun section, which are assigned to Dispinoscolex decorus, Hunanoscolex campus (= Ornatoscolex hunanensis), and Schistoscolex hunanensis. Trunk fragments of the same species with different diameter indicate the presence of several ontogenetic stages, and careful assessment revealed two types of sclerite growth patterns. One is represented by D. decorus and possibly also by S. hunanensis, in which the worm growth is accompanied mainly by the enlargement of the plates, with almost no change in the median annular zones and annulation boundaries. The other type is represented by H. campus, in which the worm growth is accompanied mainly by the widening of the median annular zones and annulation boundaries inserted with continuously more platelets and microplates. Because the specimens originally assigned to H. campus seem to represent late ontogenetic stages of O. hunanensis, we propose that H. campus is a senior synonym of the latter. This study underscores the importance of ontogenetic variation of sclerites for the taxonomy of palaeoscolecids, and highlights the significance of Orsten-type fossils in the study of metazoan ontogeny.

从严格意义上讲,古孔虫是一类旋毛虫(蜕皮虫),其特征是躯干硬骨呈棋盘格状,分为板状、小板状和微板状。华南湘西王村剖面代表了芙蓉系(上寒武统),该剖面出土了丰富多样的古小孔虫。然而,它们的硬骨在个体发育过程中的形态和排列如何变化以及这些变化对分类学的潜在影响尚不确定。在此,我们报告了王村剖面新发现的磷化躯干片段,将其归属于Dispinoscolex decorus、Hunanoscolex campus(= Ornatoscolex hunanensis)和Schistoscolex hunanensis。同一物种不同直径的树干碎片表明存在几个发育阶段,仔细评估后发现有两种硬骨生长模式。一种以 D. decorus 为代表,可能也以 S. hunanensis 为代表,在这种类型中,蠕虫的生长主要伴随着板的增大,中间环带和环带边界几乎没有变化。另一种类型以校园虫(H. campus)为代表,虫体的生长主要伴随着中间环带和环带边界的扩大,并不断插入更多的小板和微板。由于最初归入 H. campus 的标本似乎代表了 O. hunanensis 的晚期发育阶段,因此我们建议 H. campus 是后者的高级异名。这项研究强调了硬骨鱼的发育变异对于古硬骨鱼类分类学的重要性,并突出了奥尔斯滕类型化石在研究元古宙发育过程中的重要意义。
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引用次数: 0
Middle Miocene “Giraffa” from the Chinji Formation of Siwaliks, Pakistan 巴基斯坦西瓦利克Chinji组中新世中期的“Giraffa”
IF 1.7 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2024-04-01 DOI: 10.1016/j.palwor.2023.03.003
Muhammad Asim , Kiran Aftab , Muhammad Akbar Khan , Sayyed Ghyour Abbas , Muhammad Adeeb Babar , Mubashar Hussain , Sania Saeed

A comprehensive record of unique and endemic species of “Giraffapriscilla has been documented and reported from the Chinji Formation of Lower Siwaliks, Pakistan. The material comprises incisors, canines, premolars, molars, mandibular and maxillary fragments which have been recovered from the Middle Miocene localities of Dhok Bun Ameer Khatoon, Rakh Wasnal, Bhelomar, and Lawa, Punjab, Pakistan. “Giraffapriscilla is a primitive species of giraffids, having a broad crown, strong styles, and median ribs.

巴基斯坦下西瓦利克Chinji地层记录并报告了 "长颈鹿 "priscilla的独特和特有物种。这些材料包括门齿、犬齿、前臼齿、臼齿、下颌和上颌残片,均采自巴基斯坦旁遮普省 Dhok Bun Ameer Khatoon、Rakh Wasnal、Bhelomar 和 Lawa 的中新世地区。长颈鹿 "priscilla "是长颈鹿类的原始物种,具有宽阔的冠部、强壮的花柱和中肋。
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引用次数: 0
Latest Devonian–Early Mississippian conodont biostratigraphy in the Naqing section, Guizhou, South China 贵州纳庆剖面晚泥盆世-早密西西比牙形石生物地层学
IF 1.7 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2024-04-01 DOI: 10.1016/j.palwor.2023.02.004
Yu-Ze Huang , Yu-Ping Qi , Qiu-Lai Wang , Le Yao , Ji-Tao Chen

The Naqing section in Luodian (Guizhou, South China) contains a deep-water carbonate-dominated succession of the Late Devonian through the Permian. In this study, 72 conodonts species/subspecies of 11 genera were obtained from the lower part of this section including. Seven middle to late Famennian and four early to late Tournaisian conodont biozones were recognized as, in ascending order, Palmatolepis marginifera marginifera Zone, Palmatolepis marginifera utahensis Zone, Polygnathus granulosus Zone, Palmatolepis rugosa trachytera Zone, Polygnathus styriacus Zone, Palmatolepis gracilis manca Zone, Palmatolepis rugosa rugosa Zone, Siphonodella isosticha Zone, Gnathodus punctatus Zone, Gnathodus typicus-Gnathodus cuneiformis Zone, and Gnathodus typicus-Protognathodus cordiformis Zone. Several key conodont biozones across the Devonian–Carboniferous Boundary (DCB) interval found elsewhere are not discovered in the Naqing section, likely due to a conodont-barren interval in the siliciclastic facies. The middle Famennian–late Tournaisian conodont biozones illustrated in this study provide a robust chronostratigraphic framework for further research on biotic and paleo-environmental events during the Late Devonian through Early Mississippian critical greenhouse-icehouse climate transition.

罗甸(贵州,华南)纳清剖面包含晚泥盆世至二叠世以深水碳酸盐为主的演替。本研究从该剖面下部获得了 11 个属的 72 个锥齿类物种/亚种,其中包括:7 个中-晚法门世物种和 4 个早-晚梯世物种。从高到低的顺序为:Palmatolepis marginifera marginifera 区、Palmatolepis marginifera utahensis 区、Polygnathus granulosus 区、Palmatolepis rugosa trachytera 区、Palmatolepis marginifera utahensis 区、Polygnathus granulosus 区、Palmatolepis rugosa trachytera 区、Palmatolepis marginifera utahensis 区、Palmatolepis marginifera utahensis 区、Siphonodella isosticha 区、Gnathodus punctatus 区、Gnathodus typicus-Gnathodus cuneiformis 区和 Gnathodus typicus-Protognathodus cordiformis 区。其他地方发现的泥盆纪-石炭纪界线(DCB)间的几个关键性的锥齿轮生物区在那青段没有发现,这可能是由于硅质岩层中的锥齿轮生物区造成的。本研究揭示的法门期中期-晚图尔奈斯期锥齿轮生物区为进一步研究晚泥盆世至早密西西比世关键温室-冰室气候转变期间的生物和古环境事件提供了一个可靠的年代地层框架。
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引用次数: 0
Dinosaur fauna from the Lower Cretaceous of Phu Kao-Phu Phan Kham, northeastern Thailand: a review and update 泰国东北部Phu Kao Phu Phan Kham下白垩纪的恐龙区系:回顾和更新
IF 1.7 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2024-04-01 DOI: 10.1016/j.palwor.2023.03.007
Adun Samathi , Suravech Suteethorn , Tanachot Boonjarern , Krishna Sutcha , Varavudh Suteethorn

Phu Kao-Phu Phan Kham Mountain ranges, situated on the border of Nong Bua Lamphu, Udon Thani, and Khon Kaen provinces, northeastern Thailand, bear several localities that yield dinosaurs and other vertebrate faunas in the Lower Cretaceous. However, this area has received relatively little attention from the scientific community and the public. Here we review and report new dinosaur materials recovered from this region, some found decades ago, others discovered recently. We identify and discuss their taxonomy and paleobiodiversity. In the Barremian Sao Khua Formation, theropod dinosaurs were represented by spinosaurids, an early branching megaraptoran, a large indeterminate theropod, and small indeterminate theropods. Sauropods were represented by an indeterminate sauropod, a neosauropod, and titanosauriforms, including probable brachiosaurid and non-titanosaur somphospondylans. Other vertebrate groups, including crocodilians, turtles, fishes, and sharks, indicate a similar composition to other Sao Khua Formation communities in northeastern Thailand. The Sao Khua vertebrate assemblage exhibits the taxonomic diversity of dinosaur fauna and is currently the best-documented assemblage in the Early Cretaceous of Southeast Asia. The Aptian–Albian Khok Kruat Formation, however, yields less diverse vertebrates. They are represented by freshwater sharks, crocodilians, and spinosaurid theropods. This may be due to sampling bias, environmental, or taphonomic conditions. This study provides the basis for future paleontological exploration and research of Mesozoic vertebrates in northeastern Thailand.

Phu Kao-Phu Phan Kham 山脉位于泰国东北部 Nong Bua Lamphu、Udon Thani 和 Khon Kaen 三府交界处,有几个地方出土了下白垩世的恐龙和其他脊椎动物。然而,这一地区受到科学界和公众的关注相对较少。在此,我们回顾并报告了从该地区发现的新恐龙材料,有些是几十年前发现的,有些则是最近发现的。我们对它们的分类和古生物多样性进行了鉴定和讨论。在巴里米亚时期的圣库瓦地层中,恐龙的代表有棘龙类、一种早期分支的甲龙类、一种大型不确定的兽脚类恐龙和小型不确定的兽脚类恐龙。蜥脚类恐龙的代表有一种不确定的蜥脚类恐龙、一种新蜥脚类恐龙和泰坦龙类,包括可能的腕龙类和非泰坦龙类的体骨龙。其他脊椎动物类群,包括鳄鱼、海龟、鱼类和鲨鱼,显示出与泰国东北部其他老夸地层群落相似的组成。Sao Khua脊椎动物群表现出恐龙动物分类的多样性,是目前东南亚早白垩世记录最完整的脊椎动物群。然而,安普顿-阿尔卑斯Khok Kruat Formation地层的脊椎动物种类较少。它们以淡水鲨鱼、鳄鱼和棘龙类脊椎动物为代表。这可能是取样偏差、环境或岩石学条件造成的。这项研究为今后泰国东北部中生代脊椎动物的古生物学探索和研究奠定了基础。
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引用次数: 0
Vegetation dynamics in the Early–Middle Pennsylvanian coal swamps of the Nord-Pas-de-Calais Coalfield, France 法国北加莱煤田早-中宾夕法尼亚煤沼泽的植被动态
IF 1.7 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2024-04-01 DOI: 10.1016/j.palwor.2023.04.005
Azucena Molina-Solís , Christopher J. Cleal , Eliott Capel , Borja Cascales-Miñana

During recent years, different studies have focused on characterising plant diversities in the Carboniferous environments of the Variscan Foreland. One of these areas, the Nord-Pas-de-Calais Coalfield, has a Namurian–Westphalian sequence that has historically yielded abundant evidence of vegetation change, but to date, little attention has been paid to its macrofloral diversity dynamics. Here we show, for the first time, a comprehensive characterisation of the diversity changes and macroevolutionary patterns from this coalfield. The results show evidence of low levels of species richness during the Namurian, followed by an exponential diversification at the base of Westphalian. Duckmantian–Bolsovian species diversity continues to progressively increase, suggesting relatively stable conditions at that time. Maximum species richness is observed at the mid-Bolsovian followed by a major depletion during the Asturian. Overall, this pattern is quite similar to that seen in other parts of the Variscan Foreland. However, we see Nord-Pas-de-Calais vegetation diversified earlier than those in the British parts, probably because the dynamics of marine incursions and lacustrine conditions delayed its development in the latter areas. Furthermore, we also see the Asturian decline of the Nord-Pas-de-Calais coal swamp occurred later than in other basins, such as in the Pennines, Ruhr, and South Wales. This scenario suggests that the collapse of this biome was probably caused by increased sediment influx and reduced subsidence, resulting from the northwards migration of the Variscan Mountains. This further supports other studies in Europe advocating that climate change was at most only a subsidiary factor in the collapse of the Pennsylvanian coal swamp biomes.

近年来,不同的研究都侧重于描述瓦里斯坎前陆石炭纪环境中的植物多样性特征。其中一个地区--北加莱海峡煤田(Nord-Pas-de-Calais Coalfield)的纳穆尔-西弗勒(Namurian-Westphalian)序列在历史上提供了大量植被变化的证据,但迄今为止,人们很少关注其宏观植物多样性动态。在这里,我们首次展示了该煤田多样性变化和宏观进化模式的综合特征。研究结果表明,纳木耳期的物种丰富度较低,随后在威斯特伐利亚期底部出现了指数式的多样化。鸭芒纪-波尔索夫纪的物种多样性继续逐步增加,表明当时的条件相对稳定。在波尔索瓦纪中期,物种丰富度达到最高值,随后在阿斯图里亚纪出现了物种严重减少的现象。总体而言,这一模式与瓦里斯坎前陆其他地区的模式十分相似。不过,我们看到北加莱海峡植被的多样化早于英国部分地区,这可能是因为海洋入侵的动态和湖泊条件延迟了后者地区植被的发展。此外,我们还看到北加莱海峡煤炭沼泽的阿斯图里亚斯衰退期晚于其他盆地,如宾夕法尼亚、鲁尔和南威尔士。这种情况表明,这一生物群落的崩溃可能是由于瓦里斯坎山脉向北迁移导致沉积物流入量增加和沉降量减少造成的。这进一步支持了欧洲其他研究的观点,即气候变化最多只是宾夕法尼亚煤沼生物群落崩溃的一个次要因素。
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