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A new genus and species of Apocynaceae (Gentianales) seed macrofossil from the Early Miocene amber of Simojovel de Allende, Chiapas, Mexico 墨西哥恰帕斯州Simojovel de Allende早中新世琥珀中夹带科(龙胆属)种子大化石一新属、新种
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-07-03 DOI: 10.1016/j.palwor.2025.200983
Leonardo O. Alvarado-Cárdenas , Naylet K. Centeno-González , C. Sofía Islas-Hernández , Emilio Estrada-Ruiz
The Miocene amber deposit of Chiapas, Mexico, is one of the most important records in the world. It has a high diversity of plant inclusions, particularly the reproductive structures that are seldom preserved. Here we describe a new inclusion of a seed that has a narrowly oblong to linear shape, acuminate chalazal area, longitudinally striated, funicular furrow, micropylar tuft of hairs, all features found in Apocynaceae. Although Mexico is one of the centers of diversity for the family, with an estimated endemism rate of approximately 50%, the fossil record of this family in the country remains limited. Comparison with the extant and fossil seeds did not allow assignment to any known taxa, and therefore a new genus and species, Mesechitespermum endressiorum n. gen. n. sp., is proposed here. This new Miocene record supports the hypothesis that during this time the family was already present in some tropical areas of southeastern Mexico.
墨西哥恰帕斯的中新世琥珀矿床是世界上最重要的记录之一。它具有高度多样性的植物包裹体,特别是很少保存的生殖结构。在这里,我们描述了一种新的种子,它具有狭长圆形到线形,渐尖合点区,纵向条纹,索缕沟,微孔毛丛,在夹竹桃科中发现的所有特征。尽管墨西哥是该家族多样性的中心之一,估计其特有率约为50%,但该家族在该国的化石记录仍然有限。通过与现存种子和化石种子的比较,无法将其归属于任何已知的分类群,因此本文提出了一个新的属和种Mesechitespermum endressiorum n. gen. n. sp.。这个新的中新世记录支持了一种假设,即在此期间,这个家族已经出现在墨西哥东南部的一些热带地区。
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引用次数: 0
Clonal budding and a sessile stalk in conulariids suggested by a new genus from China 中国菊科植物的无性系出芽和无柄茎
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-07-03 DOI: 10.1016/j.palwor.2025.200982
Hui Min , Rui-Wen Zong , Xun-Lian Wang
The new genus Sericonularia n. gen. is established based on material from the lower Silurian Fentou Formation in Wuhan, China. This genus differs from other conulariids in having extremely thin transverse ribs, thick septa along the corners of the periderm, and flat interspaces. The reproductive mode and the sessile stalk of conulariids have long been controversial. In this study, the holotype of the type species Sericonularia gemmata n. gen. n. sp. provides additional information about possible asexual budding in conulariids; moreover, this specimen offers additional evidence on the structure and morphology of the sessile stalk. We hypothesize that once the buds could live independently, they detached from their parent and then found a suitable substrate; notably, some conulariids used sessile stalks, rather than attachment discs, during their sessile life phase. These stalks appear to have the same structure as the upper periderm. Overall, our observations provide new insights into conulariid affinities and lifestyle.
根据武汉下志留统汾头组的资料,建立了绢虫属(Sericonularia n. gen.)。这个属不同于其他圆锥类在有极薄的横棱,沿周被的角厚的隔,和平的间隙。圆锥植物的生殖方式和无柄茎长期以来一直存在争议。在本研究中,模式种Sericonularia gemmata n. gen. n. sp.的全型为圆锥植物可能的无性出芽提供了额外的信息;此外,该标本为无柄茎的结构和形态提供了额外的证据。我们假设,一旦芽能够独立生活,它们就会从亲本中分离出来,然后找到合适的基质;值得注意的是,一些圆锥植物在其无柄生命阶段使用无柄茎,而不是附着盘。这些茎似乎与上周皮具有相同的结构。总的来说,我们的观察结果提供了新的见解,以了解锥体的亲和力和生活方式。
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引用次数: 0
New insights into the paleobiogeography of teleosauroid crocodylomorphs in Southeast Asia 东南亚鳄鱼类的古生物地理学新认识
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-06-30 DOI: 10.1016/j.palwor.2025.200981
Supanut Bhuttarach , Uthumporn Deesri , Jeremy E. Martin , Sita Manitkoon , Jirasak Charoenmit , Komsorn Lauprasert
Teleosauroids are poorly known from Southeast Asia, with limited records from the Indochina Terrane of Thailand. To better understand their distribution and diversity, we describe new teleosauroid remains from two distinct geological terranes in Thailand. A nearly complete periorbital region and isolated teeth found in association from the Phu Kradung Formation (Upper Jurassic–Lower Cretaceous) of the Indochina Terrane are identified as cf. Indosinosuchus sp. The cranial specimen exhibits periorbital dimensions significantly larger than those of previously described Indosinosuchus specimens, likely representing the largest known individual of the genus and expanding our understanding of morphological diversity within this taxon. The discovery confirms the wide distribution of teleosauroids within the Phu Kradung Formation of the Khorat Plateau during the Late Jurassic to Early Cretaceous. Additionally, a frontal bone, left postorbital, and isolated teeth from the Klong Min Formation (upper Middle–Upper Jurassic) of the Sibumasu Terrane are assigned to Teleosauroidea indet. These remains, morphologically more primitive than Indosinosuchus, represent the first evidence of teleosauroids from southern peninsular Thailand and extend their known temporal range to the Middle Jurassic. Our findings demonstrate that teleosauroids were widely distributed across Thailand’s freshwater environments from the Middle Jurassic through Early Cretaceous, significantly expanding our understanding of their biogeographic distribution in Southeast Asia.
在东南亚,只有泰国的印度支那地界有有限的记录。为了更好地了解它们的分布和多样性,我们描述了来自泰国两个不同地质地形的新的远地小行星遗迹。在印度支那地系Phu Kradung组(上侏罗统-下白垩统)中发现的一个几乎完整的眶周区域和孤立的牙齿被鉴定为cf. Indosinosuchus sp.。该颅骨标本的眶周尺寸明显大于先前描述的Indosinosuchus标本,可能代表了该属已知的最大个体,并扩展了我们对该分类单元形态多样性的理解。这一发现证实了呼叻高原富克拉东组在晚侏罗世至早白垩世的广泛分布。此外,额骨、左眼眶后的和孤立的牙齿从运河分钟形成(上层中上侏罗纪)Sibumasu岩层被分配到Teleosauroidea indet。这些化石在形态上比Indosinosuchus更原始,代表了泰国半岛南部的首个远龙化石证据,并将其已知的时间范围扩展到中侏罗世。研究结果表明,从中侏罗世到早白垩世,远蜥龙广泛分布在泰国的淡水环境中,极大地扩展了我们对其在东南亚生物地理分布的认识。
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引用次数: 0
Chemostratigraphy of Albian deposits in northern Tunisia: Identification of early Albian Oceanic Anoxic Event in the southern Tethys 突尼斯北部Albian沉积物的化学地层学:特提斯南部早期Albian海洋缺氧事件的鉴定
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-06-28 DOI: 10.1016/j.palwor.2025.200980
Salma Soussi , Fares Khemiri , Amina Mabrouk El Asmi , Moez Ben Fadhel , Habib Troudi , Khaled El Asmi , Gonzalo Márquez Martinez
An integrated analysis of multiple geochemical proxies, including δ13Corg, total organic carbon (TOC), trace elements, and chemical gamma ray, coupled with a microfacies study and an age calibration using planktonic foraminiferal markers, was conducted at the Ain Asker section in northern Tunisia. This study provides significant insights into the depositional and paleoenvironmental dynamics of the mid-Cretaceous.
The carbon isotopic excursions observed at the Ain Asker section reflect a notable increase in the burial rate of marine organic carbon, specifically confined to the middle part of the Microhedbergella rischi biozone. TOC values recorded from the F2-ASK unit reach up to 2.3% combined with a δ13Corg values reaching -24.49‰, suggesting substantial accumulation of decayed marine organic matter. Although full anoxic conditions were not prevalent during deposition, evidence of oxygen-depleted bottom waters is clearly supported by the distribution patterns of redox-sensitive trace elements, such as vanadium and nickel. These findings underscore the role of partial oxygen restriction in facilitating organic matter preservation.
The enhanced burial of organic carbon is attributed to increased nutrient influx, likely driven by global sea-level transgressions and heightened surface productivity. These processes created favorable conditions for organic matter production and preservation. The integration of these sedimentological and geochemical findings with comparative records from Europe (France and Italy), China, and Mexico enables reliable correlation with the Paquier black shale level, a hallmark of OAE1b.
The Ain Asker section, through its detailed geochemical and isotopic signatures, emerges as a critical reference section for unraveling mid-Cretaceous anoxia dynamics within the southern Tethyan Realm. Its study enhances our understanding of the interplay between global paleoceanographic events and local depositional processes, shedding light on the mechanisms of organic carbon burial during a pivotal period in Earth’s history.
对突尼斯北部Ain Asker剖面进行了δ13Corg、总有机碳(TOC)、微量元素和化学伽马等多种地球化学指标的综合分析,并结合微相研究和利用浮游有孔虫标记物进行了年龄标定。该研究对中白垩世沉积和古环境动力学具有重要意义。在Ain Asker剖面观测到的碳同位素偏移反映了海相有机碳埋藏速率的显著增加,特别是局限于Microhedbergella rischi生物带中部。F2-ASK单元TOC值高达2.3%,δ13Corg值达-24.49‰,表明海相有机质大量聚集。尽管在沉积过程中完全缺氧的条件并不普遍,但氧化还原敏感的微量元素(如钒和镍)的分布模式清楚地支持了底水缺氧的证据。这些发现强调了部分限氧在促进有机物保存中的作用。有机碳埋藏的增强可归因于营养物质流入的增加,这可能是由全球海平面海侵和地表生产力提高所驱动的。这些过程为有机质的产生和保存创造了有利条件。将这些沉积学和地球化学发现与欧洲(法国和意大利)、中国和墨西哥的比较记录相结合,可以与Paquier黑色页岩水平(OAE1b的标志)进行可靠的对比。Ain Asker剖面通过其详细的地球化学和同位素特征,成为揭示特提斯王国南部白垩纪中期缺氧动力学的关键参考剖面。它的研究增强了我们对全球古海洋事件与局部沉积过程之间相互作用的理解,揭示了地球历史上关键时期有机碳埋藏的机制。
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引用次数: 0
Scleractinian corals from the Middle Jurassic Buqu Formation, Qiangtang block and their palaeogeographic implications 羌塘地块中侏罗统布曲组石珊瑚及其古地理意义
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-06-27 DOI: 10.1016/j.palwor.2025.200978
Xiu-Ping Zhu , Kun Liang , Wei-Hua Liao , Jia-Run Yin , Xin Rao , Yi-Chun Zhang
This is the first comprehensive systematic study of the Middle Jurassic scleractinian corals from the Qiangtang block, based on 225 fossil coral specimens collected from the Buqu Formation in the Biluocuo area of northern Xizang (Tibet). Three species of three genera are identified: Montlivaltia zangbeiensis, Pseudocoenia slovenica, and Kobyastraea coquandi. The assemblage is dominated by solitary Montlivaltia (92.44%), with significantly fewer compound colonies of Pseudocoenia (4.89%) and Kobyastraea (2.67%). Comparative analysis reveals that the Qiangtang coral fauna shows strong affinities with contemporaneous Tethyan assemblages, particularly showing the closest similarities to coral communities from East-Central Iran. These findings provide compelling evidence for a close palaeobiogeographic connection between the Qiangtang block and the central Tethyan region during the Middle Jurassic.
本文以西藏北部碧洛错地区布曲组225个珊瑚化石标本为基础,首次对羌塘地块中侏罗统核状系珊瑚进行了全面系统的研究。鉴定出3属3种:西藏蒙特瓦利亚(Montlivaltia zangbeiensis)、斯洛文尼亚Pseudocoenia slovenica和coquandi Kobyastraea。以单生Montlivaltia(92.44%)为主,复合菌落Pseudocoenia(4.89%)和Kobyastraea(2.67%)较少。对比分析表明,羌塘珊瑚动物群与同期的特提斯组合具有很强的相似性,特别是与伊朗中东部的珊瑚群落最相似。这些发现为中侏罗世羌塘地块与特提斯中部地区的古生物地理联系提供了有力证据。
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引用次数: 0
Calophyllum (Calophyllaceae) with high leaf mass per area from the upper Miocene of Beihai, low-latitude China 低纬北海中新世上部叶重较高的Calophyllum (Calophyllaceae)
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-06-26 DOI: 10.1016/j.palwor.2025.200979
Shu-Ran Tang , Qi-Jia Li , Hui Jia , Jian-Hua Jin , Cheng Quan
Plant development is highly influenced by ecological conditions, with different latitudes hosting vegetation with distinct component and ecological strategies. However, the ecological strategies of low-latitude plants in deep time remain poorly understood. In this study, we examine well-preserved fossil leaves from the upper Miocene of Beihai, low-latitude China, and describe three species of Calophyllum (Calophyllaceae) based on morphological and anatomical characteristics. Using leaf economic concepts, the ecological strategy of Calophyllum during the late Miocene is also evaluated. Our results suggest that all the three fossil species have leaf mass per area (LMA) higher than their nearest living relatives. It appears that the longer leaf lifespan driven by longer growing season likely contributed to the high LMA and hence the slow-return ecological strategy of Calophyllum in the late Miocene hereof. This study highlights the leaf lifespan in regulation of plant ecological strategies and provides insights into the palaeoecology of low-latitude vegetation during the late Miocene.
植物的发育受生态条件的影响较大,不同纬度的植被具有不同的组成成分和生态策略。然而,人们对低纬度植物在远古时期的生态策略了解甚少。本文对中国低纬度地区北海中新世上部保存完好的叶化石进行了研究,并基于形态学和解剖学特征对三种Calophyllum (Calophyllaceae)进行了描述。利用叶片经济学的概念,评价了晚中新世卡罗勒属植物的生态策略。结果表明,这三个化石种的叶面积质量(LMA)均高于其最近的亲缘种。较长的生长季节所导致的较长的叶片寿命可能导致了中新世晚期Calophyllum的高LMA,从而导致了缓慢回归的生态策略。本研究强调了叶片寿命对植物生态策略的调控作用,并为晚中新世低纬度植被的古生态学提供了新的见解。
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引用次数: 0
Spatiotemporal pattern of vegetation evolution in northeastern Tibetan Plateau during the Cenozoic and its climatic significance
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-06-19 DOI: 10.1016/j.palwor.2025.200977
Yun-Xia Jia , Wen-Chao Zhang , Li-Jun Tian , Zhi-Lin He , Zhao Wang , Qing-Chun Tian
The modern Asian environment comprises monsoon-dominated East Asia and the arid Asian interior. However, when and how the modern climatic environment of Asia evolved remains controversial. Quantitative and spatio-temporal paleovegetation reconstructions are crucial for addressing this issue. We compiled 10 well-dated pollen records across the northeastern Tibetan Plateau, stretching from western China into eastern China. We used the biomization method to generate a set of biome scores and landscape openness spanning the interval from the Eocene to Recent. Biome reconstructions reveal that five dominant vegetation types have attained the highest scores since ∼42 Ma, namely warm mixed forest, temperate deciduous forest, cool mixed forest, steppe, and desert. Spatially, our results reveal that the climate was predominantly wetter in the west than in the east during the Paleogene, as evidenced by the higher values of landscape openness in the west. Since ∼24 Ma this climate configuration has reversed. Temporally, our results reveal wet-dry climatic fluctuations during the Paleogene, as evidenced by the coexistence of open vegetation and forest vegetation. Then, our results reveal a reorganization of the climatic environment since ∼24 Ma, as evidenced by the presence of forest vegetation types in the east and decreasing trend of landscape openness in the west. Subsequently, our results reveal region-wide aridification since ∼8 Ma, as evidenced by the negative values of landscape openness in many regions. Together with previous paleoclimate records, we infer that the retreat of the Paratethys Sea was responsible for wet-dry fluctuations during the Paleogene. We also infer that the uplift of Central Xizang (Tibet) triggered the permanent reorganization of the Asian environment at the Oligocene/Miocene transition, and that global cooling promoted the subsequent aridification trend.
现代亚洲环境包括季风主导的东亚和干旱的亚洲内陆。然而,亚洲现代气候环境是何时以及如何演变的,仍然存在争议。定量和时空古植被重建是解决这一问题的关键。我们使用生物化方法生成了从始新世到最近的一组生物群系评分和景观开放度。生物群系重建显示,自~ 42 Ma以来,5种优势植被类型的得分最高,分别是温暖混交林、温带落叶林、凉爽混交林、草原和沙漠。从空间上看,古近纪西部比东部气候偏湿润,西部景观开放度较高。自~ 24 Ma以来,这种气候结构发生了逆转。在时间上,我们的研究结果揭示了古近纪的干湿气候波动,表现为开放植被和森林植被并存。然后,我们的研究结果揭示了自~ 24 Ma以来气候环境的重组,东部存在森林植被类型,西部景观开放性呈下降趋势。随后,我们的研究结果揭示了自~ 8 Ma以来区域范围内的干旱化,许多地区的景观开放度为负值。结合前人的古气候记录,我们认为古近纪干湿波动的主要原因是傍提提斯海的退缩。
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引用次数: 0
New larger foraminiferal taxa (Hereceina n. gen. and new Borelis species) from upper Priabonian of eastern Türkiye (Turkey): Biostratigraphic and paleoecological implications 土耳其东<s:1> rkiye上Priabonian大有孔虫新种(Hereceina n. gen.和Borelis新种):生物地层和古生态意义
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-06-13 DOI: 10.1016/j.palwor.2025.200970
Şükrü Acar, Alper Bozkurt
The Priabonian (Eocene) shallow-water carbonates of the Malatya region (eastern Türkiye) contain rich benthic foraminiferal assemblages. A new calcarinid genus (Hereceina n. gen.) with its type species (Hereceina spinigera n. gen. n. sp.) and two Borelis species (Borelis dizerae n. sp. and Borelis sozerii n. sp.) are systematically described from these deposits. A late Priabonian age, corresponding to SBZ 20, is assigned to the new taxa based on the co-occurrence of biostratigraphic markers such as Heterostegina gracilis and Spiroclypeus carpaticus. Based on microfacies studies, Hereceina spinigera n. gen. n. sp. is documented from inner to proximal middle ramp environments that were influenced by wave and current action. The introduction of these new Borelis species with Y-shaped septula from the upper Priabonian of the region provides new insights into the evolutionary history of the genus. This study highlights the significant potential of the new taxa that appeared during the Eocene–Oligocene marine extinctions to serve as valuable biostratigraphic markers at Tethyan localities.
马拉提亚地区(东基耶)普里亚盆期(始新世)浅水碳酸盐岩中含有丰富的底栖有孔虫组合。从这些沉积物中系统地描述了一个新的钙石属(Hereceina n.gen .)及其模式种(Hereceina spinigera n.gen . sp.)和两个Borelis dizerae n.sp .和Borelis sozerii n.sp .)。根据异质stegina gracilis和carpatius Spiroclypeus共存的生物地层标志,确定该新分类群的年龄为晚Priabonian,对应于sbz20。在微相研究的基础上,研究了受波浪和水流作用影响的中斜坡内至近中斜坡环境。从该地区上Priabonian引进的这些具有y形隔的Borelis新种为该属的进化史提供了新的见解。这项研究强调了在始新世-渐新世海洋灭绝期间出现的新分类群的巨大潜力,可以作为特提斯地区有价值的生物地层标志。
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引用次数: 0
Microclots in early Paleoproterozoic stromatolites from Dashiling Formation, Wutai, North China: Implications for microbial calcification and atmospheric CO2 levels during Great Oxidation Event 五台大岭组早古元古代叠层石中的微凝块:对大氧化事件时期微生物钙化和大气CO2水平的启示
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-06-09 DOI: 10.1016/j.palwor.2025.200969
Zhao-Yang Yi , Zhen Yan , Xin Wei , Kai Wang , Bing Shen , Jian-Bo Liu
The relationship between microbial calcification and atmospheric CO2 levels is a critical aspect of Earth’s biogeochemical evolution. This study presents the first detailed investigation of micro- and mesoscopic features of stromatolites from the 2.1 Ga Dashiling Formation of the Hutuo Group in Wutai area, North China, one of the best preserved microbialites during the Great Oxidation Event (GOE). The study identifies three types of microclots (microclots-1, -2 and -3) as key contributors to the formation of stromatolitic laminae in the Dashiling Formation. These microclots form layered, meshed, and dendritic microclotted laminae, characterized by alternation of light and dark laminae, with lighter laminae being sparsely distributed and darker laminae densely distributed. These alternations reflect the periodic growth and decay of microbial communities in response to dynamic environmental changes. The microclots and microclotted laminae in the Dashiling Formation exhibit evidence for microbial calcification, suggesting that atmospheric CO2 levels during the early Paleoproterozoic may have been lower than previously estimated. Previous studies have proposed that microbial calcification occurs extensively only when atmospheric CO2 declines to ∼10 times the present atmospheric level. The well-preserved microclots and calcified microbial structures in the Dashiling Formation suggest that atmospheric CO2 levels during the GOE may have approached or fallen below this threshold. While increased atmospheric O2 levels during the GOE may have influenced microbial calcification, our results indicate that declining CO2 levels played a more direct role in facilitating carbonate precipitation. These findings refine the understanding of the interplay between microbial activities, atmospheric CO2 decline, and carbonate saturation during the GOE, offering new insights into early Earth’s carbon cycle dynamics.
微生物钙化与大气CO2水平的关系是地球生物地球化学演化的一个重要方面。本文首次详细研究了华北五台地区虎沱群2.1 Ga大岭组叠层石的微观和介观特征,该叠层石是大氧化事件(GOE)时期保存最完好的微生物岩之一。研究确定了3种类型的微凝块(微凝块-1、-2和-3)是大岭组叠层石纹层形成的关键因素。这些微凝块形成层状、网状和树突状的微凝层,具有明暗相间的特征,浅色层稀疏分布,深色层密集分布。这些变化反映了微生物群落对动态环境变化的周期性生长和衰减。大岭组的微凝块和微凝片层显示出微生物钙化的证据,表明早古元古代的大气CO2水平可能低于先前的估计。以前的研究已经提出,微生物钙化只有在大气CO2下降到目前大气水平的10倍时才会广泛发生。大岭组保存完好的微凝块和钙化的微生物结构表明,GOE时期的大气CO2水平可能已经接近或低于这一阈值。虽然GOE期间大气中O2水平的增加可能影响了微生物钙化,但我们的研究结果表明,CO2水平的下降在促进碳酸盐沉淀方面发挥了更直接的作用。这些发现完善了对GOE期间微生物活动、大气二氧化碳下降和碳酸盐饱和度之间相互作用的理解,为早期地球碳循环动力学提供了新的见解。
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引用次数: 0
Anatomy and paleoclimatic implications of Cenozoic Coryphoideae palm wood species in Egypt 埃及新生代叶蕨科棕榈树种的解剖学和古气候意义
IF 1.7 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2025-06-06 DOI: 10.1016/j.palwor.2025.200968
Zainab M. El-Noamani, Wagieh E. El-Saadawi, Marwah M. Kamal El-Din, Nermeen A. Ziada
This study examines 12 well-preserved specimens collected from the Miocene Moghra and Gebel Khashab formations at six sites in the northern part of the Western Desert of Egypt. The results reveal distinct anatomical features for each of the four morphospecies (namely Palmoxylon aschersoni Schenk, P. cf. aschersoni Schenk, P. lacunosum (Unger) Felix, and P. libycum (Stenzel) Kräusel), confirming their identification and classification within the subfamily Coryphoideae. The paleoclimatic conditions for each species are interpreted based on key ecologically-based anatomical traits, such as the number of metaxylem elements and the nature of ground tissue. These findings indicate that the four recorded species reflect different paleoenvironments and depositional conditions, suggesting that they were transported by water from their original growth sites to their current locations, resulting in an intermingling of forest forms in the northern Western Desert. An overview of the fossil record of the subfamily Coryphoideae in Africa is also provided.
这项研究检查了在埃及西部沙漠北部的六个地点从中新世Moghra和Gebel Khashab地层收集的12个保存完好的标本。结果揭示了四个形态种(Palmoxylon aschersoni Schenk, P. cf. aschersoni Schenk, P. lacunosum (Unger) Felix和P. libycum (Stenzel) Kräusel)各自的解剖特征,证实了它们在蕨亚科中的鉴定和分类。每个物种的古气候条件都是基于关键的生态解剖学特征来解释的,比如韧皮部元素的数量和地面组织的性质。这些发现表明,这四种记录物种反映了不同的古环境和沉积条件,表明它们被水从原始生长地点转移到现在的位置,导致西部沙漠北部森林形式的混合。本文还概述了非洲斑蝶亚科的化石记录。
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引用次数: 0
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