Palaeoworld最新文献_第9页

New conodont fauna from the Floian (Lower Ordovician) Acoite Formation, Cordillera Oriental, Argentina 阿根廷东部科迪勒拉弗洛ian（下奥陶统）Acoite组牙形石新动物群

IF 1.7 3区地球科学 Q2 PALEONTOLOGY

Palaeoworld

Pub Date : 2025-06-06 DOI: 10.1016/j.palwor.2025.200967

E. Kristal Rueda , Guillermo L. Albanesi , Gladys Ortega

This contribution reports a new species, Trapezognathus morenensis n. sp., from the upper levels of the Acoite Formation outcropping in the El Moreno area of the Cordillera Oriental, northwestern Argentina. This species occurs together with Trapezognathus diprion, Costiconus sp., Drepanodus arcuatus, Drepanoistodus costatus, D. forceps, D. pitjanti, Erraticodon patu, Gothodus costulatus, G. andinus, Kallidontus sp., Paltodus subaequalis, Parapanderodus cf. striatus, Paroistodus parallelus, Protopanderodus rectus, Zentagnathus argentinensis, Gothodus spp., and Drepanoistodontid gen. et sp. indet. Trapezognathus morenensis n. sp. exhibits intermediate characteristics between T. diprion and T. quadrangulum, and also displays a close relationship with early species of the genus Baltoniodus. The conodont assemblage allows for the identification of the Gothodus andinus and Trapezognathus diprion zones, indicating a middle to late Floian (Early Ordovician) age. In addition, the boundary between the Gothodus andinus Zone and the succeeding Trapezognathus diprion Zone is defined by the first occurrence datum of Trapezognathus diprion within the same stratigraphic section, representing the first documented instance of this boundary. The revision of associated taxa with reported Trapezognathus spp. suggests that the definition of the Floian–Dapingian boundary still requires verification in the Cordillera Oriental of Argentina.

本文报道了阿根廷西北部东部科dillera地区El Moreno地区Acoite组露头上层的一新种Trapezognathus morenensis n. sp.。该种与Trapezognathus diprion、Costiconus sp.、arcuatus Drepanoistodus、costatus Drepanoistodus、D. forceps、D. pitjanti、Erraticodon patu、Gothodus costulatus、G. andinus、Kallidontus sp.、Paltodus subaequalis、paratanderodus cfstriatus、paristodus parallelus、Protopanderodus rectus、Zentagnathus argentinensis、Gothodus sp.和Drepanoistodontid gen. et sp. indet一起发生。摘要Trapezognathus morenensis具有介于diprion和quadrangulum之间的中间特征，并与Baltoniodus属的早期物种有密切的亲缘关系。牙形石组合可以识别出Gothodus andinus和Trapezognathus diprius带，表明其时代为弗洛世中晚期（早奥陶世）。此外，Gothodus andinus带与随后的Trapezognathus diprion带之间的边界是由同一地层剖面内的首次Trapezognathus diprion产状基准线确定的，这是该边界的首次文献记录实例。与已报道的Trapezognathus spp.相关分类群的修正表明，在阿根廷东部科迪勒拉，弗洛ian - dapingian界线的定义仍有待验证。

{"title":"New conodont fauna from the Floian (Lower Ordovician) Acoite Formation, Cordillera Oriental, Argentina","authors":"E. Kristal Rueda , Guillermo L. Albanesi , Gladys Ortega","doi":"10.1016/j.palwor.2025.200967","DOIUrl":"10.1016/j.palwor.2025.200967","url":null,"abstract":"<div><div>This contribution reports a new species, Trapezognathus morenensis n. sp., from the upper levels of the Acoite Formation outcropping in the El Moreno area of the Cordillera Oriental, northwestern Argentina. This species occurs together with Trapezognathus diprion, Costiconus sp., Drepanodus arcuatus, Drepanoistodus costatus, D. forceps, D. pitjanti, Erraticodon patu, Gothodus costulatus, G. andinus, Kallidontus sp., Paltodus subaequalis, Parapanderodus cf. striatus, Paroistodus parallelus, Protopanderodus rectus, Zentagnathus argentinensis, Gothodus spp., and Drepanoistodontid gen. et sp. indet. Trapezognathus morenensis n. sp. exhibits intermediate characteristics between T. diprion and T. quadrangulum, and also displays a close relationship with early species of the genus Baltoniodus. The conodont assemblage allows for the identification of the Gothodus andinus and Trapezognathus diprion zones, indicating a middle to late Floian (Early Ordovician) age. In addition, the boundary between the Gothodus andinus Zone and the succeeding Trapezognathus diprion Zone is defined by the first occurrence datum of Trapezognathus diprion within the same stratigraphic section, representing the first documented instance of this boundary. The revision of associated taxa with reported Trapezognathus spp. suggests that the definition of the Floian–Dapingian boundary still requires verification in the Cordillera Oriental of Argentina.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"34 5","pages":"Article 200967"},"PeriodicalIF":1.7,"publicationDate":"2025-06-06","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145048729","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

引用次数: 0

Neogene palaeoenvironments and hydrocarbon potential in the Nile Delta, Egypt: Palynological evidence from an onshore well 埃及尼罗河三角洲新近纪古环境与油气潜力：来自一口陆上井的孢粉学证据

IF 1.7 3区地球科学 Q2 PALEONTOLOGY

Palaeoworld

Pub Date : 2025-06-02 DOI: 10.1016/j.palwor.2025.200966

Mennat-Allah T. El Hussieny, Magdy S. Mahmoud, Amr S. Deaf

Palynological organic matter (POM) of the Neogene succession from the Sidi Salim-1 well, located in the onshore Nile Delta, Egypt (Eastern Mediterranean), suggests a wide range of environments, from deltaic to offshore marine. These environments were discriminated by the overall palynofacies composition, including indicative dinoflagellate cysts, mainly Spiniferites and Selenopemphix. Near-shore marine environment was interpreted for the Middle Miocene (Langhian–Serravallian) Sidi Salim Formation. Deltaic to shallow marine environments were suggested for the Qawasim (Miocene) and Kafr El Sheikh (Pliocene) formations, while the distant (offshore) marine setting was established in the Pliocene Abu Madi Formation. This deeper environment, of the Abu Madi Formation, can be used to confirm a previous documentation of an Early Pliocene progressive drowning of an incised valley, related to the Messinian Salinity Crises (MSC) events, by the late Messinian sea level drop in the Mediterranean. Suboxic to anoxic conditions existed during deposition of the investigated well succession. Anoxia was confirmed by the occurrence of imprints of pyrite crystals across much of the well succession. The occurrence of abundant Poaceae pollen may suggest widespread dry grassland vegetation during deposition of the Neogene sediments of the well. In a regional context, the Neogene environments in the Nile Delta area vary according to the relative position of the investigated sediments, due to structural, palaeogeographic and basinal settings. The recovered palynofacies fluctuated between amorphous organic matter (AOM)-dominated and phytoclast-dominated categories, mostly of the kerogen type II, which is capable of producing oil and gas. The visual assessment of the spore coloration index (SCI) of thin-walled trilete spores in the well section, shows values ranging between 5 and 8, confirming a thermally mature organic matter and, consequently, can be potential source rocks.

位于埃及尼罗河三角洲（东地中海）西迪萨利姆-1井新近系地层的孢粉有机质（POM）显示了从三角洲到近海海洋的广泛环境。这些环境可通过孢粉相的整体组成来区分，包括指示性鞭毛藻囊，主要是刺虫和硒藻。对中中新世（Langhian-Serravallian） Sidi Salim组近岸海洋环境进行了解释。中新世Qawasim组和上新世Kafr El Sheikh组为三角洲-浅海环境，上新世Abu Madi组为远海环境。Abu Madi组的这种较深的环境可以用来证实先前的文献，即上新世早期一个切口山谷的逐渐淹没，与迈西尼亚盐度危机（MSC）事件有关，该事件是由地中海迈西尼亚海平面下降引起的。在所研究的连续井沉积过程中存在亚氧至缺氧条件。在大部分井序列中发现了黄铁矿晶体的印记，证实了缺氧的存在。大量禾科花粉的出现可能表明该井新近纪沉积物沉积期间广泛存在干草地植被。在区域范围内，由于构造、古地理和盆地环境的影响，尼罗河三角洲地区的新近纪环境根据所调查沉积物的相对位置而变化。恢复的孢粉相在以无定形有机质（AOM）为主和以植物碎屑岩为主的类型之间波动，以干酪根ⅱ型为主，具有油气生产能力。对井段薄壁三棱孢子的孢子着色指数（SCI）的目视评价显示，其值在5 ~ 8之间，表明有机质热成熟，可能是潜在的烃源岩。

{"title":"Neogene palaeoenvironments and hydrocarbon potential in the Nile Delta, Egypt: Palynological evidence from an onshore well","authors":"Mennat-Allah T. El Hussieny, Magdy S. Mahmoud, Amr S. Deaf","doi":"10.1016/j.palwor.2025.200966","DOIUrl":"10.1016/j.palwor.2025.200966","url":null,"abstract":"<div><div>Palynological organic matter (POM) of the Neogene succession from the Sidi Salim-1 well, located in the onshore Nile Delta, Egypt (Eastern Mediterranean), suggests a wide range of environments, from deltaic to offshore marine. These environments were discriminated by the overall palynofacies composition, including indicative dinoflagellate cysts, mainly Spiniferites and Selenopemphix. Near-shore marine environment was interpreted for the Middle Miocene (Langhian–Serravallian) Sidi Salim Formation. Deltaic to shallow marine environments were suggested for the Qawasim (Miocene) and Kafr El Sheikh (Pliocene) formations, while the distant (offshore) marine setting was established in the Pliocene Abu Madi Formation. This deeper environment, of the Abu Madi Formation, can be used to confirm a previous documentation of an Early Pliocene progressive drowning of an incised valley, related to the Messinian Salinity Crises (MSC) events, by the late Messinian sea level drop in the Mediterranean. Suboxic to anoxic conditions existed during deposition of the investigated well succession. Anoxia was confirmed by the occurrence of imprints of pyrite crystals across much of the well succession. The occurrence of abundant Poaceae pollen may suggest widespread dry grassland vegetation during deposition of the Neogene sediments of the well. In a regional context, the Neogene environments in the Nile Delta area vary according to the relative position of the investigated sediments, due to structural, palaeogeographic and basinal settings. The recovered palynofacies fluctuated between amorphous organic matter (AOM)-dominated and phytoclast-dominated categories, mostly of the kerogen type II, which is capable of producing oil and gas. The visual assessment of the spore coloration index (SCI) of thin-walled trilete spores in the well section, shows values ranging between 5 and 8, confirming a thermally mature organic matter and, consequently, can be potential source rocks.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"34 5","pages":"Article 200966"},"PeriodicalIF":1.7,"publicationDate":"2025-06-02","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145018824","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

引用次数: 0

Cretaceous–Paleogene palynodebris from two offshore cores in Cape Basin, South Atlantic Ocean and their paleoenvironmental implications 南大西洋Cape盆地两个近海岩心白垩-古近系孢粉碎屑及其古环境意义

IF 1.7 3区地球科学 Q2 PALEONTOLOGY

Palaeoworld

Pub Date : 2025-06-02 DOI: 10.1016/j.palwor.2025.200965

A. Sandersen , L. Scott , F.H. Neumann , E.W. Bergh , M. Bachari

Palynodebris in two cores from the western South African margin provides an opportunity for investigating stratigraphical changes across the Cretaceous/Paleogene (K/Pg) boundary and the depositional environments during this interval. Changing palynodebris trends of more than 200 sidewall cores and cuttings, sampled from two offshore ocean borehole sites ca. 243 km apart and situated in the Cape Basin, reflect the regional environmental processes during the Cretaceous and early Cenozoic. Thirteen categories of microscopic structures of palynomorphs and palynodebris (phytoclasts) are identified and recorded. These categories include amorphous organic matter, marine palynomorphs, fresh-water algae, resins, black debris (charcoal), yellow-brown fragments, black-brown fragments (including microscopic charcoal), cuticles, plant tissues, woody material, sporomorphs, and fungal material such as fungal spores, fruit bodies, tubes and filaments. The relative abundances of these organic components from each site were analyzed and compared. Palynodebris compositions and patterns differ between borehole sites over time but also show some similar regional trends, e.g., growing indications of burning from the Late Cretaceous to early Paleocene. The woody component suggests that both offshore sites received similar land-derived Cretaceous and Cenozoic sediments via fluvial transport. Palynodebris was compared to previously published palynological data from the cores, showing shifts in vegetation and climate. Around and shortly after the K/Pg boundary, black debris and yellow-brown debris components fluctuate strongly and could possibly be related more to the transition from the Maastrichtian regressive events than a terminal K/Pg boundary event. Stronger marine influence, as indicated by an increase of marine palynomorphs together with geological evidence of glauconitic sandstone, Inoceramus, other bivalves, and shelly material during the Late Cretaceous and Paleogene, reflects the flooding of the basin. The stratigraphy and depositional environments contribute to a better understanding of conditions associated with the opening of the South Atlantic Ocean.

来自南非西部边缘的两个岩心孢粉碎屑为研究白垩纪/古近纪（K/Pg）界线的地层变化及其沉积环境提供了机会。在Cape盆地两个相距约243 km的海上钻孔中采集的200多个岩心和岩屑的孢粉碎屑变化趋势反映了白垩纪和早新生代的区域环境过程。鉴定并记录了孢粉和孢粉碎屑（植物碎屑）的13类显微结构。这些类别包括无定形有机物、海洋苔藓、淡水藻类、树脂、黑色碎片（木炭）、黄棕色碎片、黑棕色碎片（包括微观木炭）、角质层、植物组织、木质材料、孢子形和真菌材料，如真菌孢子、果体、管和细丝。分析比较了各地点有机组分的相对丰度。随着时间的推移，不同钻孔地点的孢粉碎屑组成和模式有所不同，但也显示出一些相似的区域趋势，例如，从晚白垩纪到早古新世，燃烧的迹象越来越多。木本成分表明，这两个海上遗址通过河流运输获得了相似的陆源白垩纪和新生代沉积物。将孢粉碎屑与先前发表的岩心孢粉学数据进行比较，显示出植被和气候的变化。在K/Pg边界附近和之后不久，黑色碎屑和黄棕色碎屑组分波动强烈，可能更多地与马斯特里赫特海退事件的过渡有关，而不是与K/Pg边界事件有关。在晚白垩世和古近纪期间，海相地貌的增加以及海绿石砂岩、头壳类、其他双壳类和壳壳类物质的地质证据表明，海相影响较强，反映了盆地的洪水。地层学和沉积环境有助于更好地了解与南大西洋开放有关的条件。

{"title":"Cretaceous–Paleogene palynodebris from two offshore cores in Cape Basin, South Atlantic Ocean and their paleoenvironmental implications","authors":"A. Sandersen , L. Scott , F.H. Neumann , E.W. Bergh , M. Bachari","doi":"10.1016/j.palwor.2025.200965","DOIUrl":"10.1016/j.palwor.2025.200965","url":null,"abstract":"<div><div>Palynodebris in two cores from the western South African margin provides an opportunity for investigating stratigraphical changes across the Cretaceous/Paleogene (K/Pg) boundary and the depositional environments during this interval. Changing palynodebris trends of more than 200 sidewall cores and cuttings, sampled from two offshore ocean borehole sites ca. 243 km apart and situated in the Cape Basin, reflect the regional environmental processes during the Cretaceous and early Cenozoic. Thirteen categories of microscopic structures of palynomorphs and palynodebris (phytoclasts) are identified and recorded. These categories include amorphous organic matter, marine palynomorphs, fresh-water algae, resins, black debris (charcoal), yellow-brown fragments, black-brown fragments (including microscopic charcoal), cuticles, plant tissues, woody material, sporomorphs, and fungal material such as fungal spores, fruit bodies, tubes and filaments. The relative abundances of these organic components from each site were analyzed and compared. Palynodebris compositions and patterns differ between borehole sites over time but also show some similar regional trends, e.g., growing indications of burning from the Late Cretaceous to early Paleocene. The woody component suggests that both offshore sites received similar land-derived Cretaceous and Cenozoic sediments via fluvial transport. Palynodebris was compared to previously published palynological data from the cores, showing shifts in vegetation and climate. Around and shortly after the K/Pg boundary, black debris and yellow-brown debris components fluctuate strongly and could possibly be related more to the transition from the Maastrichtian regressive events than a terminal K/Pg boundary event. Stronger marine influence, as indicated by an increase of marine palynomorphs together with geological evidence of glauconitic sandstone, Inoceramus, other bivalves, and shelly material during the Late Cretaceous and Paleogene, reflects the flooding of the basin. The stratigraphy and depositional environments contribute to a better understanding of conditions associated with the opening of the South Atlantic Ocean.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"34 5","pages":"Article 200965"},"PeriodicalIF":1.7,"publicationDate":"2025-06-02","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145018791","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

引用次数: 0

Dating of Early Triassic Chaohu Fauna by precise U-Pb age constraints: Temporal calibration of the emergence and first radiation of Mesozoic marine reptiles 基于精确U-Pb年龄约束的早三叠世巢湖动物群测年：中生代海洋爬行动物出现和首次辐射的时间定标

IF 1.7 3区地球科学 Q2 PALEONTOLOGY

Palaeoworld

Pub Date : 2025-05-26 DOI: 10.1016/j.palwor.2025.200963

Yi-Nuo Wang , Da-Yong Jiang , Ryosuke Motani , Ming-Tao Yao , Min Zhou , Zuo-Yu Sun , Jian-Dong Huang , Zhao-Liang Ma

The emergence time of Mesozoic marine reptiles has been controversial due to poor stratigraphic control and the lack of radiometric age constraints. We report a radiometric age of 248.7 ± 1.0 Ma (2σ), obtained through LA-ICP-MS analysis of the zircon grains sampled from a tuff which is intercalated within the fossiliferous beds of the Chaohu Fauna and layered about 7 m above the lowest fossiliferous bed. This dating confirms the Chaohu Fauna as the oldest known marine reptile fauna in the world with precise and comprehensive age constraints. Integrated with the biostratigraphic data and astrochronological analysis, our results demonstrate that marine reptiles were established at least by 3.2 million years after the Permian–Triassic boundary.

中生代海生爬行动物的出现时间由于地层控制不佳和缺乏放射性年龄约束一直存在争议。通过LA-ICP-MS分析，我们获得了248.7±1.0 Ma （2σ）的放射性年龄，这些锆石颗粒来自于位于巢湖动物群化石层中，并在最低化石层之上约7 m处的凝灰岩。这证实了巢湖动物群是世界上已知最古老的海洋爬行动物群，具有精确和全面的年龄限制。结合生物地层资料和天体年代学分析，我们的研究结果表明，海洋爬行动物至少在二叠纪-三叠纪界线后320万年建立。

引用次数: 0

Multielement conodont apparatus (Cornudina) from the Lower Triassic of China, revealed by high-resolution X-ray microtomography 中国下三叠统多元素牙形刺器（Cornudina）的高分辨率x射线显微层析成像

IF 1.7 3区地球科学 Q2 PALEONTOLOGY

Palaeoworld

Pub Date : 2025-05-26 DOI: 10.1016/j.palwor.2025.200964

Wu-Jun Xu , Zuo-Yu Sun , Shuang Liu , Cheng Ji , Hao Lu , Zhao-Liang Ma , Fei-Yang Wu , Shou-Ming Chen , Jian-Dong Huang

The apparatus of the coniform conodont genus Cornudina remains equivocal because the previous results were derived from discrete element collections rather than natural assemblages. Four exceptionally preserved natural assemblages collected from bed 719 of the upper Nanlinghu Formation in Majiashan quarry, Chaohu of Hefei City, Anhui Province, East China, were imaged using high-resolution X-ray microtomography. The consistent and repeated patterns of element juxtaposition suggest an apparatus of 15 elements, conforming to the classical 2M-9S-4P skeletal template that comprises three basic morphologies: two breviform dolabrate elements, nine elongate bipennate elements, and four angulate elements. The angulate P₁ and P₂ elements have a distinct cusp, short anterior and posterior processes and the P₂ element is twisted crescent-shaped, which supports a taxonomic assignment to Cornudina breviramulis sensu stricto Koike. Multielement comparisons among Cornudina, Neostrachanognathus, and Hindeodus apparatuses suggest Cornudina and Neostrachanognathus are closely related and possibly derived from Hindeodus. The multi-element comparison between Ordovician ‘complex’ conodont apparatuses (e.g., Paracordylodus and Phragmodus) and Triassic ‘coniform’ apparatuses (e.g., Cornudina and Neostrachanognathus) suggests that the low variation in the elemental forms of the latter is probably a secondary ‘evolution’.

锥形牙形刺属的装置仍然是模棱两可的，因为以前的结果是来自离散元素集合而不是自然组合。利用高分辨率x射线显微层析成像技术，对安徽省合肥市巢湖马家山采石场南陵湖上组719层采集的4个保存特别完好的自然组合进行了成像。元素并置的一致和重复的模式表明了一个由15个元素组成的装置，符合经典的2M-9S-4P骨架模板，包括三种基本形态：两个短形dolabate元素，九个细长的bipenate元素和四个角形元素。P1和P2呈角状，尖部明显，前后突短，P2呈弯月牙状，属于corudina breviramulis sensu stricto Koike。Cornudina、Neostrachanognathus和Hindeodus器械的多元素比较表明Cornudina和Neostrachanognathus亲缘关系密切，可能来源于Hindeodus。奥陶系“复杂”牙形器（如Paracordylodus和Phragmodus）与三叠系“形形”器（如Cornudina和Neostrachanognathus）的多元素比较表明，后者元素形式的低变化可能是二次“进化”。

{"title":"Multielement conodont apparatus (Cornudina) from the Lower Triassic of China, revealed by high-resolution X-ray microtomography","authors":"Wu-Jun Xu , Zuo-Yu Sun , Shuang Liu , Cheng Ji , Hao Lu , Zhao-Liang Ma , Fei-Yang Wu , Shou-Ming Chen , Jian-Dong Huang","doi":"10.1016/j.palwor.2025.200964","DOIUrl":"10.1016/j.palwor.2025.200964","url":null,"abstract":"<div><div>The apparatus of the coniform conodont genus Cornudina remains equivocal because the previous results were derived from discrete element collections rather than natural assemblages. Four exceptionally preserved natural assemblages collected from bed 719 of the upper Nanlinghu Formation in Majiashan quarry, Chaohu of Hefei City, Anhui Province, East China, were imaged using high-resolution X-ray microtomography. The consistent and repeated patterns of element juxtaposition suggest an apparatus of 15 elements, conforming to the classical 2M-9S-4P skeletal template that comprises three basic morphologies: two breviform dolabrate elements, nine elongate bipennate elements, and four angulate elements. The angulate P1 and P2 elements have a distinct cusp, short anterior and posterior processes and the P2 element is twisted crescent-shaped, which supports a taxonomic assignment to Cornudina breviramulis sensu stricto Koike. Multielement comparisons among Cornudina, Neostrachanognathus, and Hindeodus apparatuses suggest Cornudina and Neostrachanognathus are closely related and possibly derived from Hindeodus. The multi-element comparison between Ordovician ‘complex’ conodont apparatuses (e.g., Paracordylodus and Phragmodus) and Triassic ‘coniform’ apparatuses (e.g., Cornudina and Neostrachanognathus) suggests that the low variation in the elemental forms of the latter is probably a secondary ‘evolution’.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"34 5","pages":"Article 200964"},"PeriodicalIF":1.7,"publicationDate":"2025-05-26","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145018784","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

引用次数: 0

New findings in the Permian xyloflora of Carapacha Formation (Argentina) 阿根廷Carapacha组二叠纪木区系的新发现

IF 1.7 3区地球科学 Q2 PALEONTOLOGY

Palaeoworld

Pub Date : 2025-05-20 DOI: 10.1016/j.palwor.2025.200962

Paola Zuliani , Alexandra Crisafulli , Ricardo Melchor , Laura Vallejos Leiz

The Carapacha Formation consists of two members: the Calencó (lower) and the Urre-Lauquen (upper), dated as Permian based on its glossopterid flora. The formation outcrops in La Pampa Province, near the locality of Puelches, with sediments of continental origin, fluvio-lacustrine in type. This study describes two gymnosperm woods species. One, from the Calencó Member, is a Cordaitalean assigned to Scleromedulloxylon aveyronense found in the Permian of Aveyron, France, characterized by having a heterocellular pith with islands of sclerenchymatic cells and unorganized lacunae in transverse section. The other specimen, from the Urre-Lauquen Member, is a conifer of uncertain affinity, Zalesskioxylon uniseriatum, previously reported from the Raniganj (India), Yaguarí (Uruguay), and Tacuary (Paraguay) formations. Both woods exhibit secondary xylem features typical of Paleozoic gymnosperms, particularly concerning the radial system and the arrangement of pits in the tracheid walls. These xylological records confirm that the Carapacha Formation comprises another Permian sedimentological sequences that contain woods with diaphragm-solenoid pith such as the taxa found in the Melo (Uruguay), the Río Bonito, and the Irati (Brazil), the White Band (Africa), the Barakar, and the Raniganj (India) formations. This palaeofloristic association is part of the Australoxylon xylotaphoflora characteristic of the Gondwanan Permian. Finally, this study allows us to infer that the environmental conditions in this formation were warm with low humidity and seasonally dry.

Carapacha组由Calencó（下）和Urre-Lauquen（上）两段组成，根据其舌状植物区系可确定为二叠纪。该组露头位于拉潘帕省，靠近普埃尔切斯地区，沉积物为陆相、河流湖相。本研究描述了两个裸子植物树种。其中一个来自Calencó成员，属于法国Aveyron二叠纪的Scleromedulloxylon aveyronense，其特征是具有异细胞髓，具有岛状的厚壁细胞和横剖面上无组织的腔隙。另一个标本来自Urre-Lauquen段，是一种不确定亲缘关系的针叶树，Zalesskioxylon uniseriatum，以前报道过来自Raniganj（印度）、Yaguarí（乌拉圭）和Tacuary（巴拉圭）地层。这两种木材都具有古生代裸子植物的次生木质部特征，特别是在管胞壁的径向系统和凹坑的排列方面。这些木学记录证实，Carapacha组包含另一个二叠纪沉积序列，其中包含具有膜片螺线管髓的木材，例如在Melo（乌拉圭）、Río Bonito、Irati（巴西）、White Band（非洲）、Barakar和Raniganj（印度）组中发现的分类群。这种古植物群组合是冈瓦南二叠纪特有的Australoxylon xylotaphoflora的一部分。最后，本研究使我们推断出该地层的环境条件是温暖，低湿度和季节性干燥。

{"title":"New findings in the Permian xyloflora of Carapacha Formation (Argentina)","authors":"Paola Zuliani , Alexandra Crisafulli , Ricardo Melchor , Laura Vallejos Leiz","doi":"10.1016/j.palwor.2025.200962","DOIUrl":"10.1016/j.palwor.2025.200962","url":null,"abstract":"<div><div>The Carapacha Formation consists of two members: the Calencó (lower) and the Urre-Lauquen (upper), dated as Permian based on its glossopterid flora. The formation outcrops in La Pampa Province, near the locality of Puelches, with sediments of continental origin, fluvio-lacustrine in type. This study describes two gymnosperm woods species. One, from the Calencó Member, is a Cordaitalean assigned to Scleromedulloxylon aveyronense found in the Permian of Aveyron, France, characterized by having a heterocellular pith with islands of sclerenchymatic cells and unorganized lacunae in transverse section. The other specimen, from the Urre-Lauquen Member, is a conifer of uncertain affinity, Zalesskioxylon uniseriatum, previously reported from the Raniganj (India), Yaguarí (Uruguay), and Tacuary (Paraguay) formations. Both woods exhibit secondary xylem features typical of Paleozoic gymnosperms, particularly concerning the radial system and the arrangement of pits in the tracheid walls. These xylological records confirm that the Carapacha Formation comprises another Permian sedimentological sequences that contain woods with diaphragm-solenoid pith such as the taxa found in the Melo (Uruguay), the Río Bonito, and the Irati (Brazil), the White Band (Africa), the Barakar, and the Raniganj (India) formations. This palaeofloristic association is part of the Australoxylon xylotaphoflora characteristic of the Gondwanan Permian. Finally, this study allows us to infer that the environmental conditions in this formation were warm with low humidity and seasonally dry.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"34 5","pages":"Article 200962"},"PeriodicalIF":1.7,"publicationDate":"2025-05-20","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145018780","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

引用次数: 0

The oldest known Curvicubitidae (Hemiptera) from the Permian (Wuchiapingian) of Zhangshu City, Jiangxi Province, southern China 江西省张树市二叠纪（五家坪纪）已知最古老的弯蝶科（半翅目）

IF 1.7 3区地球科学 Q2 PALEONTOLOGY

Palaeoworld

Pub Date : 2025-05-20 DOI: 10.1016/j.palwor.2025.200959

Qian-Qi Zhang , Lin Mu , Lei Zhang , Xu-Heng Du , Hai-Chun Zhang

As the earliest record of the family Curvicubitidae Hong, 1984, Sinocurvicubitus qingjiangensis n. gen. n. sp. is established based on a cicada tegmen collected from the Laoshan Member of the Leping Formation in the Qingjiang Basin, Jiangxi Province, southern China. This new taxon is characterised by vein R forked at basal 1/3rd tegminal length, indentation at basal 2/3rds length, vein M first forked on nodal line, more developed M and less developed CuA. A Wuchiapingian age for this new species is suggested by the co-occurrence of the ammonoids Araxoceras tricarinatus and Pseudogastrioceras sp. The new find not only extends the known palaeogeographical range of curvicubitids in China, but also suggests an origin of this family in the Palaeozoic. This new record, combined with other records of Curvicubitidae, suggests a possibility of simultaneously northward and southward migration and dispersal of this family during the late Permian to the Late Triassic.

Sinocurvicubitus qingjiangensis n. gen. n. sp.根据江西清江盆地乐平组崂山段的蝉甲，于1984年建立，是curvicutidae科最早记录。该新分类群的特征为脉R在基部1/3顶尖处分叉，脉M在基部2/3处凹陷，脉M在节线上先分叉，脉M较发达，脉CuA较不发达。这一新发现不仅扩大了曲丘类动物在中国已知的古地理分布范围，而且表明曲丘类动物起源于古生代。这一新的记录，结合其他的曲丘虫科记录，表明该科在晚二叠世至晚三叠世期间可能同时向北和向南迁移和分散。

引用次数: 0

Phylogenetic evolution of Neorotalia mecatepecensis to Miogypsinidae from Rajamandala Formation, West Java, Indonesia 印度尼西亚西爪哇Rajamandala组mecatepecneorotalia meatepecensis至Miogypsinidae的系统进化

IF 1.7 3区地球科学 Q2 PALEONTOLOGY

Palaeoworld

Pub Date : 2025-05-20 DOI: 10.1016/j.palwor.2025.200961

Nadila Novandaru , Khoiril Anwar Maryunani , Rubiyanto Kapid , Aswan , Wahyu Budhi Khorniawan

At Tagogapu in West Java, the Miogypsinidae larger benthic foraminifera from the Rajamandala Limestone have been studied for biometric trends in evolution. Observed taxa include Neorotalia mecatepecensis, Paleomiogypsina sp., Miogypsinella bornea, Miogypsinoides complanatus, and Miogypsinoides formosensis, with biometric parameters (X, Y, Z, AX, and γ) revealing distinct phylogenetic trends through heterochrony. Neorotalia mecatepecensis morphotype A is identified as the basal species, while Neorotalia mecatepecensis morphotype B diverges through nepionic retardation. The transition to Paleomiogypsina sp. is marked by the development of auxiliary chambers. The transition to Miogypsinella bornea is suggested by the addition of secondary foramen and more auxiliary chambers. The evolution to Miogypsinoides complanatus is characterized by the planispiral nepionic, loss of an apertural lip, and broader auxiliary chamber arrangements. The speciation of Miogypsinoides formosensis is recognized by the peramorphic addition of late adult stage auxiliary chambers. Biostratigraphic analysis and strontium isotopic dating place the studied taxa within the Te 2-3 to Te 4 Letter Stage (late Oligocene). This research highlights the biostratigraphic application of biometric methods in a lineage with heterochronous evolutionary trends, and proposes AX as a robust metric for the descendant peramorphic forms.

在西爪哇的Tagogapu，研究了来自Rajamandala石灰岩的Miogypsinidae大型底栖有孔虫的生物特征进化趋势。观察到的分类群包括Neorotalia mecatepecensis、Paleomiogypsina sp.、Miogypsinella bornea、miogypsinides planatus和miogypsinides formosensis，其生物特征参数（X、Y、Z、AX和γ）通过异时性揭示了不同的系统发育趋势。Neorotalia mecatepecensis形态型A被确定为基种，而Neorotalia mecatepecensis形态型B通过nepionic迟滞分化。向Paleomiogypsina sp.过渡的标志是辅助室的发育。通过增加次级孔和更多的辅助腔室，提示向密氏肺菌过渡。向平腹小齿进化的特征是平螺旋形的唇形，失去开口唇形，辅助腔室布置更宽。台湾miogypsinides formosensis的形态是由成虫后期辅助腔的变质性添加来识别的。生物地层分析和锶同位素定年将研究类群确定在晚渐新世的第2-3 ~第4字母阶段。本研究强调了生物测量方法在具有异时进化趋势的谱系中的生物地层学应用，并提出AX作为后代变质岩形式的稳健度量。

{"title":"Phylogenetic evolution of Neorotalia mecatepecensis to Miogypsinidae from Rajamandala Formation, West Java, Indonesia","authors":"Nadila Novandaru , Khoiril Anwar Maryunani , Rubiyanto Kapid , Aswan , Wahyu Budhi Khorniawan","doi":"10.1016/j.palwor.2025.200961","DOIUrl":"10.1016/j.palwor.2025.200961","url":null,"abstract":"<div><div>At Tagogapu in West Java, the Miogypsinidae larger benthic foraminifera from the Rajamandala Limestone have been studied for biometric trends in evolution. Observed taxa include Neorotalia mecatepecensis, Paleomiogypsina sp., Miogypsinella bornea, Miogypsinoides complanatus, and Miogypsinoides formosensis, with biometric parameters (X, Y, Z, AX, and γ) revealing distinct phylogenetic trends through heterochrony. Neorotalia mecatepecensis morphotype A is identified as the basal species, while Neorotalia mecatepecensis morphotype B diverges through nepionic retardation. The transition to Paleomiogypsina sp. is marked by the development of auxiliary chambers. The transition to Miogypsinella bornea is suggested by the addition of secondary foramen and more auxiliary chambers. The evolution to Miogypsinoides complanatus is characterized by the planispiral nepionic, loss of an apertural lip, and broader auxiliary chamber arrangements. The speciation of Miogypsinoides formosensis is recognized by the peramorphic addition of late adult stage auxiliary chambers. Biostratigraphic analysis and strontium isotopic dating place the studied taxa within the Te 2-3 to Te 4 Letter Stage (late Oligocene). This research highlights the biostratigraphic application of biometric methods in a lineage with heterochronous evolutionary trends, and proposes AX as a robust metric for the descendant peramorphic forms.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"34 5","pages":"Article 200961"},"PeriodicalIF":1.7,"publicationDate":"2025-05-20","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145018792","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

引用次数: 0

A cranium of “Gazella” nihensis from Pliocene of Qinghai-Tibetan Plateau and the differentiation of early Antilopina

IF 1.7 3区地球科学 Q2 PALEONTOLOGY

Palaeoworld

Pub Date : 2025-05-20 DOI: 10.1016/j.palwor.2025.200960

Shi-Qi Wang , Xiao-Yong Zhang , Yi-Kun Li , Xiao-Xiao Zhang , Qi-Gao Jiangzuo

Fossils of Antilopina were often referred to and classified under the genus name “Gazella”, except for some Miocene spiral horned antelopes; however, their true taxonomic attribution is difficult to determine due to the significant morphological variation observed in large fossil samples. Species of two extant genera within Antilopina that are widely distributed in China are Gazella subgutturosa and Procapra spp. Assigning abundant fossil specimens identified as “Gazella” from China to these two extant groups is particularly challenging. Molecular phylogenetic studies have revealed that Gazella and Procapra belong to two distinct clades within Antilopina. Therefore, the study of fossil “Gazella” is crucial for understanding the differentiation and evolutionary history of Antilopina, which includes a diverse array of genera that lived in Eurasia and Africa. In this article, we describe a newly discovered cranium of “Gazella” nihensis from the Pliocene Epoch on the Qinghai-Tibetan Plateau. The new cranium exhibits several distinctive features: the braincase is considerably elongated, with the occipital condyles protruding caudally beyond the paroccipital process; the cranial-facial axis is weakly curved, and the horn cores are strongly inclined caudally. The morphology is distinct from most other members of Antilopina. We testified the view that “Gazella” nihensis is possibly related with the living Procapra because of these specialized features. Interestingly, in “Gazella” nihensis, the elongated braincase that is less bent from the facial axis resembles that of the living Litocranius walleri; and the strongly caudally inclined horn core that of the living Ammodorcas clarkei and Antidorcas marsupialis. These similarities may be the result of parallel evolution, but further research is needed to explore this interesting resemblance.

除了一些中新世的螺旋角羚羊外，Antilopina的化石经常被称为Gazella，并被归类为属名；然而，由于在大量化石样本中观察到明显的形态差异，它们的真正分类归属难以确定。在中国广泛分布的羚羊属（Antilopina）中现存的两个属是Gazella subgutturrosa和Procapra spp，将来自中国的大量被鉴定为“Gazella”的化石标本分配给这两个现存的类群尤其具有挑战性。分子系统发育研究表明，Gazella和Procapra属于Antilopina中两个不同的分支。因此，对“Gazella”化石的研究对于了解Antilopina的分化和进化历史至关重要，Antilopina包括生活在欧亚大陆和非洲的多种属。新颅骨表现出几个独特的特征：脑壳相当长，枕髁尾侧突出，超出枕旁突；颅面轴呈弱弯曲，角核向尾端强烈倾斜。其形态不同于Antilopina的大多数其他成员。由于这些特殊的特征，我们证明了“Gazella”nihensis可能与现存的原甲龙有关的观点。有趣的是，在“Gazella”nihensis中，较长的脑壳与面部轴的弯曲程度较低，与现存的walleri Litocranius相似；现存的clarkei Ammodorcas和Antidorcas marsupialis的角核强烈地向尾端倾斜。这些相似性可能是平行进化的结果，但需要进一步的研究来探索这种有趣的相似性。

{"title":"A cranium of “Gazella” nihensis from Pliocene of Qinghai-Tibetan Plateau and the differentiation of early Antilopina","authors":"Shi-Qi Wang , Xiao-Yong Zhang , Yi-Kun Li , Xiao-Xiao Zhang , Qi-Gao Jiangzuo","doi":"10.1016/j.palwor.2025.200960","DOIUrl":"10.1016/j.palwor.2025.200960","url":null,"abstract":"<div><div>Fossils of Antilopina were often referred to and classified under the genus name “Gazella”, except for some Miocene spiral horned antelopes; however, their true taxonomic attribution is difficult to determine due to the significant morphological variation observed in large fossil samples. Species of two extant genera within Antilopina that are widely distributed in China are Gazella subgutturosa and Procapra spp. Assigning abundant fossil specimens identified as “Gazella” from China to these two extant groups is particularly challenging. Molecular phylogenetic studies have revealed that Gazella and Procapra belong to two distinct clades within Antilopina. Therefore, the study of fossil “Gazella” is crucial for understanding the differentiation and evolutionary history of Antilopina, which includes a diverse array of genera that lived in Eurasia and Africa. In this article, we describe a newly discovered cranium of “Gazella” nihensis from the Pliocene Epoch on the Qinghai-Tibetan Plateau. The new cranium exhibits several distinctive features: the braincase is considerably elongated, with the occipital condyles protruding caudally beyond the paroccipital process; the cranial-facial axis is weakly curved, and the horn cores are strongly inclined caudally. The morphology is distinct from most other members of Antilopina. We testified the view that “Gazella” nihensis is possibly related with the living Procapra because of these specialized features. Interestingly, in “Gazella” nihensis, the elongated braincase that is less bent from the facial axis resembles that of the living Litocranius walleri; and the strongly caudally inclined horn core that of the living Ammodorcas clarkei and Antidorcas marsupialis. These similarities may be the result of parallel evolution, but further research is needed to explore this interesting resemblance.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"34 5","pages":"Article 200960"},"PeriodicalIF":1.7,"publicationDate":"2025-05-20","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145018829","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

引用次数: 0

Middle Campanian to Maastrichtian Foraminifera and biomarkers: Seawater incursion events in the Songliao Basin, Northeast China 松辽盆地中坎帕阶—马斯特里赫特阶有孔虫及其生物标志：海水入侵事件

IF 1.7 3区地球科学 Q2 PALEONTOLOGY

Palaeoworld

Pub Date : 2025-05-16 DOI: 10.1016/j.palwor.2025.200958

Ke-Ying Yue , Sha Li , Tian Jiang , Dang-Peng Xi , Bo-Wen Song , Yu Liang , Li-Xin Sun , Tian-Fu Zhang , Hai-Chun Zhang , Xiao-Qiao Wan , Bo Wang

The Songliao Basin, spanning approximately 260,000 km² in Northeast China, is one of the largest oil-bearing lacustrine basins. Seawater incursion events have previously been discovered in the Turonian (lower and the uppermost Qingshankou Formation) to the Santonian and early Campanian (the first and second members of the Nenjiang Formation). We newly discovered benthic and planktonic foraminifera in the middle Campanian to Maastrichtian (Fifth Member of the Nenjiang Formation, Sifangtai, and Mingshui formations) in the ZKD-1 borehole. The Planktonic foraminifera include Archaeoglobigerina blowi, A. cretacea, and Globotruncanella sp. The benthic foraminifera include Anomalinoides sp., Gavelinella sp., and Haplophragmoides sp. Among them, A. blowi, A. cretacea, Anomalinoides sp., Gavelinella sp., Haplophragmoides sp. are widely distributed in the marine strata of the Upper Cretaceous. We tested the marine biomarker compounds including C₃₀ 24-n-propyl cholestane and C₃₀ 24-iso-propyl cholestane, indicating that the Songliao Basin experienced seawater incursion events to varying degrees during the middle to late Campanian. We also found abundant charophyte fossils in the ZKD-1 borehole, including Atopochara trivolvis ulanensis, Latochara yuananensis, Feistiella anluensis, Mesochara biacuta, M. gobica, and M. leiocarpa. Among them, the brackish water charophyte fossils of Feistiella anluensis were first found in Songliao Basin.

松辽盆地位于中国东北，面积约26万平方公里，是中国最大的含油湖盆之一。在吐鲁番统（青山口组下段和上段）至三东统和早坎帕纪（嫩江组一段和二段）均发现过海水入侵事件。在ZKD-1钻孔新发现坎帕阶中—马斯特里赫特（嫩江组五段、四方台组、明水组）底栖和浮游有孔虫。浮游有孔虫包括Archaeoglobigerina blowi、A.白垩纪、Globotruncanella sp.底栖有孔虫包括Anomalinoides sp.、Gavelinella sp.、Haplophragmoides sp.，其中A. blowi、A.白垩纪、Anomalinoides sp.、Gavelinella sp.、Haplophragmoides sp.广泛分布于上白垩统海相地层中。通过对C30 24-正丙基胆甾和C30 24-异丙基胆甾等海洋生物标志物的检测，表明松辽盆地在坎帕拉世中晚期经历了不同程度的海水入侵事件。在zpd -1钻孔中还发现了丰富的蕨类化石，包括Atopochara trivolvis ulanensis、Latochara yuananensis、Feistiella anluensis、Mesochara biacuta、M. gobica和M. leiocarpa。其中，在松辽盆地首次发现安露费斯提拉（Feistiella anluensis）半咸淡水蕨类化石。

{"title":"Middle Campanian to Maastrichtian Foraminifera and biomarkers: Seawater incursion events in the Songliao Basin, Northeast China","authors":"Ke-Ying Yue , Sha Li , Tian Jiang , Dang-Peng Xi , Bo-Wen Song , Yu Liang , Li-Xin Sun , Tian-Fu Zhang , Hai-Chun Zhang , Xiao-Qiao Wan , Bo Wang","doi":"10.1016/j.palwor.2025.200958","DOIUrl":"10.1016/j.palwor.2025.200958","url":null,"abstract":"<div><div>The Songliao Basin, spanning approximately 260,000 km2 in Northeast China, is one of the largest oil-bearing lacustrine basins. Seawater incursion events have previously been discovered in the Turonian (lower and the uppermost Qingshankou Formation) to the Santonian and early Campanian (the first and second members of the Nenjiang Formation). We newly discovered benthic and planktonic foraminifera in the middle Campanian to Maastrichtian (Fifth Member of the Nenjiang Formation, Sifangtai, and Mingshui formations) in the ZKD-1 borehole. The Planktonic foraminifera include Archaeoglobigerina blowi, A. cretacea, and Globotruncanella sp. The benthic foraminifera include Anomalinoides sp., Gavelinella sp., and Haplophragmoides sp. Among them, A. blowi, A. cretacea, Anomalinoides sp., Gavelinella sp., Haplophragmoides sp. are widely distributed in the marine strata of the Upper Cretaceous. We tested the marine biomarker compounds including C30 24-n-propyl cholestane and C30 24-iso-propyl cholestane, indicating that the Songliao Basin experienced seawater incursion events to varying degrees during the middle to late Campanian. We also found abundant charophyte fossils in the ZKD-1 borehole, including Atopochara trivolvis ulanensis, Latochara yuananensis, Feistiella anluensis, Mesochara biacuta, M. gobica, and M. leiocarpa. Among them, the brackish water charophyte fossils of Feistiella anluensis were first found in Songliao Basin.</div></div>","PeriodicalId":48708,"journal":{"name":"Palaeoworld","volume":"34 5","pages":"Article 200958"},"PeriodicalIF":1.7,"publicationDate":"2025-05-16","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144170130","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}

引用次数: 0